Beruflich Dokumente
Kultur Dokumente
Without doubt, this excellent catalogue, compiled with diligence and knowledge, will be an important tool for anyone,
not only specialists on Balkan Carabidae, and will improve our
insights into the systematics and distribution of carabid beetles.
The hope of the author to place Albania with this catalogue side
by side to Bulgaria (certainly the best-known regional fauna of
Carabidae in the Balkan Peninsula, catalogued in 1995 by the
same author and his father) is surely fulfilled.
David Wrase
ANNOTATED CATALOGUE
of the
Carabid Beetles
of Albania
(Coleoptera: Carabidae)
by
B.V. Guorguiev
SOFIAMOSCOW
2007
PENSOFT Publishers
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Contents
Introduction 7
Background of the study 7
Aims of the study 8
Short description of the studied area 8
Treatment of the taxa 10
Material and depositories of the collections 10
Acknowledgments 11
Abbreviations 13
Topographic treatment of some localities 14
Systematic part 16
Results 187
Conclusions 188
Taxonomic structure of the Albanian Carabidae (Table 1) 189
Appendix 193
References 205
Index 224
Introduction
The ground beetle fauna of Albania is not very well known taking into account the
fact that an overall investigation based upon studied material and critical survey on
the literature has never been made. So far, our knowledge of this fauna is still largely
due to Apfelbecks well-known monograph (Apfelbeck, 1904) and other works of the
same author published in the period 1901-1930 (Apfelbeck, 1901a; 1901b; 1902a;
1903; 1904; 1905; 1906; 1907a; 1907b; 1909; 1911; 1916; 1917; 1918a; 1918b; 1918c;
1919a; 1919b; 1930). Later, important faunistic contributions were made also by Csiki
(1940), Schatzmayr (1943b), and Hieke et al. (1986). Dealing with the Bulgarian ground
beetles, Hieke & Wrase (1988) mentioned 296 species (Nebriinae 14, Cicindelinae
5, Carabinae 10, Omophroninae 1, Brachininae 10, Siagoninae 1, Scaritinae
12, Trechinae 58, Harpalinae 185) for Albania without detailed records. For political reasons, in the period 1945-1990 not many faunistic records were published, and
relatively few materials from the country were available in the collections. Since 1990
collecting ground beetles there became possible again. Recently, 363 species (Nebriinae 10, Cicindelinae 8, Carabinae 18, Omophroninae 1, Brachininae 11,
Siagoninae 1, Scaritinae 20, Apotominae 1, Trechinae 55, Harpalinae 238)
have been mentioned for Albania in the most recent catalogue of Palaearctic Coleoptera by Lbl & Smetana (2003).
Examination of the available collections added information about 55 new species
for the country plus 119 species omitted in the last work as well as many new faunistic
data for other species. The subsequent critical survey of the literature data brought to
the conclusion that 543 species of Carabidae live in Albania. Moreover, some taxonomic changes and a lot of comments for many taxa have been made.
a high diversity of the focused animal groups the ground beetles. Finally, it was
necessary to sort out some interpretation of toponyms in the region, which were
incorrectly treated in the past (for example, some authors have erroneously cited
records or have referenced old collection material as being from Albania).
Map of Albania
MAP OF ALBANIA
mer the rainfall is just about 1/10 from the annual sum, and most plains and hilly parts
of the country are characterized with heavy summer droughts. The days with average
temperature more than 10 C are about 300 (in the seacoast plains) and favor the development of Mediterranean flora and fauna. The evergreen sclerophilous Mediterranean
forests are developed in the western part of Albania up to altitude of 400-500 m and in
some other regions higher. These forests consist of maquises (mosaically distributed
in the country) where Olea sp., Quercus coccifera, Q. ilex, other Quercus spp., Laurus nobilis,
Phyllirea angustifolia, Pistacia lentiskus, Juniperus oxicedrus and other dominate. The following
higher zone is occupied by xeric oak-hornbeam forests. The two zones are ones mostly
suffering from human activities in many places those forests are destroyed and, in are
replaced by farmlands or secondary xeric formations including mostly Paliurus spinacristi. The belt of the mesophilous oak-hornbeam and beech forests is developed at
altitude from 900-1000 up to 1600-1800 m a.s.l. This vegetation is more abundant in the
northern parts of Albania, where the mesophilous forests form the upper forest border. Normally in Northern Albania the coniferous communities are above the beech
forests, but elsewhere inversions in the distribution of the deciduous and coniferous
forests, owing to local humidity conditions, may occur. The orophytic (subalpine and
alpine) grass belt starts from 1900-2000 m a.s.l. It may be noted that the intrazonal
riverside communities consist mainly of Platanus orientalis.
Treatment of taxa
The arrangement of the taxa followed the order used in the recent Palaearctic Catalogue (Lbl & Smetana, 2003), with only two exceptions: genus Sinechostictus (Bembidiini) is regarded as separate genus, according to the recent studies on the phylogeny
of Trechitae (Grebennikov, 1997; Grebennikov & Maddison, 2005), and the arrangement of Tachyini follows Sciaky & Vigna Taglianti (2003). Synonyms are included if
they consider taxa from Albania.
and A. Winneguth, A. Penther and H. Zerny (most part), H. Priesner (most part), L.
Mader, C. Lona, J. Mller. F. Hieke, G. Mller-Motzfeld and L. Behne (Hieke et al.,
1986), as well as the materials collected in the second half of XX century by H. Peks
and S. Peks. The second important depository is that in the Naturhistorisches Museum
Wien, Vienna, Austria. From aspect of the species diversity the materials from Albania
there are comparable with these in Berlin. The most significant ground beetle collections in Vienna are these of V. Apfelbeck and his laboratory assistant A. Winneguth, E.
Csiki, A. Penther and H. Zerny, H. Priesner, L. Mader (most part), A. Bischoff and A.
Winkler (most part). Additionally, small parts of the collections of Matzenauer, C. Ravasini and C. Lona, J. Mller, L. Weirather and J. Meschnigg are also housed in Vienna.
Another place of important carabid collection is the Hungarian Natural History
Museum, Budapest. The most important collections here are these of V. Apfelbeck
and A. Winneguth, E. Csiki (most part) and Xh. Murraj. Other small collections housed
in Budapest are those of M. Grabowski, L. Mader, C. Ravasini and C. Lona, A. Winkler and A. Bischoff. Recently, Fehr et al. (2004) also collected material of Carabidae
from Albania that is currently in process of preparation.
An important place as depository of type material from the Balkans is the collection of the Deutsches Entomologisches Institut in Mncheberg, Germany; some of
the types deposited there have been reviewed. Materials from one American and two
Balkan museums have been examined too. The most substantial collection among
three is that housed in the National Museum of Natural History in Sofia. It has been
collected by Bulgarian expeditions to Albania after 1992 and at present it is containing about 160 species. The other two collections are: i/ the one in the Museum of
Natural Sciences, Tiran (Albania), which in part was identified and published by
Murraj (1968), and ii/ that in the Carnegie Museum of Natural History, Pittsburgh
(USA), consisting of recently collected material of 57 species.1
Acknowledgments
The most important part of the material was examined in the course of the authors
work under the European Union-funded Integrated Infrastructure Initiative Synthesys, which is actually the prime cause for the completion of this work (Applications DE-TAF-725, AT-TAF-758 and HU-TAF-817). For the work with the collec1
Except the aforementioned collections, there are other four European museums that may house more
or less large number of Carabidae from Albania. At the present time a significant part of the former
collection of V. Apfelbeck is preserving in the Zemaljski muzej Bosne i Hercegovine in Sarajevo
(Bosnia Herzegovina). The Museo Civico di Storia Naturale in Trieste (Italy) and the Museo Civico di
Storia Naturale in Milano (Italy) house the most part of the materials collected by the Italian expeditions
in Albania before the Second World War. The first one is remarkable with the collections of C. Ravasini,
C. Lona and J. Mller, and the second one - with that of A. Schatzmayr. Most probably, the collection
of the Narodni Museum in Prague (Czech Republic) also contains materials from Albania.
ACKNOWLEDGEMENTS
11
tions and the materials on loan I am indebted to Prof. V. Andoni (MNST), Dr. P.
Beron (NMNHS), Dr. R. Booth (BMNH), Mr. R. Davidson (CMNH), Mr. B. Jaeger
(ZMHU), Dr. A. Popov (NMNHS), Dr. H. Schnmann (NMW), Dr. Gy. Szel
(HNHM), Dr. M. Uglih (ZMHU) and Dr. L. Zerche (DEI).
With pleasure, I record my deep appreciation to the colleagues who contributed
this study giving their unpublished data (marked with the paragraphs Material included or Additional material included): in the first place Prof. Dr. A. Vigna Taglianti
(Universit di RomaLa Sapienza, Roma, Italy) as well as Mr. P. Bulirsch (Prague,
Czech Republic), Dr. F. Hieke (ZMHU), Mr. S. Hristovski (Institute of Biology, Skopje, Republic of Macedonia), Mr. B. Jaeger (ZMHU), Prof. Xh. Murraj (Tiran, Albania) and Mr. D.W. Wrase (Berlin, Germany). It is worth mentioning that, Prof. Dr. A.
Vigna Taglianti told me that several years ago he began the preparation of a checklist
of the Carabidae of Albania, being still not completed. Generously, he entrusted to
me the draft of this work which included a moderate number of records collected
from the Italian expeditions to Albania carried out during the last 15 years. Dr. F.
Hieke made valuable comments and improvements in the part about genus Amara, as
well as kindly provided me the data of both unavailable papers and papers in press.
The following colleagues also revised and improved different parts of the catalogue:
Dr. P. Bonavita (Bembidiini; Universit di RomaLa Sapienza, Roma, Italy), Mr. P.
Bulirsch (Scaritinae), Mr. J. Gebert (Cicindelinae; Rohne, Germany), Dr. P.M. Giachino (Speluncarius, Tapinopterus; Settore Fitosanitario Regionale, Torino, Italy), Mr. B.
Jaeger (Stenolophina), Mr. P. Neri (Bembidiini; Forl, Italy), Dr. B. Kataev (Harpalini;
ZISP), Mr. L. Picciau (Speluncarius, Tapinopterus; Museo Regionale de Scienze Naturali,
Torino), Dr. L. Toledano (Bembidiini; Museo Civico di Storia Naturale, Verona, Italy), Prof. Dr. A. Vigna Taglianti (whole draft, and especially Bembidiini) and Mr. D.W.
Wrase (whole draft, and especially Harpalini).
For the determination of material, I thank Dr. D. Fedorenko (Dyschirius; Moscow
Pedagogical State University, Moscow, Russia), Dr. F. Hieke (Amara), Dr. B. Kataev
(Anisodactylus, Harpalus, Ophonus, Trichotichnus, Acupalpus, Bradycellus), both Mr. G. Ledoux
and Mr. Ph. Roux (Nebria; Paris, France), Mr. P. Neri (Bembidion), Mr. M. Pavesi (Syntomus, Olisthopus; MSNM, Italy) and Mr. D.W. Wrase (Notiophilus, Harpalus, Ophonus,
Parophonus, Microlestes).
Special thanks are due to Prof. Dr. A. Vigna Taglianti, Mr. D.W. Wrase, and Mr. I.
Stoyanov (Central Laboratory of General Ecology, Sofia, Bulgaria), who made careful reviews and proposed important changes of the earlier versions of the manuscript on which this paper is based. Willingly, again Mr. I. Stoyanov helped me with
the revision of the English text. Prof. Dr. A. Casale (Universit di Sassari, Sassari)
kindly allowed me the reproduction of his drawing of Abanotrechus beroni, and Dr. P.
Stoev (NMNHS) gave me the basis of the map.
I owe deep gratitude to my late farther for his strong and enthusiastic support in the
initial stages of the work, as well as to my mother and my wife for their tolerance.
12
Abbreviations
I. Abbreviations of collections studied upon the European Union-funded Integrated Infrastructure Initiative Synthesys by the author
DEI
Deutsches Entomologisches Institut, Mncheberg, Germany
HNHM Hungarian Natural History Museum (Magyar Termszettudomnyi Mzeum),
Budapest, Hungary
MNHUB Museum fr Naturkunde der Humboldt Universitt zu Berlin, Bereich Zoologisches Museum, Berlin, Germany
NMW
Naturhistorisches Museum Wien, Vienna, Austria
II. Abbrevuiations of other collections studied by the author
BMNH The Natural History Museum, London, United Kingdom
CMNH Carnegie Museum of Natural History, Pittsburgh, USA
MNST Museum of Natural Sciences, Tirana, Albania
NMNHS National Museum of Natural History, Sofia, Bulgaria
III. Abbreviations of collections studied by other colleagues
MCG
Museo Civico di Storia Naturale Giacomo Doria, Genoa, Italy
MHNG Musum dHistoire Naturelle, Geneva, Switzerland
MNHP Musum National dHistoire Naturelle, Paris, France
MSNM Museo Civico di Storia Naturale di Milano, Milano, Italy
MTKD Museum fr Tierkunde, Dresden, Germany
NMPC National Museum Prague (Nrodni muzeum v Praze), Prague, Czech Republic
ZISP
Zoological Institute, St. Petersburg, Russia
ZSM
Zoologische Staatssammlung, Munich, Germany
cAVT
private collection A. Vigna Taglianti, Roma, Italy
cBJ
private collection B. Jaeger, Berlin, Germany
cFC
private collection F. Cassola, Rome, Italy
cLR
private collection G. Ledoux & P. Roux, Paris, France
cPB
private collection P. Bulirsch, Prague, Czech Republic
cPN
private collection P. Neri, Forl, Italy
cSH
private collection S. Hristovski, Skopje, Macedonia
cWH
private collection W. Heinz, Schwanfeld, Germany
cWR
private collection D.W. Wrase, Berlin, Germany
cXM
private collection Xh. Murraj, Tiran, Albania
ABBREVIATIONS
13
IV. Other abbreviations and symbols used in the systematic part of the catalogue
! taxa still not well defined
? taxa with doubtful or questionable occurrence in Albania
[ ] taxa that most probably inhabit Albania; they were recorded from neighbouring
region
# taxa incorrectly indicated for Albania, and therefore excluded from the catalogue
f. female specimen/s
m. male specimens/s
s. specimen/s
15
SYSTEMATIC PART
Family CARABIDAE Latreille, 1802
Subfamily NEBRIINAE Laporte, 1834
Tribe NEBRIINI Laporte, 1834
1. Genus Eurynebria Ganglbauer, 1891
Notes. New genus for the fauna of Albania.
1.1. Eurynebria complanata (Linnaeus, 1767)
Material studied. Apfelb. Albanien, 4 s. (MNHUB); Albania: Valona. / complanata orientalis Typus!
Apfb., 1 s. (HNHM); Valona. Albania / Typus! Apf., 1 s. (HNHM); Valona / v. albanica
Cotypus! Apfb, 4 s. (HNHM); Sdalbanien Kopstein / Eurynebria complanata var. albanica Apf., 2
s. (NMW); Alban. Wght. Valona / Eurynebria complanata var. albanica Apf., 8 s. (NMW); Alban.
Wght. Valona, 4 s. (MNHUB); Albanien Avlona 08 / E. complanata albanica Coll. Apfelbeck, 4 s.
(HNHM); Valona Albanien V.1908 H. Hopp, 2 s. (MNHUB); Ruskuli Alb., Mader / Eurynebria
complanata var. albanica Apf., 2 s. (NMW); Shng 25.X.1965 kol. Murraj, 2 s. (MNHUB).
Additional material included (studied A. Vigna Taglianti). Shn Pjeter, Durrs District,
29.V.2004, 1 f., A. Paparistos leg. (cAVT).
Notes. Eurynebria complanata orientalis Apfelbeck, E. complanata albanica and E. complanata var.
albanica Apfelbeck are nomen nudum. The species was recorded hitherto only for European
Turkey on the Balkans (Farka & Janata, 2003: 78). New species for the fauna of Albania.
17
19
21
paratypes of Nebria aetolica tomoricensis (NMW); Tomori Toik Alban. mer. / Winkler, Lona,
Bischoff, V.1931, 1 s. (NMW); Albania leg. Bischoff 1932 / Tojk Tomor, 10 paratypes of
Nebria aetolica tomoricensis (MNHUB); Albania 1936 leg. Ramadan coll. Bischoff / Maja e
Papingut, 1 holotype and 26 paratypes of Nebria aetolica papingutica (HNHM; MNHUB);
Albania leg. Bischoff 1937 / Maja e Policanit, 2 paratypes of Nebria aetolica papingutica
(HNHM); Albania leg. Bischoff 1937 / Mal i That, 2 s. (HNHM).
[Nebria macedonica rambouseki Maan, 1938]
Notes. This geographical race was described from the Macedonian part of the Jablanitsa Mt.
(Maan, 1938: 57; Maan, 1945: 56). So far there is no sure record from Albania.
Subgenus Boreonebria Jeannel, 1937
3.6. Nebria rufescens (Strm, 1768)
3.6.1. Nebria rufescens rufescens (Strm, 1768) [= gyllenhali (Schnherr, 1806)]
References. Nebria gyllenhali: Hieke & Wrase, 1988: 22. 12. Nebria rufescens rufescens: Farka &
Janata, 2003: 88.
Material studied. Albania leg. Bischoff 1935 / Shklzen, 4 s. (MNHUB).
Notes. First detailed record for Albania.
Subgenus Eunebria Jeannel, 1937
3.7. Nebria jockischi Sturm, 1815
3.7.1. Nebria jockischi jockischi Sturm, 1815
References. Nebria jockischi: Hieke & Wrase, 1988: 22.
Material studied. Albania leg. Bischoff 1936 / Gjallica, 3 s. (MNHUB); Albania leg.
Bischoff 1936 / Korab, 8 s. (MNHUB).
Notes. Farka & Janata (2003: 90) have omitted N. jockischi for Albania. The species is here
confirmed for the country with first detailed records.
Subgenus Nebria s.str.
3.8. Nebria brevicollis (Fabricius, 1792)
References. Nebria brevicollis: Apfelbeck, 1904: 54 (Velipoja; Vlor); Apfelbeck, 1909: 297
(Vlor); Schatzmayr, 1943b: 95 (Mal i Krujes; Lushnje; Stan-Karbunar; Berat); Hieke & Wrase,
1988: 22; Farka & Janata, 2003: 92. Nebria brevicollis var. salina: Apfelbeck, 1909: 297 (Vlor).
Material studied. Albania Cerrik / 18.IV.1958 leg. Xh. Murraj, 5 s. (HNHM); Albania
Sauk / 2.V.1958 leg. Xh. Murraj, 3 s. (HNHM); Albania Dajti / 2.6.1959 leg. Xh. Murraj,
1 s. (HNHM); Tiran, 10.V.1961, 1 s. (MNST); Mal i Dajtit Mt., 10-11.V.1965, 3 s. (MNST);
Tiran Park, 8.III.1998, 54 s., shaded lake edge under leaves and wet, rotten wood, R. Ward
leg. (CMNH; NMNHS); Albania Tomor Mts. Kajala e Tomorit, 26.V.2004, K. Harmos & D.
Murnya, 1 s. (HNHM).
22
Additional material included (studied A. Vigna Taglianti). Tiran, Mal i Dajtit Mt.,
7.VI.2002, 1 f., A. Paparistos leg. (cAVT); Qesarat, near Memaliaj (Tepelen District), valley
of Vijos River, 300-360 m, 18.IV.2003, 3 m., 2 f., P. Crucitti & C. Cavalieri leg. (cAVT);
Humelic (between Tepelen and Gjirokastr), 350 m, 23.IV.2003, 1 m., 2 f., P. Crucitti & C.
Cavalier leg. (cAVT).
Notes. Except the material studied above I have seen additionally more than 20 s. from different
localities in MNHUB.
3.9. Nebria kratteri Dejean, 1831
3.9.1. Nebria kratteri kratteri Dejean, 1831 [= valonensis Apfelbeck, 1904]
References. Nebria kratteri: Apfelbeck, 1904: 52 (South Albania); Apfelbeck, 1907a: 498, 501,
503 (Mal i Shenjt Mt.: top region, 1500 m); Schatzmayr, 1928: 46; Holdhaus, 1956: 7 (Mal i
Dajtit Mt.); Magistretti, 1965: 79; Ledoux & Roux, 1990: 94. Nebria kratteri valonensis: Apfelbeck,
1904: 55 (Vlor); Bnninger, 1925: 274; Maan, 1939a: 19 (Vlor); Jeanne, 1974: 66. Nebria
kratteri ssp.: Schatzmayr, 1943b: 94 (cave Stere Gropa e Vasos near Dukati). Nebria kratteri
kratteri: Farka & Janata, 2003: 92.
Material studied. Lumi Dukati Alban. mer. / Winkler, Lona, Bischoff, V.1931, 1 s. (NMW);
Mal-i-Dajtit Tiran / Albania Bischoff 1931, 2 s. (DEI); Tomor: Kulmak Albania mer.
/ Winkler, Lona, Bischoff, V.1931, 5 s. (NMW); Logara juni / Albania Bischoff 1932, 1
s. (DEI); Tomor-Buranj juli 1932 / Albania Bischoff 1932, 1 s. (DEI).
Notes. The record Chani Driskos bei Jannina (Apfelbeck, 1904: 55) refers to Northwestern
Greece. The locality at Mal i Shenit Mt. (Apfelbeck, 1907a), representing the northernmost
known spot in the species range, needs verification. In MNHUB, I have seen more than 100
specimens from different localities, which remained unregistered.
# Nebria salina Fairmaire & Laboulbne, 1854
References. Nebria brevicollis var. salina: Apfelbeck, 1909: 297 (Vlor).
Notes. This Atlantic species does not inhabit the Balkan Peninsula, so that the record for it
referred to N. brevicollis.
23
24
25
Additional material included (studied F. Cassola). Rapsh, 15.V.2002, 1 s., A. Paparistos leg.
(cFC); Qesarat, near Memaliaj (Tepelen District), valley of Vijos River, 300-360 m, 18.IV.2003,
1 m., 1 f., P. Crucitti & C. Cavalieri leg. (cFC); Vasjar (Tepelen District), 18.IV.2003, 1 m., P.
Crucitti & C. Cavalieri leg. (cFC).
Notes. By the lack of modern study, the intraspecific relations of C. campestris s.l. in the Balkan
Peninsula are obscure. For this reason all the data about it from Albania are referred to ssp.
olivieria provisionally. The ssp. pontica Fischer von Waldheim, 1828 inhabits only the Black Sea
Coast, so that the record for this race is referred to ssp. olivieria. Except the material studied
above I have also seen additional specimens from Gjallica Lums and the Korab Mt. in HNHM.
# Cicindela hybrida Linnaeus, 1758
References. Cicindela hybrida: Apfelbeck, 1904: 5 (Vlor); Csiki, 1940: 209 (Kula Ljums); Hieke
& Wrase, 1988: 10.
Notes. I follow the view of Gebert (1995) and refer the literature records for C. hybrida to C.
monticola albanica.
6.2. Cicindela monticola Menetries, 1832
6.2.1. Cicindela monticola albanica Apfelbeck, 1909
References. Cicindela hybrida: Apfelbeck, 1904: 5 (Vlor); Csiki, 1940: 209 (Kula Ljums); Hieke
& Wrase, 1988: 10. Cicindela hybrida albanica: Apfelbeck, 1909: 297 (Vlor, loc. typ.; Berat, loc.
typ.); Apfelbeck, 1917: 219 (Kiri River near Mesi, loc. typ.; Merdita: Munella, loc. typ.; Vlor,
loc. typ.); Apfelbeck, 1918a: 75 (Shkodr, loc. typ.; Orish, loc. typ.; Vlor, loc. typ.); Apfelbeck,
1930: 161; Murraj, 1968: 33 (Golem; Suhagora; Tomor Mt.; Shngjin); Cassola, 1973: 26. Cicindela
sahlbergi albanica: Gebert, 1995: 16 (Buen River near Shkorr; Mesi; Vlor; Merdita: Munella;
Berat). Cicindela monticola albanica: Puchkov & Matalin, 2003: 106.
Material studied. Skutari Bojana / C. v. albanica Apfb. Coll. Apfelbeck, 2 s. (HNHM);
Albania Kula Ljums / Akad. Balk. Exp. Csiki 1917, 1 s. (HNHM); Albania coll. Lichtn.,
1 s. (HNHM); Shkumbi - Kavajes, 7.V.1958, 1 f. (MNST); Golem, 24.III.1961, 2 m., 3 f.
(MNST); Divjake, 18.VI.1964, 1 f. (MNST); Tepelen District, Vijos River, 2.VI.1994, 1 s., G.
Blagoev leg. (NMNHS); Radona, Shkiela, 21.VI.1994, 2 s., B. Guorguiev leg. (NMNHS);
Leskovac - Barmachi, 880 m, 2.VI.1995, 1 s., G. Blagoev leg. (NMNHS).
Notes. Apfelbeck published four consecutive descriptions of the ssp. albanica (Apfelbeck, 1909;
Apfelbeck, 1917; Apfelbeck, 1918a; Apfelbeck, 1930: 161), the oldest one is correctly treated as valid.
6.3. Cicindela sylvatica Linnaeus, 1758
6.3.1. Cicindela sylvatica sylvatica Linnaeus, 1758
References. Cicindela sylvatica sylvatica: Puchkov & Matalin, 2003: 107.
6.4. Cicindela transversalis Dejean, 1822
6.4.1. Cicindela transversalis transversalis Dejean, 1822 [= riparia Dejean, 1822]
References. Cicindela hybrida riparia: Apfelbeck, 1907a: 496 (Kiri River near Mesi).
Notes. Puchkov & Matalin (2003: 108) have omitted the species for Albania, therefore its
occurrence in Albania needs further confirmation.
COLEOPTERA: CARABIDAE: CICINDELINAE
27
28
29
Additional material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollats Mt.,
1900 m, 10.VII.1993, 1 m., L. Latella leg. (cAVT).
# subgenus Eucarabus Ghin, 1876
# Carabus catenulatus Scopoli, 1763
# Carabus catenulatus catenulatus Scopoli, 1763 [= catenatus Panzer, 1803; albanensis Ghin, 1885]
References. Carabus catenatus var. albanensis: Apfelbeck, 1904: 35.
Notes. Probably, the original record for this species from Albania is that in the description of
albanensis Ghin, 1885. Apfelbeck (1904: 35) has based indirectly on this record, indicating no
exact locality for the county. I cannot find the original description of this variation, and believe
the species does not inhabit Albania (see also Turin et al., 2003).
[subgenus Hygrocarabus C.G. Thomson, 1875]
[Carabus variolosus Fabricius, 1787]
[Carabus variolosus nodulosus Creutzer, 1799]
References. Carabus variolosus nodulosus: Breuning, 1926: 19; Breuning, 1928a: 113; Buresch &
Kantardjieva, 1928: 88. Carabus nodulosus: Turin et al., 2003: 240.
Notes. Actually, the record Albanien: Ljuboten (Breuning, 1926: 22) concerns the territory
of Republic of Macedonia (see also Breuning, 1935: 1169). The following authors had followed
this data mentioning the species for Albania. In spite of this, C. variolosus may inhabit the
marginal northern or northeastern regions of the country.
[subgenus Limnocarabus Ghin, 1876]
[Carabus clathratus Linnaeus, 1761]
[Carabus clathratus auraniensis J. Mller, 1903]
Notes. I cannot find any exact record for this species from Albania even though Turin et al.
(2003: 155) showed in the map it occurs there. This rare Carabus-species may inhabit the
country as Purkyn (1928) described the variation epirensis Purkyn, 1928 (synonym of C.
clathratus auraniensis) from the Greek Epirus.
Subgenus Megodontus Solier, 1848
10.4. Carabus caelatus Fabricius, 1801
10.4.1. Carabus caelatus caelatus Fabricius, 1801 [= sarajevoensis Apfelbeck, 1890; malissorum
Apfelbeck, 1919; metalkanus Apfelbeck, 1919]
References. Carabus caelatus sarajevensis sic!: Apfelbeck, 1907a: 498, 502 (Mal i Shenjt Mt.: top
region, 1500 m). Carabus caelatus: Apfelbeck, 1907a: 500; Apfelbeck, 1917: 220 (Merdita);
Apfelbeck, 1930: 162 (Merdita); Horion, 1941: 41; Bezina, 1999: 73. Carabus caelatus malissorum:
Apfelbeck, 1919a: 226-267 (North Albanian Alps, Maja e Jezerz, loc. typ.); Apfelbeck, 1919b:
45-46 (North Albanian Alps, Maja e Jezerz, loc. typ.). Carabus caelatus caelatus n. metalkanus:
Breuning, 1935: 1297 (Merdita; Gjalic e Lumes; Sarisaltik near Kruja; Male Dojtitz; Mal e
Krujes); Breuning, 1978: 258. Carabus caelatus caelatus m. malissorum: Breuning, 1935: 1298 (North
Albanian Alps; Maja e Jezerc); Breuning, 1978: 258. Carabus caelatus metalkanus: Arnold, 1939:
189. Carabus caelatus metalkanus v. malissorum: Arnold, 1939: 189. Carabus caelatus caelatus: Bezina,
1999: 73; Bousquet et al., 2003: 163. Carabus caelatus sarajevoensis: Turin et al., 2003: 253.
COLEOPTERA: CARABIDAE: CARABINAE
31
Material studied. North Albanian Alps, Bog, 1400-1800 m, 1.VI.1993, 2 m., P. Beron & B.
Petrov leg. (NMNHS); North Albanian Alps, Bog, 1000-1100 m, 5-9.VI.1993, 3 m., P. Beron
& B. Petrov leg. (NMNHS).
Additional material included (studied A. Vigna Taglianti). Theth, VII.1993, 1 m., 1 f., P.
Dioli leg. (cAVT); Tropoia, valley of Valbona River, 900 m, 5.VII.1993, 1 f., L. Latella leg.
(cAVT); Valbona, Kollats Mt., 1800 m, 9.VII.1993, 1 f., F. Sirtori leg. (cAVT).
Notes. Apfelbeck published two description of the form malissorum (Apfelbeck, 1919;
Apfelbeck, 1920), the older one is correctly treated as valid.
10.4.2. Carabus caelatus dalmatinus Duftschmid, 1812 [= procerus Reitter, 1885]
References. Carabus caelatus dalmatinus: Apfelbeck, 1904: 25 (Velipoja near Lezh); Wohlberedt,
1909: 127 (near Liqen i Shkodrs); Bezina, 1999: 74; Bousquet et al., 2003: 163; Turin et al.,
2003: 253 (Velipoja; Lezh). Carabus dalmatinus procerus: Apfelbeck, 1918c: 160 (North Albanian
Alps). Carabus caelatus var. procerus: Apfelbeck, 1918c: 163 (North Albanian Alps, 1300-2000 m).
Carabus caelatus dalmatinus n. procerus: Breuning, 1935: 1300 (Lezh District); Breuning, 1978: 258.
Material included (studied A. Vigna Taglianti). Durrs, 13,VI.1998, 1 f., A. Paparistos leg. (cAVT).
Notes. The mentioning of the locality Vilajet (Turin et al., 2003: 253) is misunderstanding.
Actually this Turkish word is an equivalent to the English one district.
10.5. Carabus croaticus Dejean, 1826
10.5.1. Carabus croaticus durmitorensis Apfelbeck, 1904 [= ljubetensis Apfelbeck, 1917; babinjensis
Apfelbeck, 1919]
References. Carabus croaticus babinjensis: Apfelbeck, 1919a: 226 (North Albanian Alps, loc. typ.);
Apfelbeck, 1919b: 45 (North Albanian Alps, loc. typ.); Krtschmer & Drovenik, 1977: 215 (Abb.1);
Turin et al., 2003: 254 (Babinje Planina Mt.). Carabus croaticus ljubetensis n. ljubetensis: Breuning,
1935: 1291 (North Albanian Alps; Babinje Planina Mt.); Breuning, 1978: 257. Carabus croaticus
ljubetensis m. durmitorensis: Breuning, 1935: 1291 (Korab Mt.); Breuning, 1978: 257. Carabus croaticus:
Horion, 1941: 41; Tabakovi-Toi, 1988: 135; Bezina, 1999: 74. Carabus croaticus durmitorensis:
Bezina, 1999: 74 (North Albanian Alps); Bousquet et al., 2003: 163. Carabus croaticus durmitorensis
var. babinjensis: Bezina, 1999: 74 (North Albanian Alps). Carabus croaticus durmitorensis var. ljubetensis:
Bezina, 1999: 74. Carabus croaticus ljubetensis: Turin et al., 2003: 254.
Notes. Apfelbeck published three consecutive descriptions of Carabus croaticus ljubetensis
(Apfelbeck, 1917; Apfelbeck, 1918a; Apfelbeck, 1930), the second one has long time been
accredited to be correct. However it is wrong and the original description must be assigned to
the oldest one. The same author gave also two descriptions of Carabus croaticus babinjensis (Apfelbeck,
1919a; Apfelbeck, 1919b), as for the present it is not clear which of both has priority.
10.6. Carabus violaceus Linnaeus, 1758
10.6.1. Carabus violaceus azurescens Dejean, 1826 [= rilvensis Kolbe, 1887; zabljakensis Eidam,
1927; shljebensis Mandl, 1985]
References. Carabus violaceus azurescens n. zabljakensis: Breuning, 1935: 1261 (Maranajt Mt.; Merdita);
Breuning, 1978: 254. Carabus violaceus azurescens m. zabljakensis: trba, 1945: 152 (Maranajt Mt.;
Merdita). Carabus violaceus: Hieke & Wrase, 1988: 15. Carabus violaceus azurescens var. shljebensis:
Bezina, 1999: 77 (North Albanian Alps). Carabus violaceus azurescens: Turin et al., 2003: 260.
32
10.6.2. Carabus violaceus dryas Gistl, 1857 [= merditanus Apfelbeck, 1917; shardagensis Apfelbeck,
1917; korabensis Csiki, 1944]
References. Carabus violaceus merditanus: Apfelbeck, 1917: 220 (Merdita: Zep Mt. near Fandi,
loc. typ.); Apfelbeck, 1918a: 75-76 (Merdita: Zep Mt., loc. typ.); Apfelbeck, 1930: 162 (Merdita:
Zep Mt., loc. typ.); Cleu, 1968: 28 (Merdita); Cleu, 1969: 167. Carabus violaceus azurescens m.
merditana: Breuning, 1935: 1262 (Vermosh; Zep Mt. near Oroshi; Peshtrik Mt.; Gjalic e
Lumes); Breuning, 1978: 254 (Zep Mt. near Oroshi). Carabus violaceus zabljakensis v. merditanus:
Arnold, 1939: 186. Carabus violaceus var. shardagensis: Csiki, 1940: 210 (Gjalic Mt.; Korab Mt.).
Carabus violaceus var. korabensis: Csiki, 1944: 50-51 (Korab Mt., loc. typ.). Carabus violaceus azurescens
m. merditanus: trba, 1945: 152 (Vermosh; Zeb Mt.). Carabus violaceus shardagensis: Cleu, 1968:
28 (Korab Mt.); Cleu, 1969: 167 (Korab Mt.). Carabus picenus peristericus: Tarrier, 1974: 242
(Korab Mt.). Carabus violaceus dryas: Bezina, 1999: 77; Bousquet et al., 2003: 165; Turin et al.,
2003: 260. Carabus violaceus dryas var. merditanus: Bezina, 1999: 77 (Merdita: Zep Mt.; Gjalic
e Lumes). Carabus violaceus dryas var. korabensis: Bezina, 1999: 77 (Korab Mt.).
Material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollats Mt., 2250 m,
11.VII.1993, 1 m., L. Latella leg. (cAVT); Tropoia, Valbona, Kollats Mt., 2000 m, 14.VII.1993,
rests of 1 s., A. Gobetti leg. (cAVT).
Notes. Apfelbeck published three descriptions of both C. violaceus merditanus and C. violaceus
shardagensis, respectively in Serbo-Croatian, in Latin and in German (Apfelbeck, 1917; Apfelbeck,
1918a; Apfelbeck, 1930), the second ones have long time been accredited to be correct. However
it is incorrect and the original descriptions must be assigned to the oldest ones. A revision of
the type of the var. shardagensis Apfelbeck, 1917 is needed to elucidate to which subspecies,
azurescens or dryas, it belongs. Csiki (1944: 50, Fig. 1a) represented a figure of this variation
which let me assume that it belongs rather to the former subspecies.
# Carabus violaceus exasperatus Duftschmid, 1812 [= vlasuljensis Apfelbeck, 1894]
References. Carabus violaceus vlasuljensis: Apfelbeck, 1907a: 502 (Merdita); Apfelbeck, 1918c:
160, 163 (North Albanian Alps, Vermosh).
Notes. The easternmost known records of ssp. exasperatus come from Bosnia Herzegovina, so that
the literature data for it should be referred to some of both subspecies that really live in Albania.
Subgenus Oreocarabus Ghin, 1876
Notes. In my opinion, C. hortensis, C. neumeyeri and C. preslii represent separate species by
reason of its clear and constant differences in both the overall aspect of the pronotum and
the penis (Casale et al., 1982: 185, 190 fig. 75, 195). Externally, the first species can be easily
distinguished from the other two in the more quadrate aspect of the pronotum (vs. subovate
or trapezium-shaped), the lateral border of pronotum posteriorly sinuate (vs. straight to slightly
sinuate) and in the lateral lobes of pronotum clearly horizontal or slightly oblique (vs. oblique).
Additionally C. neumeyeri has lateral borders of pronotum posteriorly more convergent than
those in C. hortensis. The following nomenclatorial acts are proposed: Carabus (Oreocarabus)
neumeyeri Schaum, 1856, stat. nov. for Carabus (Oreocarabus) hortensis neumeyeri Schaum, 1856;
Carabus (Oreocarabus) preslii Dejean, 1830, stat. rest. for Carabus (Oreocarabus) hortensis preslii
Dejean, 1830 (see Casale et al., 1982: 194).
33
34
35
et al., 1986: 183 (Mal i Dajtit Mt.: Shkall Prisk); Bezina, 1999: 62; Bousquet et al., 2003: 187;
Turin et al., 2003: 278. Carabus coriaceus excavatus dalmaticus: Schatzmayr, 1943b: 94 (Lushnje;
Karbunara e Madhe; Dukati, south of Vlor). Carabus coriaceus joniensis: Bezina, 1999: 62;
Bousquet et al., 2003: 187. Carabus coriaceus joniensis: var. albanicus: Bezina, 1999: 62 (Maja e
Tartarit Mt.). Carabus coriaceus mediterraneus: Turin et al., 2003: 278.
Material studied. Sauk, 18.II.1960, 1 m. (MNST); Ib, 3.VII.1962, 1 f. (MNST); Biz,
27.VII.1966, 1 f. (MNST); Tiran, 27.III.1968, 1 f. (MNST); Tiran Park, 22-31.X.1997, 2 s.,
R. Ward leg. (CMNH); Tiran Park, 1-4.I.1998, 2 s., R. Ward leg. (CMNH).
Additional material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollats Mt.,
2250 m, 11.VII.1993, rests of 1 s., L. Latella leg. (cAVT); Tropoia, Valbona, Kollats Mt., 2000
m, 14.VII.1993, 1 f., A. Gobetti leg. (cAVT); Qesarat near Memaliaj (Tepelen District), valley
of Vijos River, 300-360 m, 18.IV.2003, 2 m., P. Crucitti & C. Cavalieri leg. (cAVT).
Note: According to Turin et al. (2003) C. coriaceus mediterraneus Born, 1906 has transionian
distribution and lives in Albania.
Subgenus Tachypus Weber, 1801
10.14. Carabus cancellatus Illiger, 1798
10.14.1. Carabus cancellatus alessiensis Apfelbeck, 1901 [= hypsobius Apfelbeck, 1918]
References. Carabus cancellatus var. alessiensis: Apfelbeck, 1901b: 121 (Lezh, loc. typ.). Carabus cancellatus
alessiensis: Apfelbeck, 1904: 30; 33 (Velipoja near Lezh); Apfelbeck, 1907a: 496 (Buen River Outfall);
Breuning, 1933: 594 (Velipoja near Lezh; Shkodr; Pulaj); Arnold, 1939: 161 (North Albania);
Breuning, 1978: 187 (Velipoja); Bezina, 1999: 41 (Velipoja near Lezh); Bousquet et al., 2003: 195;
Turin et al., 2003: 194. Carabus cancellatus subgraniger var. alessiensis: Bernau, 1913: 198 (Lezh). Carabus
cancellatus emarginatus: Bernau, 1913: 200. Carabus cancellatus: Hieke & Wrase, 1988: 17.
Material studied. Shkodr, 14.V.1963, 2 m. (MNST).
Notes. The type locality of the form hypsobius Apfelbeck, 1918c: 163 is Vunaj (in Albanian)
or Vusanje (in Serbo-Croatian) in the North Albanian Alps. It is situated in the Montenegrin
territory very closely to Albania. C. cancellatus emarginatus Duftschmid, 1812 is a valid subspecies
reaching to Bosnia Herzegovina in the east.
Subgenus Tomocarabus Reitter, 1896
10.15. Carabus convexus Fabricius, 1775
10.15.1. Carabus convexus dilatatus Dejean, 1826 [= chionophilus Apfelbeck, 1904; miliusi Breuning, 1927]
References. Carabus convexus var.: Apfelbeck, 1907a: 498 (Mal i Shenjt Mt.: top region, 1500
m). Carabus convexus dilatatus: Breuning, 1933: 874 (Vlor; Gjirokastr); Bousquet et al., 2003:
198; Turin et al., 2003: 230. Carabus convexus dilatatus n. miliusi: Breuning, 1933: 876 (North
Albanian Alps; Merdita); Breuning, 1978: 214. Carabus convexus miliusi: Arnold, 1939: 178. Carabus
convexus var. chionophilus: Csiki, 1940: 210 (Korab Mt.). Carabus convexus: Hieke & Wrase, 1988:
19. Carabus convexus dilatatus var. miliusi: Bezina, 1999: 33.
Material studied. Mal i Dajtit Mt., 11.V.1965, 1 m. (MNST); North Albanian Alps, Maje
Radohimes, 1900-2200, 28.V.1993, 1 m., 1 f., P. Beron & B. Petrov leg. (NMNHS); North Albanian
Alps, Maje Radohimes, 2400-2550 m, 29.V.1993, 1 m., 1 f., P. Beron & B. Petrov leg. (NMNHS);
36
North Albanian Alps, Bog, 1400-1800 m, 1.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS);
North Albanian Alps, Bog, 1800-1900 m, 20-23.VI.1993, 3 f., P. Beron & B. Petrov leg. (NMNHS).
? subgenus Trachycarabus Ghin, 1876
? Carabus scabriusculus Olivier, 1795
? Carabus scabriusculus bulgarus Lapouge, 1908
References. Carabus scabriusculus bulgarus: Turin et al., 1993: 42.
Notes. This species is questionable for Albania due to lack of exact records. On the other
side, it was quoted for the neighboring countries Montenegro and Serbia.
37
Notes. Winkler (1939) synonymized the ssp. albanicus with ssp. montenegrinus. Maan (1940c)
accepted this point of view, but Mandl (1989) treated both as separate subspecies. Samples of
the ssp. montenegrinus, ssp. albanicus, as well as the holotype and paratypes of the ssp. dinaricus
have been studied in NMW. Differences between specimens from different localities exist
mostly in the sizes of the body and shape of the pronotum, but they also have occurred in the
limits of one and the same population. In reality characters that can support well-defined
geographical races in the Western Balkans, excluding the well-differentiated ssp. travnikanus
Apfelbeck, 1904 (type series in NMW) and the not studied ssp. jakupicensis Maan, 1943, are
absent. Therefore the following synonymy is proposed: Cychrus semigranosus albanicus Hopp,
1929, syn. rest. of Cychrus semigranosus montenegrinus Apfelbeck, 1904; Cychrus semigranosus dinaricus
Mandl, 1989, syn. nov. of Cychrus semigranosus montenegrinus Apfelbeck, 1904. In MNHUB, I
have furthermore seen 69 s. from different localities, which remained unregistered.
Dragot / Albania leg. Bischoff 1936, 1 s. (MNHUB); Bojana Shkoder / Albania leg. Bischoff
1937, 1 s. (MNHUB); Tepelen District, Vijos River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS).
39
(Merdita: Zep Mt. near Fandi, loc. typ.); Casale & Vigna Taglianti, 1983: 29, 39, 41 (Merdita:
Zep Mt.; Shen Thanas Mt. near Vlor); Hrka, 1988: 293-294 (Mal i Shenjt Mt.; Zep Mt.;
Mal i Dajtit Mt.; Mal i Krujs Mt.; Shen Ili; Maja e Tartarit Mt.; Cafa Kjors; Tomor Mt.; Shen
Thanas Mt.; Llogora); Wrase, 1991: 18. Aptinus merditanus var. granulipennis: Apfelbeck, 1917:
227 (South Albania: Shen Thanas Mt. near Vlor, loc. typ.; Northwestern Greece: Peristeri Mt.
near Joannina, loc. typ.); Apfelbeck, 1918b: 89 (South Albania: Shen Thanas Mt. near Vlor,
loc. typ.; Northwestern Greece: Peristeri Mt. near Joannina, loc. typ.); Apfelbeck, 1930: 172
(South Albania: Shen Thanas Mt. near Vlor, loc. typ.; Northwestern Greece: Peristeri Mt.
near Joannina, loc. typ.). Aptinus merditanus merditanus: Hrdlika, 2003: 212.
Material studied. Merdita M. Schit / Coll. Apfelbeck / merditanus Typ. Apf., 1 m. (HNHM);
1 s. from afa Kjores, 53 s. from Llogora and Logara, 8 s. from Maja e Tartarit, 40 s. from
Mal i Dajtit, 1 s. from Mal i Krujes, 10 s. from Shen Ili, 1 s. from Tomor (MNHUB).
Notes. Apfelbeck published three consecutive descriptions for both A. merditanus and A.
merditanus var. granulipennis (Apfelbeck, 1917; Apfelbeck, 1918b; Apfelbeck, 1930) and the second
ones have long time been accredited to be valid. Actually this is incorrect and the original
descriptions must be assigned to the oldest ones. Aptinus lugubris rufipes (Apfelbeck, 1907a:
499) is nomen nudum (see also Hrka, 1988: 292). In that case, Apfelbeck had most probably the
intention to introduce a name for A. merditanus earlier than the year of 1917.
41
Notes. Hrdlika (2003: 214) has omitted the species for Albania; therefore its occurrence in
the country needs further confirmation.
Subgenus Cnecostolus Reitter, 1919
15.11. Brachinus bayardi Dejean, 1831
References. Brachinus bayardi: Schatzmayr, 1943b: 112 (Mal i Krujes).
Material studied. Liqen i Butrintes Vivari / leg. Bischoff Albania 1934, 2 s. (MNHUB);
Gjormi / Albania leg. Bischoff 1935, 1 s. (MNHUB).
Notes. Hrdlika (2003: 214) has omitted the species for Albania. Here it is confirmed for the
country.
15.12. Brachinus bipustulatus Quensel, 1806
References. Brachinus bipustulatus: Apfelbeck, 1904: 354 (Velipoja; Vlor); Apfelbeck, 1909:
298 (Vlor); Reitter, 1919: 135; Hieke & Wrase, 1988: 166; Hrdlika, 2003: 214.
Material studied. Scutari / collectio Paganetti, 1 s. (NMW); Mustajbeg Velipoja Albania,
1 s. (MNHUB); Durazzo Apfelbeck, 1 s. (HNHM); Oboti Bojana, 4 s. (HNHM); Alban.
Wgth. Valona, 1 s. (NMW); Durazzo / Albania leg. Bischoff 1931, 1 s. (MNHUB);
Myzeqe / Albania leg. Bischoff 1936, 1 s. (MNHUB); Sukthi bis Gurse / Albania leg.
Bischoff 1936, 1 s. (MNHUB).
15.13. Brachinus exhalans (P. Rossi, 1792)
References. Brachinus exhalans: Apfelbeck, 1904: 354 (Velipoja; Vlor); Apfelbeck, 1909: 298
(Vlor); Schatzmayr, 1943b: 112 (Lushnje; Mal i Krujes); Hrdlika, 2003: 214.
Material studied. Avlona, 3 s. (HNHM); Avlona Albania mer., 1 s. (HNHM); Albanien
Avlona v. Oertzen., 5 s. (MNHUB); Durazzo Apfelbeck, 2 s. (HNHM); Alban. Wgth.
Valona, 6 s. (NMW); Murshi / Albania leg. Bischoff 1936, 111 s. (MNHUB); Radona,
Shkiela, 21.VI.1994, 1 s., B. Guorguiev leg. (NMNHS).
Species insertae sedis
15.14. Brachinus nigricornis Gebler, 1829 [= incertus Brull, 1834]
References. Brachinus incertus: Apfelbeck, 1904: 350 (Velipoja near Lezh); Apfelbeck, 1909:
298 (Vlor); Hieke & Wrase, 1988: 165. Brachinus nigricornis: Hrdlika, 2003: 216.
Material studied. Durazzo, 1 s. (HNHM); Albania Avlona, 5 s. (HNHM); Bojana
Mundung, 3 s. (MNHUB); Alban. Wgth. Valona, 12 s. (NMW).
43
44
45
47
48
49
Material studied. Durazzo / Albania leg. Bischoff 1931, 1 s. (MNHUB); Himara Juni
/ Albania leg. Bischoff 1932, 2 s. (MNHUB); Liqen i Butrintes Vivari / leg. Bischoff
Albania 1934, 1 s. (MNHUB); Valona / Albania leg. Bischoff 1936, 4 s. (MNHUB); 2 km
N Durrs, 24.V.1993, 1 s., P. Stoev & D. Zaprjanova leg. (NMNHS); Shkodr, 12.VI.1993, 2 s.,
P. Beron & B. Petrov leg. (NMNHS).
20.2. Scarites terricola Bonelli, 1813
20.2.1. Scarites terricola terricola Bonelli, 1813
Material studied. Liqen i Butrintes Vivari / leg. Bischoff Albania 1934, 2 s. (MNHUB);
Bojana Shkoder / Albania leg. Bischoff 1937, 1 s. (MNHUB); Albania Sauk / 16.VI.1958
leg. Xh. Murraj, 1 s. (HNHM); Saranda District, Butrinti, 16.IV.1994, 1 s., S. Beshkov leg.
(NMNHS); Prmeti District, Pelombasi, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS).
Notes. New species for the fauna of Albania.
Subgenus Scallophorites Motschulsky, 1858
20.3. Scarites buparius (Forster, 1771)
References. Scarites buparius: Hieke et al., 1986: 184 (Vlor).
Material examined. Vom Alb. Amerik Institut / Zweifellos Alb. Konvenienc, 1 s.
(MNHUB); S Albanien Vlora, Strand p.7.58 leg. Grebenscikov, 1 s. (DEI).
Additional material included (studied A. Vigna Taglianti). Tiran, botanical garden, 6.VI.1998,
1 s., A. Paparistos leg. (cAVT).
Notes. Balkenohl (2003: 233) has omitted the species for Albania. Here it is confirmed for the country.
51
Material studied. Shklzn Alban., 3 s. (NMW); Albania Buneevce / Akad. Balk. Exp.
Csiki 1917, 14 s. (HNHM); Albania Kula Lums / Akad. Balk. Exp. Csiki 1918, 4 s.
(HNHM); Alban. Exp.18 Kula Ljums 26.VI.-3.VII., 62 s. (NMW); Skutari, Alb. Klenka, 3
s. (NMW); Vojusa Shkoder 1934, 1 s. (NMW); Dajtit Alb. Bischoff 1935, 8 s. (NMW);
Cucale / Albania leg. Bischoff 1936, 1 s. (NMW); Tomor Mt., 21.IX.1961, 2 m., 3 f., Xh.
Murraj leg. (MNST); Tepelen District, Vijos River, 2.VI.1994, 2 s., G. Blagoev leg. (NMNHS).
Notes. Bonavita & Vigna Taglianti (2005) concluded that A. caraboides and A. nebulosum are
two distinct species, the first monotypic and living in Central Europe, the second polytypic
(with four subspecies) and living in South Europe and Anatolia. The record of Murraj (1968)
about Bembidion elegans is based on misidentification. Except the material mentioned above the
present author has seen further more than 40 s. in MNHUB, which remained unregistered.
22.3. Asaphidion rossii (Schaum, 1857)
References. Asaphidion rossii: Hieke et al., 1986: 186 (Uji Ftohte, south of Tepelen); Marggi
et al., 2003: 243.
Material studied. Elbasan Alb., Mader, 1 s. (NMW); Tiran / Albania leg. Bischoff
1931, 2 s. (NMW; MNHUB); Bushek Alb. Elbasan Bischoff 1932, more than 45 s. (NMW;
MNHUB); Ibe Erzen / Albania leg. Bischoff 1932, 2 s. (NMW; MNHUB); Gjirakastr
Ung. Albania 1937 Bischoff 1934, 3 s. (NMW); Dajtit, Albania Bischoff 1935, 8 s. (NMW);
Mal i Dajtit / Albania leg. Bischoff 1935, 11 s. (MNHUB); Berat / Albania leg. Bischoff
1936, 2 s. (NMW); Mal i Dajtit / Albania leg. Bischoff 1936, 2 s. (MNHUB); Vojusa
Tepelene 1936, 4 s. (NMW); Bojana Shkoder / Albania leg. Bischoff 1937, 1 s. (MNHUB);
Lem i Kuri Ura Mes / Albania leg. Bischoff 1937, 1 s. (MNHUB); Mal i Dajtit / Albania
leg. Bischoff 1939, 2 s. (MNHUB).
22.4. Asaphidion stierlini (Heyden, 1880)
References. Asaphidion flavipes: Csiki, 1940: 212 (Kula Ljums).
Material studied. Albania Kula Lums / Akad. Balk. Exp. Csiki 1918, 3 s. (HNHM).
Notes. New species for the fauna of Albania.
53
Notes. In HNHM, I found single specimen identical with that cited by Csiki (1940) to belong
to B. quadripustulatum. The occurrence of B. quadrimaculatum in Albania is very probable due to
the distribution of this common species on the Balkans.
23.1. Bembidion quadripustulatum Audinet-Serville, 1821
23.1.1. Bembidion quadripustulatum quadripustulatum Audinet-Serville, 1821 [= quadriguttatum
(Olivier, 1795) nec Fabricius, 1775]
Material studied. Skutari Bojana / Apfelbeck, 1 s. (NMW); Alban., Wgth. Valona /
Apfelbeck, 1 s. (NMW); Elbasan, Alb. 15.IX.18 Dr. H. Priesner, 1 s. (NMW); Albania
Kula Lums / Akad. Balk. Exp. Csiki 1918, 1 s. (HNHM); Skutari, Alb. Klenka, 3 s.
(NMW); Elbasan Alb., Mader, 1 s. (NMW); Bojana - Ufer b. Shkoder, Alb. / Alb. Winkler
1934, 1 s. (NMW); Durazzo, Alb. Bischoff , 1 s. (NMW); Tiran Park, 1.III.1998, 1 s., R.
Ward leg. (CMNH).
Notes. New species for the fauna of Albania.
Subgenus Bembidionetolitzkya E. Strand, 1929
Notes. This is one of the most complex subgenera from the genus in the Balkans. Apparently,
the center of diversity of this subgenus in the Peninsula is situated in the northern half of
Albania and the adjacent territories of Montenegro and Kosovo. The taxonomical problems
with several yet not specified forms inhabiting the studied territory deserve special endeavour.
Species group tibiale
23.2. Bembidion rhodopense Apfelbeck, 1902
References. Bembidion rhodopense: Hieke & Wrase, 1988: 47 (Korab Mt.); Guorguiev &
Guorguiev, 1995: 106; Guorguiev et al., 1997: 14; Sakalian & Guorguiev, 1997: 46.
Material studied. Albanien, 5 s. (NMW); Shklzn Alb. Bischoff 1935, 1 s. (NMW);
Korab / Albania leg. Bischoff 1935, 2 s. (NMW; MNHUB); Korab / Albania leg.
Bischoff 1936, 5 s. (NMW; MNHUB); Korab, Alb. Guri Kuq Bischoff 1936, 27 s. (NMW;
MNHUB).
Notes. Marggi et al. (2003: 246) have omitted the species for Albania. Here it is confirmed for
the country.
23.3. Bembidion tibiale (Duftschmid, 1812)
References. Bembidion tibiale: Apfelbeck, 1907a: 496 (Kiri River near Mesi); Hieke & Wrase,
1988: 47.
Material studied. Shklzn Alban., 1 s. (NMW); Shklzn / Albania Bischoff 1935, 9
s. (MNHUB).
Notes. Marggi et al. (2003: 246) have omitted the species for Albania. Here it is confirmed for
the country.
54
55
Notes. In both the shape and the internal structure of the median lobe this species seems
most related to B. fasciolatum. On the other hand, in the wider margin of the pronotum and
bigger size of the body, it resembles B. ascendens.
23.10. Bembidion concoeruleum Netolitzky, 1943
References. Bembidion coeruleum: Apfelbeck, 1907a: 498 (Fani i Madh River near Oroshi); Hieke &
Wrase, 1988: 47. Bembidion concoeruleum: Netolitzky, 1943a: 114 (Oroshi; Shkodr; Dukati); Marggi et
al., 2003: 245. Bembidion coeruleum concoeruleum: Hieke et al., 1986: 185 (Borshi near Vlor; Uji Ftohte,
south of Tepelen). Ocydromus concoeruleus: Bonavita & Vigna Taglianti, 1993: 84.
Material studied. Orosi (Ptr.) Alban. spt., 5 s. (HNHM; NMW); Skutari Mesi / Apfelb.
Alban. spt. IV.V.05, 1 s. (NMW); Merdita Oroshi / Apfelb. Alban. spt. IV.V.05, 3 s.
(NMW); Prokletija, 1 s. (NMW); Prokletija 1300-2000 m / Alban. montenegr. Grenze,
Penther14, 1 s. (NMW); Dukati, 1 s. (NMW); Skutari, Alb. Klenka, 4 s. (NMW); Skutari,
Alb. Mader, 1 s. (NMW); Lumi, Benges Tepelene, Alb. m., 2 s. (NMW); Argirokastron
Albania mai 1929 Weirather, 6 s. (NMW); Lumi Dukati Alban. mer. / Winkler, Lona,
Bischoff V.1931, 1 s. (NMW); Korab, Alb. Guri Kuq Bischoff 1936, 2 s. (MNHUB); Tepelen
District, Vijos River, 2.VI.1994, 2 m., G. Blagoev leg. (NMNHS).
Notes. All previously published data for B. coeruleum Audinet-Serville, 1821 from the Balkans
refer to this species (see also Bonavita & Vigna Taglianti, 1993).
# Bembidion fasciolatum (Duftschmid, 1812)
References. Bembidion fasciolatum: Apfelbeck, 1907a: 496, 498 (Kiri River near Mesi; Fani i
Madh River near Oroshi); Hieke et al., 1986: 185 (Uji Ftohte near Tepelen); Marggi et al.,
2003: 245.
Notes. I failed to find material of this species from the Balkan Peninsula in the collections,
excluding a single specimen from the locality Travnik. There are two places with this
name - one in Slovenia and one in Bosnia Herzegovina. B. fasciolatum is removed from the
list of the Albanian Carabidae. The records cited for it in the past most probably concern
either an unnamed species (the record near Oroshi) or B. brunoi (the records by Mesi River
and in South Albania).
! Bembidion sp. (ex. gruppo fasciolatum)
References. Bembidion fasciolatum: Apfelbeck, 1907a: 498 (Fani i Madh River near Oroshi);
Marggi et al., 2003: 245.
Material studied. Orosi (Ptr.) Alban. spt., 6 s. (DEI; HNHM; MNHUB).
Notes. The specimens from DEI and HNHM have been identified as Bembidion merditanum
Apfelbeck as two of them from the first depository were incorrectly designated as syntypes
of this form. In fact, Bembidion merditanum Apfelbeck is nomen nudum. The series from Oroshi
may belong to separate species.
[subgenus Bracteon Bedel, 1879]
[Bembidion litorale (Olivier, 1790) [= elegans Germar, 1824]]
References. Bembidion elegans: Murraj, 1968: 33 (Tomor Mt.).
Notes. Murraj (1968) cited this species from the Tomor Mt. Among the material loaned from
MNST, I found five specimens from this locality bearing label data as the published ones and
COLEOPTERA: CARABIDAE: TRECHINAE
57
identified by the author as B. elegans. In fact, all the specimens concern Asaphidion nebulosum
balcanicum. For the present B. litorale is treated as an expected species for Albania because it
inhabits most of the Balkan countries.
Subgenus Chlorodium Motschulsky, 1864
23.11. Bembidion splendidum Sturm, 1825
23.11.1. Bembidion splendidum pincum De Monte, 1957
References. Bembidion splendidum: Apfelbeck, 1907a: 495 (Buen River).
Material studied. Scutari Bojana / Apfelb. Alban. spt. IV.V.05, 5 s. (NMW); Vojusa
Tepelene / Albania leg. Bischoff 1936, 1 s. (NMW); Bojana Shkoder / Albania leg.
Bischoff 1937, 3 s. (NMW).
Notes. Marggi et al. (2003: 248) have omitted the species for Albania. Here it is confirmed for
the country. According to Bonavita and Vigna Taglianti (2005) the Balkan populations of the
species belong to ssp. pincum.
Subgenus Diplocampa Bedel, 1896
23.12. Bembidion assimile Gyllenhal, 1810
References. Bembidion assimile: Apfelbeck, 1904: 111 (Velipoja); Hieke & Wrase, 1988: 57.
Material studied. Mustajbeg Velipoja Albania, 2 s. (HNHM).
Notes. Marggi et al. (2003: 248) have omitted the species for Albania. Here it is confirmed for
the country.
23.13. Bembidion fumigatum (Duftschmid, 1812)
References. Bembidion fumigatum: Apfelbeck, 1904: 111 (Velipoja near Lezh); Hieke & Wrase,
1988: 57.
Notes. Marggi et al. (2003: 248) have omitted the species for Albania. Actually, further
confirmation for the country is needed.
Subgenus Emphanes Motschulsky, 1850
23.14. Bembidion axillare (Motschulsky, 1844)
23.14. Bembidion axillare occiduum Margi & Huber, 2001 [= rivulare (Dejean, 1831)]
References. Bembidion rivulare: Hieke & Wrase, 1988: 58.
Material studied. Durrs Alban., 1 s. (NMW); Albanien Ruskuli 24.IX.18 Dr. H. Priesner,
1 s. (NMW); Ruskuli Alb., Mader, 1 s. (NMW); Durazzo, Alb. Ravasini, Lona, 2 s. (HNHM);
Durrazo, Alb. Bischoff , 4 s. (NMW); Durazzo / Albania leg. Bischoff 1931, 9 s.
(MNHUB); Valona, Alb. Bischoff 1936, 2 s. (NMW); Radona, Shkiela, 21.VI.1994, 3 s., B.
Guorguiev leg. (NMNHS).
Notes. Marggi et al. (2003: 248) have omitted the species for Albania. Here B. axillare is
confirmed for the country with first detailed records.
58
59
Notes. Marggi et al. (2003: 249) have omitted the species for Albania. Here it is confirmed for
the country.
Subgenus Euperyphus Jeannel, 1941
23.19. Bembidion combustum Mntries, 1832
23.19.1. Bembidion combustum combustum Mntries, 1832
References. Bembidion combustum: Apfelbeck, 1907a: 498, 501 (Fani i Madh River near Oroshi);
Hieke et al., 1986: 185 (Uji Ftohte, south of Tepelen); Hieke & Wrase, 1988: 48. Bembidion
combustum combustum: Marggi et al., 2003: 249.
Material studied. Albania, Berat, 1 s. (NMW); Merdita Oroschi / Apfelb. Alban. spt. IV.V.05,
9 s. (NMW); Oroi (Ptr.) Alban. spt., 11 s. (HNHM; MNHUB); Berat, Alb. Ravasini, Lona, 3 s.
(NMW); Skutari, Alb. Klenka, 1 s. (NMW); Tiran, Alb. Alf. Bischoff , 2 s. (NMW); Cucale
Geb. / Albania leg. Bischoff 1931, 1 s. (NMW); Elbasan Juni / Albania leg. Bischoff 1932,
1 s. (MNHUB); Lum i Tirans Alb. V.1934 Mjr. Strupi, 2 s. (NMW); Lum Erzen Alb. V.1934
Mjr. Strupi, 2 s. (NMW); Tiran / lgt. Bischoff Albania 1934, 2 s. (MNHUB); Tiran / lgt.
Bischoff Albania 1935, 2 s. (MNHUB); Tomor Mt., 21.IX.1961, 1 m., Xh. Murraj leg. (MNST).
23.20. Bembidion scapulare Dejean, 1831
23.20.1. Bembidion scapulare oblongum Dejean, 1831
References. Bembidion oblongum: Apfelbeck, 1907a: 498 (Fani i Madh River near Oroshi).
Bembidion scapulare oblongum: Netolitzky, 1943b: 17; Hieke & Wrase, 1988: 50.
Material studied. Merdita Oroschi / Apfelb. Alban. spt. IV.V.05, 4 s. (NMW); Berat Alb.
Dr. J. Mller18, 1 s. (NMW); Elbasan, Alb. 4.IX.18. Dr. H. Priesner, 1 s. (NMW); Elbasan
Alb., Mader, 3 s. (NMW); Lumi Dukati Alban. mer. / Winkler, Lona, Bischoff, V.1931, 1 s.
(NMW); Elbasan Juni / Albania leg. Bischoff 1932, 1 s. (MNHUB); Tomorica / Albania
leg. Bischoff 1932, 1 s. (MNHUB); Tirane / leg. Bischoff Albania 1934, 2 s. (NMW;
MNHUB); Tepelen District, Vijos River, 2.VI.1994, 2 s., G. Blagoev leg. (NMNHS).
Notes. Marggi et al. (2003: 250) have omitted the species for Albania. Here it is confirmed for
the country. Presently the subspecific status of the Balkan populations remains unsettled.
23.21. Bembidion testaceum (Duftschmid, 1812)
23.21.1. Bembidion testaceum testaceum (Duftschmid, 1812)
Material studied. Berat, Alb. Ravasini, Lona, 1 s. (NMW); Bojana-Schkodr Alb., Bischoff
1934, 1 s. (NMW).
Notes. New species for the fauna of Albania.
Subgenus Metallina Motschulsky, 1850
23.22. Bembidion lampros (Herbst, 1784)
References. Bembidion lampros: Apfelbeck, 1904: 83 (Velipoja; Vlor); Apfelbeck, 1909: 297
(Vlor; Sop); Csiki, 1940: 212 (Gjalic Mt.); Hieke & Wrase, 1988: 44; Marggi et al., 2003: 251.
60
Material studied. Albania, Mts. Gyalica Lums / Akad. Balk. Exp. Csiki 1918, 1 s. (HNHM);
North Albanian Alps, Bog, 1000-1100 m, 30.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS);
North Albanian Alps, Bog, 1600 m, near cave B 52, 20.VI.1994, 1 s., T. Ivanova leg. (NMNHS);
Tiran Park, 22-31.X.1997 / 1-9.IX.1997 / 8.II.1998 / 11.II.1998 / 15.II.1998 / 22.II.1998, 9
s., from grass and soil around tree trunks, R. Ward leg. (CMNH).
Notes. Except the specimens mentioned above, I have seen additionally nearly 100 s. in NMW
and MNHUB, which remained unregistered.
23.23. Bembidion properans (Stephens, 1828)
References. Bembidion lampros var. properans: Csiki, 1940: 212 (Kula Ljums). Bembidion properans:
Hieke & Wrase, 1988: 44.
Material studied. Albania Kula Lums / Akad. Balk. Exp. Csiki 1917, 1 s. (HNHM); Elbasan
Alb., Mader, 1 s. (HNHM); Lumi i Tirans / Albania leg. Bischoff 1935, 1 s. (MNHUB);
Mal i Dajtit / Albania leg. Bischoff 1935, 2 s. (MNHUB); Tiran / Albania leg. Bischoff
1935, 4 s. (MNHUB); Sukthi bis Gurse / Albania leg. Bischoff 1938, 2 s. (MNHUB);
Tiran Park, 8.II.1998, 2 s., from grass and soil around tree trunks, R. Ward leg. (CMNH).
Notes. Marggi et al. (2003: 251) have omitted the species for Albania. Here it is confirmed for
the country. Except the specimens mentioned above, in NMW the present author has seen
other specimens, which remained not registered.
Subgenus Microserrullula Netolitzky, 1921
23.24. Bembidion quadricolle (Motschulsky, 1844) [= inserticeps Chaudoir, 1850; cyaneomicans Maan, 1933]
References. Bembidion inserticeps: Apfelbeck, 1907a: 495 (Buen River). Bembidion quadricolle:
Hieke & Wrase, 1988: 42; Nitzu, 2003: 180.
Material studied. Albania Scutari, 1 s. (MNHUB); Skutari Bojana, 2 s. (HNHM); Skutari
Bojana / Apfelb. Alban. spt. IV.V.05, 5 s. (NMW; MNHUB); Berat, Alb. Ravasini, Lona,
1 s. (NMW); Berat, Alb. Ravasini, Lona / cyaneomicans det. Paul Meyer, 1 s. (NMW); Berat
/ Albania leg. Bischoff 1931, 1 s. (NMW); Lumi Berati Alban. mer. / Winkler, Lona,
Bischoff, V.1931, 1 s. (NMW); Tepelene / Albania leg. Bischoff 1932, 5 s. (NMW;
MNHUB); Ura Erzen / Albania leg. Bischoff. 1932, 2 s. (NMW); Bojana Shkoder /
Albania leg. Bischoff 1934, 2 s. (NMW).
Notes. Marggi et al. (2003: 251) have omitted the species for Albania. Here B. quadricolle is
confirmed for the country.
Subgenus Nepha Motschulsky, 1864
23.25. Bembidion callosum Kster, 1847
23.25.1. Bembidion callosum callosum Kster, 1847 [= laterale Dejean, 1831; pseudolaterale Netolitzky,
1943a: 103]
References. Bembidion laterale: Apfelbeck, 1904: 108 (Velipoja). Bembidion laterale pseudolaterale:
Netolitzky, 1943a: 103.
COLEOPTERA: CARABIDAE: TRECHINAE
61
Material included (studied P. Bonavita). Maja e Tartarit / Albania leg. Bischoff 1936, 1 f. (NMW).
Notes. Marggi et al. (2003: 252) have omitted the species for Albania. Here it is confirmed for
the country.
23.26. Bembidion caucasicum (Motschulsky, 1844) [= substriatum Chaudoir, 1846]
References. Bembidion substriatum: Apfelbeck, 1907a: 501 (Merdita); Csiki, 1940: 213 (Shklzen
Mt.; Gjalic Mt.). Bembidion caucasicum: Mller, 1937: 124; Hieke & Wrase, 1988: 56.
Material studied. Merdita Munela, 2 s. (NMW); Albania Galica Lums / Akad. Balk.
Exp. Csiki 1917, 4 s. (HNHM); Albania, Mts Gyalicalums / Akad. Balk. Exp. Csiki 1917,
1 s. (HNHM); Albania Sklsen / Akad. Balk. Exp. Csiki 1917, 1 s. (HNHM); Tomor:
Tomor Alban. mer. / Winkler, Lona, Bischoff, V.1931, 1 s. (NMW); Gjallica, Alb. Bischoff ,
1 s. (NMW); Maja e Jezerc Albania Bischoff leg., 2 s. (NMW); Korab, Guri Kua Alban.
Bischoff 35, 5 s. (NMW); Shklzen, Alb. Bischoff 1935, 1 s. (NMW); Maja e Policanit /
Albania 1936 leg. Ramadan coll. Bischoff , 1 s. (NMW); Korab / Albania leg. Bischoff
1936, 1 s. (NMW); Shklzen, Alb. Bischoff 5.8.1936, 3 s. (HNHM; NMW); North Albanian
Alps, Maje Radohimes, 2200-2400 m, 29.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS).
Additional material included (studied A. Vigna Taglianti). Tropoia, Kollats Mt., Qafa-eRubs, 2000 m, 8.VII.1993, 2 f., L. Latella leg. (cAVT).
Notes. Marggi et al. (2003: 252) have omitted the species for Albania. Here it is confirmed for
the country. Besides the material studied above, in NMW and MNHUB I have seen other
numerous specimens from the massifs of the North Albanian Alps, Korab Mt., Koritnik Mt.,
Gjalic Mt. and Maja e Policanit Mt.
23.27. Bembidion vseteckai Maan, 1936
23.27.1. Bembidion vseteckai dissimile G. Mller, 1943
References. Bembidion genei illigeri: Mller, 1943: 13; Hieke et al., 1986: 185 (Uji Ftohte, south
of Tepelen; Kula e Lums near Kuks); Hieke & Wrase, 1988: 55. Bembidion illigeri: Marggi et
al., 2003: 253.
Material studied. Albania Kula Lums / Akad. Balk. Exp. Csiki 1918, 3 s. (HNHM); Elbasan,
Alb. Leop. Mader leg., 2 s. (NMW); Kruja Alb. Leop. Mader, 2 s. (NMW); Terbaci Alban.
mer. / Winkler, Lona, Bischoff, V.1931, 1 s. (NMW); Mal i Dajtit Mt., 6.V.1962, 6 s. (MNST).
Notes. Except the specimens studied above I have seen several additional ones in NMW,
which remained unregistered. Bonavita (2005), as well P. Bonavita and A. Vigna Taglianti
(personal communication), noticed that all records or literature data about B. genei illigeri
Netolitzky, 1914 from the southern parts of Balkan Peninsula concern B. vseteckai dissimile.
Subgenus Notaphemphanes Netolitzky, 1920
23.28. Bembidion ephippium (Marsham, 1802)
References. Bembidion ephippium: Hieke & Wrase, 1988: 46.
Material studied. Valona, Albania, 4 s. (NMW); Durazzo Alb. Dr. J. Mller, 5 s. (NMW);
Durazzo / Albania leg. Bischoff 1931, 2 s. (MNHUB); Durres / Albania leg. Bischoff
1932, 9 s. (MNHUB).
62
Notes. Marggi et al. (2003: 253) have omitted the species for Albania. Here it is confirmed for
the country with first detailed records.
Subgenus Notaphus Dejean, 1821
23.29. Bembidion varium (Olivier, 1795) [= rumelicum Apfelbeck, 1902; dyrrhachii De
Monte, 1947]
References. Bembidion varium: Apfelbeck, 1894: 547; Apfelbeck, 1904: 85 (Velipoja near Lezh);
Hieke & Wrase, 1988: 46; Marggi et al., 2003: 254. Bembidion varium dyrrhachii: De Monte, 1947:
52 (Durrs, loc. typ.). Bembidion rumelicum: Guorguiev & Guorguiev, 1995: 100.
Material studied. Bojana Oboti, Alb., 1 s. (NMW); Oboti Schirei / Apfelbeck, 1 s.
(NMW); Durazzo / Albania leg. Bischoff 1931, 12 s. (MNHUB); Durres / Albania leg.
Bischoff 1932, 7 s. (MNHUB); Drin Shkoder / Albania leg. Bischoff 1937, 1 s. (NMW);
Radona, Shkiela, 21.VI.1994, 4 s., B. Guorguiev leg. (NMNHS); Tiran Park, 22.II.1998, 30
s., from grasses along small sluggish stream, R. Ward leg. (CMNH).
Notes. Except the material studied above the present author has seen more specimens in
NMW, which remained unregistered.
Subgenus Ocydromus Clairville, 1806
23.30. Bembidion aetolicum Apfelbeck, 1901
23.30.1. Bembidion aetolicum aetolicum Apfelbeck, 1901
References. Bembidion aetolicum: Netolitzky, 1943b: 12 (Tomor Mt.). Bembidion aetolicum
aetolicum: Mller-Motzfeld, 1996b: 164 (Tomor Mt.: Buranj; Tomor Mt.: Kulmak); Marggi
et al., 2003: 254.
Material studied. Tomor: Kulmak Albania mer., 1 s. (NMW); Tomor Buranj Alb.
10.4.1932 Alf. Bischoff , 4 s. (NMW); Barsh-i-Vjetr Wildbach-Quellen Bischoff
21.VII.1940, 2 s. (NMW).
23.31. Bembidion decorum (Zenker in Panzer, 1799)
23.31.1. Bembidion decorum decorum (Zenker in Panzer, 1799)
References. Bembidion decorum: Apfelbeck, 1907a: 496, 498 (Kiri River near Mesi; Fani i Madh
River near Oroshi); Hieke et al., 1986: 185 (Uji Ftohte, south of Tepelen; Lur e Vjetr near
Lura Mt.); Hieke & Wrase, 1988: 49. Bembidion decorum decorum: Mller-Motzfeld, 1996b: 153
(Shkodr; Mal i Dajtit Mt.; Shklzen Mt.; Bushek near Elbasan; Tiran; Zeza-Brche near
Kruja; Lumi i Benes near Tepelen; Tomor Mt.: Kulmak; Miloti; Gjirokastr; Prespa Lake;
Cafa Stams; Kiri River near Mesi); Marggi et al., 2003: 255.
Material studied. Tepelen District, Vijos River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS);
Leskovac - Barmashi, 880 m, 2.VI.1995, 1 s., G. Blagoev leg. (NMNHS).
Notes. Except the specimens studied above, I have seen several more ones in NMW, which
remained unregistered.
63
Material studied. Korab, Guri-Kua Alban. Bischoff 35, 1 s. (NMW); Tomor-Tojk Alban
1935 Bischoff leg. / pindicum var.? dtt. Netolitzky, 1 s. (NMW); Tomor - Tojk Alban. 1935
Bischoff leg., 2 s. (NMW).
Notes. Marggi et al. (2003: 257) have omitted the species for Albania. The specimens studied
have been found to be conspecific with three syntypes of B. pindicum labelled Apfelbeck
Peristeri / (Peryphus) pindicum typ. Apf. / Typus (NMW), therefore the species is here
confirmed for Albania.
23.36. Bembidion praeustum Dejean, 1831
23.36.1. Bembidion praeustum praeustum Dejean, 1831
References. Bembidion praeustum: Hieke et al. 1986: 186 (Uji Ftohte, soth of Tepelen); Hieke
& Wrase, 1988: 52. Bembidion praeustum praeustum: Marggi et al., 2003: 257.
Material studied. Dukati / Apfelbeck, 1 s. (NMW); Tomor: Kulmak Albania mer. /
Ravasini & Lona, 1 s. (NMW); Lumi i Benes Tepelene, Alb. m. / Alb. Winkler, 1 s.
(NMW); Bushek b. Elbasan, Alb. Alf. Bischoff , 1 s. (NMW); Maje Shbogut Alnania mer. /
Winkler, Lona, Bischoff, V.1931, 2 s. (NMW); Humara juni / Albania leg. Bischoff 1932,
2 s. (MNHUB); Llogora / Albania leg. Bischoff 1932, 9 s. (MNHUB); Maranaj, Alb. Bischoff,
1934, 1 s. (NMW); Lumi i Tirans Alb. V.1934 Mjr. Strupi, 1 s. (NMW); Kleyr / Albania
leg. Bischoff 1936, 1 s. (NMW); Boge ober d. Quelle / Albania leg. Mueller 1939, 1 s.
(NMW); Borsh-i-Vjetr Wildbach-Quellen Bischoff 21.VII.1940, 8 s. (HNHM; NMW);
Borsh, Alb. Bischoff 1940, 1 s. (NMW); Tirane, Alb. dch. Winkler 42, 1 s. (NMW); Tomor
Mt., 21.IX.1961, 1 m., Xh. Murraj leg. (MNST); Prmeti District, Pelombasi, 2.VI.1994, 1 s., G.
Blagoev leg. (NMNHS); Tepelen District, Vijos River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS);
Mal i Dajtit Mt., 20 km E Tiran, 1000 m, 9.V.1995, 1 s., L. Penev leg. (NMNHS).
Notes. A part of the material in NMW has been determined as B. praeustum viridifluum J. Mller,
1929. However, I refer provisionally all the specimens studied to the nominotypical race.
23.37. Bembidion signatipenne Jacquelin du Val, 1852
References. Bembidion signatipenne: Hieke & Wrase, 1988: 53.
Notes. Marggi et al. (2003: 257) have omitted B. signatipenne for Albania. Actually, the species
needs further confirmation for the country.
23.38. Bembidion stolfai J. Mller, 1943
References. Bembidion stolfai: Mller, 1943: 12 (North Albanian Alps: Okol near Bog, loc.
typ.; Maja e Jezerc); Netolitzky, 1943a: 41 (North Albanian Alps: Okol near Bog; Maja e
Jezerc); Marggi et al., 2003: 257.
Material studied. Maja e Jezerce Albania sept. / Alb. Winkler 1934 / coll. Netolitzky
/ stolfai m. n. sp. det. J. Mller / Co-typus / Coll. Netolitzky, 1 f. syntype (NMW).
Notes. The syntype of B. stolfai was studied and compared with specimens of B. reiseri reiseri
Apfelbeck, 1902 from Bosnia Herzegovina (including a syntype labelled reis. 1900 Cvrstica
/ Testediolum bosnicum typ. Apf. / Reiseri / Typus), topotypical material of B. reiseri vranense
Apfelbeck, 1902 and extensive material of B. balcanicum balcanicum. B. stolfai is distinct from the
other two species in the pronotum wider than long being narrower in the anterior part and in
COLEOPTERA: CARABIDAE: TRECHINAE
65
the elytra having distinct transverse microsculpture. Besides, it differs from B. reiseri s.l. in the
wrinkles on outer basal fovea of pronotum clear and long (vs. reduced), the shoulders of
elytra distinct (vs. round) and the sides of elytra after middle parallel (vs. slightly round). B.
stolfai is also living sympatrically with B. balcanicum balcanicum in the massif of Maja e Jezerc.
Subgenus Odontium LeConte, 1848
23.39. Bembidion foraminosum Sturm, 1825
References. Bembidion foraminosum: Hieke & Wrase, 1988: 43.
Material studied. Tiran / Albania leg. Bischoff 1931, 2 s. (MNHUB).
Notes. Marggi et al. (2003: 257) have omitted the species for Albania. Here it is confirmed for
the country with first detailed record.
23.40. Bembidion striatum (Fabricius, 1792)
References. Bembidion striatum: Netolitzky, 1918: Beilage zu Heft 4-6 (Shkodr; Oboti);
Geisthardt, 1975: 8. Bembidion striatul sic!: Panin, 1942: 567 (Shkodr).
Material studied. Oboti Schirei, 1 s. (NMW); Skutari Bojana, 5 s. (NMW); Skutari
Bojana / Apfelb. Alban. spt. IV.V.05, 4 s. (NMW).
Notes. Marggi et al. (2003: 257) have omitted the species for Albania. Here it is confirmed for
the country.
Subgenus Omoperyphus Netolitzky, 1931
23.41. Bembidion hypocrita Dejean, 1831
23.41.1. Bembidion hypocrita illyricum Netolitzky, 1918
References. Bembidion hypocrita illyricum: Mller, 1918b: 89; Mller, 1926: 66; Netolitzky, 1943a: 122.
Material studied. Skutari, Alb. 4.X.18 Dr. H. Priesner, 1 s. (NMW); Skutari, Alb. Mader,
1 s. (NMW); Argyrkastron, Alb. leg. Weirather, 1 s. (NMW); Mal i Dajtit / Albania leg.
Bischoff 1935, 3 s. (NMW; MNHUB); Shkelzn, Alb. Bischoff, 5.8.1936, 1 s. (NMW);
Bischoff 21.VII.1940 Brsh-i-Vjetr Wildbach-Quellen, 7 s. (NMW); Brsh-i-Vjetr Alb.
21.7.1940 Bischoff , 1 s. (MNHUB).
Notes. Marggi et al. (2003: 258) have omitted the species for Albania. Here it is confirmed for
the country with first detailed records.
Subgenus Peryphanes Jeannel, 1941
23.42. Bembidion brunnicorne Dejean, 1831
23.42.1. Bembidion brunnicorne brunnicorne Dejean, 1831
References. Bembidion brunnicorne: Netolitzky, 1943a: 50; Csiki, 1940: 213 (Kula Ljums).
Material studied. Albania Kula Ljums, 1 s. (NMW); Albania Kula Lums / Akad. Balk.
Exp. Csiki 1918, 7 s. (HNHM); Skutari, Alb. 4.X.18 Dr. H. Priesner, 1 s. (NMW); Kruja,
Alban., Mader, 1 s. (NMW); Tomor: Kulmak Albania mer., 3 s. (NMW); Mal i Dajtit /
66
Albania leg. Bischoff 1935, 2 s. (HNHM; MNHUB); Maranaj 1937 / Albania bei Skutari,
1 s. (NMW); Brsh i Vjetr Wildbach-Quellen Bischoff, 21.VII.1940, 4 s. (NMW).
Notes. Marggi et al. (2003: 258) have omitted the species for Albania. Here it is confirmed for
the country. Besides the material listed above, I have seen additionally more than 70 s. from
different localities in MNHUB, which remained unregistered. All the material in the last museum
was identified as B. milleri.
23.43. Bembidion dalmatinum Dejean, 1831
23.43.1. Bembidion dalmatinum dalmatinum Dejean, 1831
References. Bembidion dalmaticum sic!: Anonymous, 1907: 286. Bembidion dalmatinum: Apfelbeck,
1907a: 498 (Fani i Madh River near Oroshi); Csiki, 1940: 213 (Gjalic Mt.); Schatzmayr, 1943b:
95 (Dukati; Lushnje); Hieke & Wrase, 1988: 53. Chlaenius nigricornis: Murraj, 1968: 33 (Tiran;
Tomor Mt.). Bembidion dalmatinum dalmatinum: Marggi et al., 2003: 258.
Material studied. Tomor Mt., 21.IX.1961, 9 s., Xh. Murraj leg. (MNST); Theth, 800-1200 m,
28.V.1993, 7 s., P. Beron & B. Petrov leg. (cPN; NMNHS); North Albanian Alps, Bog, 14001800 m, 1.VI.1993, 5 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bog,
1000-1100 m, 5-9.VI.1993, 5 s., P. Beron & B. Petrov leg. (NMNHS); Reshen District, Merkurth,
11.VI.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); Prmeti District, Pelombashi, 2.VI.1994,
1 s., G. Blagoev leg. (NMNHS); Tepelen District, Vijos River, 2.VI.1994, 27 s., G. Blagoev
leg. (NMNHS); Mal i Dajtit Mt., 20 km E Tiran, 1000 m, 9.V.1995, 1 s., L. Penev leg. (NMNHS);
Leskoec - Barmashi, 880 m, 2.VI.1995, 2 s., G. Blagoev leg. (NMNHS); 5 km Tepelen, Drina
River, 3.VI.1995, 2 s., G. Blagoev leg. (NMNHS); Ljukove near Vlor, 6.VI.1995, 4 s., G.
Blagoev leg. (NMNHS); Tiran Park, 1-9.XI.1997, 1 s., R. Ward leg. (CMNH).
Notes. About the record of Murraj (1968) above see the notes under Chlaenius nigricornis.
Besides the material studied, the present author has seen numerous other specimens in NMW
and MNHUB, which remained unregistered.
23.44. Bembidion deletum Audinet-Serville, 1821
23.44.1. Bembidion deletum deletum Audinet-Serville, 1821
Material studied. Borsh, Alb. Bischoff 1940, 1 s. (NMW).
Notes. New species for the fauna of Albania.
23.45. Bembidion grandipenne Schaum, 1862
23.45.1. Bembidion grandipenne grandipenne Schaum, 1862
References. Bembidion grandipenne: Apfelbeck, 1907a: 498, 501 (Fani i Madh River near Oroshi);
Hieke & Wrase, 1988: 54.
Material studied. Oroschi (Ptr.) Alban. spt., 4 s. (HNHM; MNHUB); Merdita Oroschi /
Apfelb. Alban. spt. IV.V.05, 3 s. (NMW); Albania Malija / Akad. Balk. Exp. Csiki 1917,
1 s. (HNHM); Skutari Alb., Mader, 1 s. (NMW); Lumi i Benes Tepelene, Alb. m., 2 s.
(NMW); Tomor: Kulmak Albania mer., 1 s. (NMW); Ibe-Erzen / Albania leg. Bischoff
1932, 2 s. (NMW; MNHUB); Tomor-Buranj juli 1932 / Albania leg. Bischoff 1932, 1 s.
(MNHUB); Shklzn, Alb. Bischoff 1935, 1 s. (NMW); Borsh-i-Vjetr Wildbach-Quellen
Bischoff 21.VII.1940, 4 s. (NMW; MNHUB); Borsh, Alb. Bischoff 1940, 1 s. (NMW).
COLEOPTERA: CARABIDAE: TRECHINAE
67
Notes. Marggi et al. (2003: 259) have omitted the species for Albania. Here it is confirmed for
the country.
# Bembidion milleri Jacquelin du Val, 1852
# Bembidion milleri milleri Jacquelin du Val, 1852
References. Bembidion milleri: Hieke & Wrase, 1988: 54.
Notes. According to Mller-Motzfeld (personal communication in 1994) B. milleri does not
live in the Balkans. P. Bonavita and A. Vigna Taglianti (personal communication) stated that it
occurs only in Slovenia and probably in Serbia. Here the upper opinions are followed, though
Marggi et al. (2003: 258-259) noted the species from the Balkan Peninsula.
Subgenus Peryphus Dejean, 1821
23.46. Bembidion bualei Jacquelin du Val, 1852
23.46.1. Bembidion bualei bualei Jacquelin du Val, 1852 [= albanicum J. Mller, 1935]
References. Bembidion andreae: Apfelbeck, 1904: 92 (Durrs). Bembidion andreae albanicum: Mller,
1935: 152 (Berat, loc. typ.; Tiran; Shkodr); Netolitzky, 1943a: 27 (Berat); Hieke et al. 1986:
186 (Uji Ftohte, south of Tepelen); Hieke & Wrase, 1988: 51; Guorguiev & Guorguiev,
1995: 111; Guorguiev et al., 1997: 15. Bembidion andreae bualei: Panin, 1942: 582 (Berat);
Schatzmayr, 1943b: 95 (Ura Zezes). Bembidion cruciatum albanicum: Marggi et al., 2003: 260.
Material studied. Alban. Exped. Gjalica Ljuma 17.-26.VI.18, 6 s. (NMW); Bojana Shkodr
Alb., Bischoff 1937 / Albania leg. Bischoff 1937, 1 s. (HNHM).
Notes. According to a recent study (Coulon, 2006) the Balkan populations of the species
related to B. cruciatum Dejean, 1831 belong to B. bualei bualei, and the form albanicum is a synonym
of the latter. Except the material studied above, I have seen more specimens from different
localities in NMW and MNHUB, which remained unregistered.
23.47. Bembidion femoratum Sturm, 1825
23.47.1. Bembidion femoratum femoratum Sturm, 1825
Material studied. Scutari, Alb. Klenka, 1 s. (NMW); Bojana, Alb. Shkodr Bischoff 1934,
31 s. (NMW); Albania leg. Bischoff 1937, 1 s. (NMW); Bojana Shkodr Albania leg. Bischoff
1937, 4 s. (NMW); 5 km from Tepelen, Drina River, 3.VI.1995, 2 s., G. Blagoev leg. (NMNHS).
Notes. New species for the fauna of Albania.
23.48. Bembidion subcostatum (Motschuslky, 1850)
23.48.1. Bembidion subcostatum vau Netolitzky, 1913 [= javurkovae Fassati, 1944]
References. Bembidion subcostatum: Hieke & Wrase, 1988: 52. Bembidion subcostatum javurkovae: Hieke
et al. 1986: 186 (Borshi near Vlor; Lur e Vjetr near Lura Mt.); Marggi et al., 2003: 263.
Material studied. Alban. Exp.18 Kula Ljums 26.VI.-3.VII., 4 s. (NMW); Scutari Alb. Dr.
J. Mller, 1 s. (NMW); Berat, Alb. sch. Winkler, 1 s. (NMW); Tirana Alb. dch. Winkler 42,
2 s. (NMW); Bushek Juni / Berat Alban. mer. / Albania leg. Bischoff 1932, 4 s. (NMW);
Tepelen District, Vijos River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Bureli, 22.VI.1994,
1 s., B. Guorguiev leg. (NMNHS).
68
Notes. Netolitzky identified the specimens in NMW as ssp. vau, ssp. javurkovae Fassati, 1944 or
ssp. spartanum Fassati, 1944. However, in a forthcoming paper these three taxa are synonymized
(Guorguiev & Lobo, in press). In HNHM I have seen numerous additional specimens mostly
from the localities Korab and Kula Lums, and in MNHUB - other more than 200 s., as all
those data remained unregistered.
23.49. Bembidion tetracolum Say, 1823
23.49.1. Bembidion tetracolum tetracolum Say, 1823
References. Bembidion ustulatum (Linnaeus) (sic!): Csiki, 1940: 213 (Kula Ljums; Korab Mt.);
Murraj, 1968: 33 (Tomor Mt.).
Material studied. Dukati / Apfelbeck, 1 s. (NMW); Merdita M. Scheit / Mali Scheit
Apfelbeck, 1 s. (NMW); Alban. Vermosa Penther, 1 s. (NMW); Kruja, Alban., Mader, 4
s. (NMW); Berat Alban. mer., 2 s. (NMW); Albania leg. Bischoff 1936, 2 s. (NMW);
Klcyr / Albania leg. Bischoff 1936, 1 s. (NMW); Bojana Schkoder / Albania leg.
Bischoff 1937, 2 s. (NMW); Tomor Mt., 21.IX.1961, 1 m., Xh. Murraj leg. (MNST); Ljukova,
100 m, 5.VI.1995, 1 m., 1 f., G. Blagoev leg. (NMNHS).
Notes. Marggi et al. (2003: 263) have omitted the species for Albania. Here it is confirmed
for the country. The record of Murraj (1968) is included here since the revision of one male
from the Tomor Mt., having the same label data as the published material, has been found
to be correct.
Subgenus Philochthus Stephens, 1828
23.50. Bembidion decolor Apfelbeck, 1911
References. Bembidion decolor: Apfelbeck, 1911: 210 (Buen River; Vlor); Apfelbeck, 1916:
340-341 (Buen River; Vlor); Netolitzky, 1943a: 63 (Elbasan; Kruja); Hieke & Wrase, 1988:
62; Marggi et al., 2003: 263.
Material studied. Alban. Wgth Valona / Coll. Apfelbeck, 1 m. (HNHM); Albania Kula
Lums / Akad. Balk. Exp. Csiki 1918, 1 s. (HNHM).
Notes. Apfelbeck published two descriptions of this species, the first in Latin and SerboCroatian and second one in Latin and German (Apfelbeck, 1911; Apfelbeck, 1916). The older
one is correctly treated as valid.
23.51. Bembidion guttula (Fabricius, 1792)
23.51.1. Bembidion guttula guttula (Fabricius, 1792)
References. Bembidion guttula: Hieke & Wrase, 1988: 61.
Material studied. Elbasan Alb., Mader, 1 s. (NMW); Tomor: Kulmak Alban. mer. / Alb.
Winkler leg., 1 s. (NMW); Maje Tartarit / Albania leg. Bischoff 1936, 1 s. (MNHUB).
Notes. Marggi et al. (2003: 264) have omitted the species for Albania. Here it is confirmed for
the country with first detailed records.
69
70
71
Korab / Akad. Balk. Exp. Csiki 1918, 15 s. (HNHM); Korab Preshk, Alb. Bischoff 1936,
1 s. (HNHM).
Additional material included (studied A. Vigna Taglianti). Tropoia, Kollats Mt., Qafa-eRubs, 2000 m, 8.VII.1993, 4 m., 1 f., L. Latella leg. (cAVT).
Notes. Marggi et al. (2003: 269) have omitted the species for Albania. Here it is confirmed for
the country. Besides the material studied above, I have seen other numerous specimens from
different localities in NMW and MNHUB, which remained unregistered.
Subgenus Trepanes Motschulsky, 1864
23.61. Bembidion articulatum (Panzer, 1796)
References. Bembidion articulatum: Apfelbeck, 1907a: 496 (Kiri River near Mesi); Marggi et al.,
2003: 270.
Material studied. Skutari Bojana, 1 s. (NMW); Skutari Alb. Dr. J. Mller, 1 s. (NMW);
Vojusa Tepelene / Albania leg. Bischoff 1936, 1 s. (NMW); Bojana Shkoder / Albania
leg. Bischoff 1937, 3 s. (NMW).
23.62. Bembidion maculatum Dejean, 1831
23.62.1. Bembidion maculatum maculatum Dejean, 1831
Material studied. Tirana, Alb. Ravasini, Lona, 1 s. (NMW).
Notes. New species for the fauna of Albania.
23.63. Bembidion octomaculatum (Goeze, 1777)
References. Bembidion octomaculatum: Apfelbeck, 1904: 110 (Velipoja near Lezh); Apfelbeck,
1907a: 496 (Kiri River near Mesi); Hieke & Wrase, 1988: 60.
Material studied. Alban. Wgth. Valona / Apfelbeck, 1 s. (NMW); Fieri, Albania a Semeni,
Mller, 4 s. (NMW); Skutari, Alb. Prof. Dr. Mller, 2 s. (NMW); Bojana-Ufer b. Shkoder, Alb.
/ Alb. Winker 1934, 1 s. (NMW); Bojana Shkoder / Albania leg. Bischoff 1937, 1 s. (NMW).
Notes. Marggi et al. (2003: 270) have omitted the species for Albania. Here it is confirmed for
the country.
72
73
74
75
76
Notes. After the original description, only Zamotailov (2003: 283) noted this species for
Albania without any detailed records. However, the type locality, Vunaj (in Albanian) or
Vusanje (in Serbo-Croatian) is situated in the Montenegrin section of the North Albanian
Alps very close to Albanian territory (Apfelbeck, 1918c: 160-161, 164-165). The massifs
of Kom Mt. and Zljeb Mt. also lie outside the investigated territory. Due to lack of any
reliable record from Albania D. malissorum is treated here as questionable. Very probably it
inhabits the country.
33.1. Deltomerus parumpunctatus Zamotailov, 1990
References. Deltomerus parumpunctatus: Zamotailov, 1990: 133, 135 (North Albanian Alps: afa
Borit, loc. typ.); Zamotailov, 1992: 56; Zamotailov, 2003: 283.
Material studied. Wgth. 1918 Menina pl. N. alb. alp. / Deltomerus parumpunctatus m. paratype
V. N. Kurnakov det. 1970, 1 m. designated by mistake as paratype (NMW); Wgth. 1918
Menina pl. / Deltomerus parumpunctatus m. alotype V. N. Kurnakov det. 1970, 1 f. designated
by mistake as paratype (NMW).
Notes. Zamotailov (1990: 135) have noted no paratypes in the description of D. parumpunctatus.
For this reason, the two specimens in NMW designated as paratypes should not be considered
as part of the type series.
Subgenus Paradeltomerus Apfelbeck, 1908
33.2. Deltomerus paradoxus Apfelbeck, 1908
Notes. Apfelbeck published three consecutive descriptions for both the subgenus and the
species (Apfelbeck, 1908; Apfelbeck, 1917; Apfelbeck, 1930), the oldest ones are correctly
treated as valid.
? Deltomerus paradoxus paradoxus Apfelbeck, 1908
References. Deltomerus paradoxus: Zamotailov, 1992: 56. Deltomerus paradoxus paradoxus:
Zamotailov, 2003: 283.
Notes. Hieke (1981: 80) stated by mistake that D. paradoxus is described from Albania.
In reality, the nominotypical form was described from the alpine zone of the Shar
Planina Mts. (Apfelbeck, 1907c; Apfelbeck, 1908). Dinchev & Atanasov (1998) stated
that only 1 % of the territory of the last mountain massif is situated in Albania. Actually
all specimens of the nominotypical race in the collections originate from Macedonia or
the Kosovo Region of Serbia. Due to lack of reliable data the subspecies is treated as
doubtful for Albania.
33.2.1. Deltomerus paradoxus korabensis J. Mller, 1936
References. Deltomerus paradoxus korabensis: Mller, 1936: 108-110 (Korab Mt., loc. typ.; Bjeshka
Matroxh); Zamotailov, 2003: 283. Deltomerus paradoxus: Zamotailov, 1992: 56.
Material studies. Nearly 15 s. from Korab Mt. (MNHUB).
77
79
1983: 249; Casale & Laneyrie, 1982:199; Genest & Juberthie, 1994: 592 (cave at Shen Elias
Mt., south of Vlor); Magrini et al., 1995b: 25 (Llogora Mt., cave Stera Kullxhive, south of
Vlor); Moravec et al., 2003: 304.
Material studied. Stere gropa de Vasos / Reza e Kanalit, Alban. m. / Winkler, Lona,
Bischoff, V.1931, 8 syntypes (NMW); Shen-ELias Juni 1932 / Albania leg. Bischoff 1932,
2 s. (MNHUB); Shen-Elias / leg. Bischoff Albania 1934, 1 f. (HNHM); Shen-ELias Juni
1932 / leg. Bischoff Albania 1934, 3 s. (MNHUB).
Species group fuchsi
38.5. Duvalius fuchsi Scheibel, 1937
References. Duvalius fuchsi: Scheibel, 1937a: 15-18, 20 (nameless cave near Llogora, south of
Vlor, loc. typ.); Turin, 1981: 111; Casale, 1983: 249; Casale & Laneyrie, 1982: 199; Genest &
Juberthie, 1994: 592 (cave at Llogora, south of Vlor); Moravec et al., 2003: 303.
Material studied. Hhle in der Llogora / Albania leg. Bischoff 1935, 1 syntype (MNHUB);
Hhle i.-d. Llgora / Bischoff mai 1935, 4 syntypes (MNHUB).
37.6. Duvalius pominii Schatzmayr, 1943
References. Duvalius pominii: Schatzmayr, 1943b: 96-97, 99 (cave Stere e Vogel and cave Stere
e Madhe near Dukati, south of Vlor, loc. typ.); Turin, 1981: 112; Genest & Juberthie, 1994:
592 (cave Stere e Vogel and cave Stere e Madhe near Dukati); Moravec et al., 2003: 305.
38.7. Duvalius regiszogui Meschnigg, 1936
References. Duvalius regiszogui: Meschnigg, 1936: 46-47 (cave Homberg-Hhle Maja e Tanasit
Mt., loc. typ.); Scheibel, 1937a: 18, 20 (Maja e Tanasit Mt., south of Vlor); Schatzmayr, 1943b:
99 (Maja e Tanasit Mt.: cave Homberg); Turin, 1981: 113; Casale, 1983: 249; Casale & Laneyrie,
1982:199; Genest & Juberthie, 1994: 592 (cave Homberg, Maja e Tanasit Mt., south of
Vlor); Moravec et al., 2003: 305.
II. North Aegean groups
Species group leonhardi
38.8. Duvalius sturanyi (Apfelbeck, 1904)
38.8.1. Duvalius sturanyi stenocephalus (Apfelbeck, 1918)
References. Trechus stenocephalus: Apfelbeck, 1918c: 164 (North Albanian Alps, Vermosh, 1100
m, loc. typ.). Duvalius leonhardi sturanyi: Jeannel, 1928: 548 (North Albanian Alps: Vermosh,
1100 m). Duvalius sturanyi stenocephalus: Scheibel, 1937a: 18-20 (North Albanian Alps: Vermosh,
Menina Planina Mt.); Pretner, 1978: 130, 135, 148, 167-168 (Vermosh; Menina Planina Mt.);
Moravec et al., 2003: 305. Duvalius leonhardi stenocephalus: Maan, 1939c: 60 (Vermosh). Duvalius
sturanyi: Casale & Laneyrie, 1982:199; Genest & Juberthie, 1994: 592 (North Albanian Alps:
Menina Planina Mt. near Vermosh).
COLEOPTERA: CARABIDAE: TRECHINAE
81
Material studied. Cukali Gebirge N-Albanien Lona leg. 1914, 4 s. (NMW); Scpedha nGiuka
Zukali Albanien Lona Giuno 1914, 1 s. (DEI); Cukale Albania sept. / Bischoff 1930 coll.
A. Winkler, 5 s. (NMW); Cucale Geb. Hhle von Vukaj / Albania leg. Bischoff 1931, 1 s.
(MNHUB); Shpella Kodra orit / Cukale Geb. / leg. Bischoff Albania 1934, 3 s.
(MNHUB); Shpella Kodra Carit Thep / Cukale / Albania leg. Bischoff 1935, 8 s.
(MNHUB); Shpella Luga en Vukaj / Cukale / Albania leg. Bischoff 1935, 1 s.
(MNHUB); Shpella Rasa-Kunors Vukaj / Cukale / Albania leg. Bischoff 1935, 4 s.
(MNHUB); Shpella uka Vjit / Cucale / Albania leg. Bischoff 1936, 2 s. (MNHUB);
Shpella Goll / Cukale / Albania leg. Bischoff 1936, 1 s. (MNHUB).
Notes. N. lonae lonae is sympatric and syntopic with N. malissorum (J. Mller, 1914) in most of
the caves where they have been found. The report of this subspecies from the area of Rapsh
(Genest & Juberthie, 1994: 592) is erroneous since this region is populated by N. lonae atavus.
The nominotypical subspecies differs from both ssp. atavus and ssp. shkodrensis in the wider
and more rounded, almost not narrowed to apex, elytra.
39.1.2. Neotrechus lonae atavus J. Mller, 1934 [= bischoffi Meschnigg, 1936]
References. Neotrechus bischoffi: Meschnigg, 1936: 44-46 (North Albanian Alps: cave Shpella
e lugstit, Hot-Brigje near Rapsh, loc. typ.); Schatzmayr, 1943b: 98 (cave Shpella e lugstit,
Hot-Brigje near Rapsh); Casale & Laneyrie, 1983: 180. Neotrechus lonae atavus: Scheibel, 1936:
227-228 (cave Shpella e lugstit, Hot Brgje near Rapsh); Scheibel, 1937a: 20 (cave Hot
Brigje); Moravec et al., 2003: 315. Neotrechus lonai sic! atavus: Absolon & Maan, 1947: 2;
Maan, 1947: 11 (Rapsh); Pretner, 1978: 147, 166 (cave Shpella e lugstit, Hot Brgje near
Rapsh); Casale & Laneyrie, 1983: 180; Genest & Juberthie, 1994: 592 (cave Shpella e Lugstit,
cave Hot Brigje near Rapsh).
Material studied. Shpella e Lugstit / Brigie Hot / leg. Bischoff Albania 1934, 7 syntypes
of Neotrechus bischoffi (MNHUB); Shpella e Lugstit / Brigie Hot / Albania leg. Bischoff
1935, 11 syntypes of Neotrechus bischoffi (MNHUB).
Notes. Scheibel (1936) established the synonymy of N. bischoffi with N. lonae atavus. Regardless
of that, Schatzmayr (1943b) and Casale & Laneyrie (1983) listed the former as valid species.
The type localities of both N. lonae atavus and N. suturalis pretneri Scheibel, 1936 coincide and
are located in Montenegro very close to the Albanian border.
39.1.3. Neotrechus lonae shkodrensis Scheibel, 1936
References. Neotrechus lonae shkodrensis: Scheibel, 1936: 228-229 (cave Shpella kufinit, KoplikutHotit near Ivanaj, north of Liqen i Shkodrs, loc. typ.); Scheibel, 1937a: 20 (Ivanaj); Moravec
et al., 2003: 315. Neotrechus lonai sic! shkodrensis: Absolon & Maan, 1947: 2; Maan, 1947: 11
(Ivanaj); Pretner, 1978: 147, 166 (cave Shpella kufinit, Koplikut-Hotit near Ivanaj); Casale &
Laneyrie, 1983: 180; Drovenik & Peks, 1994: 45; Genest & Juberthie, 1994: 592 (cave Kufinit
et Kopikut, Hotit pes near Ivanaj, north of Liqen i Shkodrs); Drovenik & Peks, 1999: 51.
Material studied. Shpella Kufin Hot Koplicut Ivanaj / Albania leg. Bischoff 1935, 4
syntypes (MNHUB); Shpella Kufin Hot Koplicut Ivanaj / Albania leg. Bischoff 1937, 4 s.
(MNHUB).
Notes. Drovenik & Peks (1994: 45) and Drovenik & Peks (1999: 51) indicated that the type
locality of this subspecies is situated in Montenegro. Actually it is located in Albania, very
close to the state border.
83
85
1925: 23 (Tomor Mt.: Kulmak); Horion, 1941: 181; Schatzmayr, 1943b: 96 (Dukati; cave Stere
e Madhe near Dukati); Casale & Laneyrie, 1983: 147; Hieke & Wrase, 1988: 36; Nonveiller et
al., 1994: 9, 15; Guorguiev & Guorguiev, 1995: 75; Guorguiev et al., 1997: 11; Sakalian &
Guorguiev, 1997: 46.
Material studied. Kruja, Alban., Mader, 2 s. (HNHM).
Additional material included (studied A. Vigna Taglianti). Durrs, Plepa, 15.VIII.1995, 1
m., D. Antonelli leg. (cAVT).
Notes. In MNHUB I have seen also additional 39 s., which remained not registered.
40.5.2. Trechus subnotatus ljubetensis Apfelbeck, 1908
References. Trechus subnotatus ljubetensis: Moravec et al., 2003: 341.
Material studied. Albanien Avlona v. Oertzen, 2 s. (MNHUB); Albania / ljubetensis Apf.
R. Jeannel, 1 m. (NMW).
Notes. Apfelbeck published three consecutive descriptions of this taxon (Apfelbeck, 1908;
Apfelbeck, 1917; Apfelbeck, 1930), the oldest one is correctly treated as valid. The identification
of the two specimens from Vlor is doubtful since the ssp. ljubetnesis is normally treated as
local endemic of the Shar Planina Mts.
? species group cardioderus
? Trechus irenis Csiki, 1912 [= cardioderus balcanicus Jeannel, 1927]
References. Trechus cardioderus balcanicus: Tabakovi-Toi, 1988: 137.
Notes. Tabakovi-Toi (1988) noted the range of this taxon reaching Albania. Due to lack of
reliable records from the country the species is treated as questionable for Albania.
Species group pulchellus
40.6. Trechus merditanus Apfelbeck, 1906
References. Trechus merditanus: Apfelbeck, 1906: 1163 (Merdita: Mal i Shenjt Mt. and Zep
Mt., loc. typ.); Apfelbeck, 1907a: 498-499, 503 (Mal i Shenjt Mt.: top region, 1500 m); Jeannel,
1927: 539-540 (Merdita: Mal i Shenjt Mt. and Zep Mt.); Pawlowski, 1972: 312-314; Casale &
Laneyrie, 1983: 151; Moravec et al., 2003: 336.
Material studied. Merdita M. Scheit / merditanus n. sp. Apfelb., 2 s. probably syntypes (NMW).
Notes. According to Jeannel (1927), T. merditanus is a vicarious species of T. priapus.
40.7. Trechus priapus K. Daniel, 1902
40.7.1. Trechus priapus priapus K. Daniel, 1902
References. Trechus priapus: Meixner, 1939: 309; Pawlowski, 1973: 249; Hieke & Wrase, 1988:
38; Nonveiller et al., 1994: 9; Guorguiev & Guorguiev, 1995: 77; Guorguiev et al., 1997: 10.
Trechus priapus priapus: Jeannel, 1927: 537 (North Albanian Alps; Gjalic Mt.); Moravec et al.,
2003: 338.
Material studied. Alban. Exped. Gjalica Ljums 17.-26.VI18, 2 s. (NMW).
Additional material included (studied A. Vigna Taglianti). Tropoia, Kollats Mt., Qafa-eRubs, 2000 m, 8.VII.1993, 2 m., 1 f., L. Latella leg. (cAVT).
Notes. Except the two specimens above, in MNHUB I have seen additionally nearly 800 s.,
which remained unregistered.
86
87
Casale & Laneyrie, 1982: 153. Trechus kobingeri bulgaricus: Guorguiev et al., 1997: 10. Trechus
kobingeri kobingeri: Moravec et al., 2003: 333.
Material studied. Maja e Jezerce Albania sept., 1 m. (NMW); Cucale / Albania leg.
Bischoff 1936, 1 s. (MNHUB).
Notes. Trchus kobingeri bulgaricus Pawlowski, 1972 is a junior synonym of T. kobingeri pawlowskianus
P. Moravec & Lompe, 2003, and its record for Albania is erroneous. This race is known only
from a limited region in Western Bulgaria.
# Trechus obtusiusculus Ganglbauer, 1889
References. Trechus obtusiusculus: Nonveiller et al., 1994: 9.
Notes. The species is recorded with detailed records only from Bosnia Herzegovina and
Montenegro. The data for Albania is too doubtful, so that it is removed from the species list.
40.12. Trechus ravasinianus Lorenz, 1998 [= ravasinii Jeannel, 1925]
References. Trechus ravasinii: Jeannel, 1925: 24-25 (Tomor Mt.: Kulmak, Toik, Abbas Ali, loc.
typ.); Jeannel, 1927: 570-571 (Tomor Mt.: Abbas Ali, Kulmak, Toik); Casale & Laneyrie, 1982:
153. Trechus ravasinianus: Moravec et al., 2003: 339.
Material studied. Tomor: Tomor Albania mer. / lg. Winkler Mai 1931, 1 f. (HNHM);
Tomor: Kulmak Albania mer. / lg. Winkler Mai 1931, 12 s. (NMW).
Notes. Except the localities mentioned above, in MNHUB I have seen more than 100 s.,
which remained unregistered.
! Trechini gen. indet.
! Trechini gen. & sp. indet.
Material included (studied A. Vigna Taglianti). Tropoia, Kollats Mt., Mali-i-That Mt., cave
Shpella e Haxise, 8.VII.1993, rests of 1 s., L. Latella leg. (cAVT).
Notes. According to Vigna Taglianti (personal communication) the parts found in the cave
Shpella e Haxise may belong to an unknown genus close to Pheggomisetes Knirsch, 1923.
88
89
Notes. Murraj (1968: 33) cited this species from two localities (Tiran; Tomor Mt.). Among
the material loaned from MNST, I found out 9 s. from the second place having the same label
data as the material published by Murraj as Chlaenius nigricornis. Actually, all the nine specimens
refer to Bembidion dalmatinum.
43.3. Chlaenius nitidulus (Schrank, 1781)
References. Chlaenius nitidulus: Hieke & Wrase, 1988: 149; Kirschenhofer, 2003: 350.
Material studied. Maja Qori 2050 m, Albania, 1 s. (NMW); Sauk, 5.III.1961, 1 m. (MNST).
Notes. First detailed records for Albania.
43.4. Chlaenius terminatus Dejean, 1826
References. Chlaenius terminatus: Schatzmayr, 1943b: 100 (Stan-Karbunar; Skrofotina; Vlor).
Material studied. Oboti Bojana, 3 s. (HNHM).
Notes. Kirschenhofer (2003: 350) has omitted the species for Albania. Here it is confirmed
for the country.
43.5. Chlaenius tristis (Schaller, 1783)
43.5.1. Chlaenius tristis tristis (Schaller, 1783)
References. Chlaenius tristis: Hieke & Wrase, 1988: 149. Chlaenius tristis tristis: Kirschenhofer, 2003: 350.
Material studied. Oboti Bojana, 5 s. (HNHM); Tiran / Albania leg. Bischoff 1936, 1 s.
(MNHUB).
Notes. First detailed records for Albania.
43.6. Chlaenius vestitus (Paykull, 1790)
References. Chlaenius vestitus: Csiki, 1940: 219 (Kula Ljums); Schatzmayr, 1943b: 100 (StanKarbunar); Kirschenhofer, 2003: 351.
Material studied. Bojana Oboti, Alb., 1 s. (NMW); Lumi Dukatit Alban. mer., 4 s. (NMW);
Maje Tartarit Albania mer., 1 s. (NMW); Albania Kula Lums / Akad. Balk. Exp. Csiki 1918,
1 s. (HNHM); Alban. Exped. Kula Ljums 14.20.VII.18, 1 s. (NMW); Elbasan, Alb. 15.IX.18 Dr.
H. Priesner, 4 s. (NMW); Alessio, Alb. Matzenauer, 2 s. (NMW); Miloti, Alb. Matzenauer, 1 s.
(NMW); 40 s. from different localities (MNHUB); Librazhd, Prenjas, 750 m, 7.V.1995, 1 s., S.
Golovach, B. Petrov, P. Stoev leg. (NMNHS); Drina River, 3.VI.1995, 1 s., V. Sakalain leg. (NMNHS).
Additional material included (studied A. Vigna Taglianti). Humelic (between Tepelen
and Gjirokastr), 350 m, 23.IV.2003, 2 f., P. Crucitti & C. Cavalieri leg. (cAVT).
Subgenus Chlaenites Motschulsky, 1860
43.7. Chlaenius spoliatus (P. Rossi, 1792)
43.7.1. Chlaenius spoliatus spoliatus (P. Rossi, 1792)
References. Chlaenius spoliatus: Schatzmayr, 1943b: 99 (Durrs); Hieke & Wrase, 1988: 147.
Chlaenius spoliatus spoliatus: Kirschenhofer, 2003: 351.
90
91
Bischoff 1936, 1 s. (MNHUB); Ivan Geb Korce / Albania 1936 leg. Ramadan coll. Bischoff ,
1 s. (MNHUB); Tiran, 27.IX.1958, 1 m. (MNST); Tiran, 10.V.1962, 1 m. (MNST).
Subgenus Epomis Bonelli, 1810
43.11. Chlaenius circumscriptus (Duftschmid, 1812)
References. Chlaenius circumscriptus: Kirschenhofer, 2003: 353.
Material studied. Gjormi / Albania leg. Bischoff 1935, 1 s. (MNHUB); Liqen i Butrintes
/ Albania leg. Bischoff 1936, 1 s. (MNHUB).
Notes. First detailed records for Albania.
43.12. Chlaenius dejeani (Dejean, 1831)
References. Chlaenius dejeani: Apfelbeck, 1904: 154 (Vlor); Schatzmayr, 1943b: 99 (Lushnje);
Hieke & Wrase, 1988: 147. Epomis dejeani: Apfelbeck, 1909: 298 (Vlor).
Material studied. Gjormi / Albania leg. Bischoff 1935, 1 s. (MNHUB); Murthi / Albania leg.
Bischoff 1936, 2 s. (MNHUB); Sukthi bis Gurse / Albania leg. Bischoff 1938, 1 s. (MNHUB).
Notes. Kirschenhofer (2003: 353) has omitted the species for Albania. Here it is confirmed
for the country.
Subgenus Trichochlaenius Seidlitz, 1887
43.13. Chlaenius aeneocephalus Dejean, 1826
43.13.1. Chlaenius aeneocephalus aeneocephalus Dejean, 1826
References. Chlaenius aeneocephalus: Apfelbeck, 1904: 156 (Vlor); Apfelbeck, 1909: 298 (Vlor);
Hieke & Wrase, 1988: 148.
Notes. Kirschenhofer (2003: 355) has omitted the species for Albania, so that it needs of
further confirmation for the country.
93
95
Material studied. Albania Avlona v. Oertzen, 1 s. (MNHUB); Brataj Ramica Albania mer.
/ Winkler, Lona, Bischoff, V.1931, 1 s. (NMW); M. e Tartarit juni 1932 / Albania leg.
Bischoff 1932, 1 s. (MNHUB); Tiran / Albania leg. Bischoff 1935, 1 s. (MNHUB);
Borshne / Albania leg. Bischoff 1940, 1 s. (MNHUB); Borsh, 15-28.V.1961, 1 f. (MNST);
Ljukove, 100 m, 5.VI.1995, 1 m., G. Blagoev leg. (NMNHS).
Additional material included (studied S. Hristovski). Dhermi, 200 m, 12.VII.2006, 2 s., M.
Komnenov leg. (cSH).
96
97
98
99
i Shenjit / leg. Bischoff Albania 1934, 2 s. (MNHUB); Okol Shkalla / Albania leg.
Bischoff 1935, 1 s. (MNHUB); Shklzn / Albania leg. Bischoff 1935, 8 s. (MNHUB);
AL (Tirana) 8 km N Psene. E Tirana 1450 m. 30.IV.1993 leg. H. + S. Peks, 2 m. (MNHUB);
North Albanian Alps, Bog, 1000-1100 m, 5-9.VI.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS);
Cafasan, 1230 m, 27.VII.1993, 1 s., P. Stoev & D. Zaprjanova leg. (NMNHS).
56.5. Harpalus rufipalpis Sturm, 1818
56.5.1. Harpalus rufipalpis rufipalpis Sturm, 1818 [= rufitarsis (Duftschmid, 1812)]
References. Harpalus rufitarsis: Apfelbeck, 1904: 196 (Berat); Apfelbeck, 1907a: 499 (Mal i
Shenjt Mt.: top region, 1500 m); Apfelbeck, 1909: 298 (Berat); Hieke et al., 1986: 188 (Biz
near Shngjergji); Hieke & Wrase, 1988: 138. Harpalus rufipalpis rufipalpis: Kataev & Wrase,
2003: 381.
Material studied. Mal i Dajtit / Albania leg. Bischoff 1932, 2 s. (MNHUB); Mal i Shenjit
/ leg. Bischoff Albania 1934, 6 s. (MNHUB); Gjallica / Albania leg. Bischoff 1935, 1 s.
(MNHUB); Cucale / Albania leg. Bischoff 1936, 4 s. (MNHUB); Maranaj / Albania leg.
Bischoff 1937, 2 s. (MNHUB); Albania Sauk / 27.VII.1959 leg. Xh. Murraj, 3 s. (HNHM).
Notes. In addition to the material studied above, in NMW I have seen more specimens, which
remained unregistered.
56.6. Harpalus sulphuripes Germar, 1824
56.6.1. Harpalus sulphuripes sulphuripes Germar, 1824
References. Harpalus sulphuripes: Apfelbeck, 1904: 196 (Vlor); Anonymous, 1907: 284;
Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Apfelbeck, 1909: 298 (Vlor);
Schauberger, 1926: 32 (Vlor; Pasha Liman); Hieke et al., 1986: 188 (Tomor Mt.: Abbas
Ali, 2401 m); Hieke & Wrase, 1988: 139. Harpalus sulphuripes sulphuripes: Kataev & Wrase,
2003: 383.
Material studied. More than 50 s. from different localities (MNHUB); North Albanian Alps,
Bog, 1000-1100 m, 5-9.VI.1993, 2 s., P. Beron, B. Petrov leg. (NMNHS); Saranda District,
Ljukove, 5.V.1995, 1 s., P. Stoev leg. (NMNHS); Gjirokastra, 6.V.1994, 1 s., P. Stoev leg.
(NMNHS); 12 km NE Erseka, 900 m, 12.V.1995, 1 s., L. Penev leg. (NMNHS).
Additional material included (studied S. Hristovski). Dhermi, 200 m, 12.VII.2006, 1 s., M.
Komnenov leg. (cSH).
Species group atratus
56.7. Harpalus atratus Latreille, 1804 [= subsinuatus (Duftschmid, 1812)]
References. Harpalus atratus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);
Hieke et al., 1986: 187 (Mal i Dajtit Mt.: Shkall Prisk); Hieke & Wrase, 1988: 137; Kataev &
Wrase, 2003: 373. Harpalus atratus var. subsinuatus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top
region, 1500 m).
Material studied. Albania Zalle-Dardha / 23.V.1958 leg. Xh. Murraj, 1 s. (HNHM); more
than 60 s. from different localities (MNHUB).
100
101
Golovach, B. Petrov, P. Stoev leg. (NMNHS); Llogora Pass, 1025 m, 6.VI.1995, 3 s., G. Blagoev
leg. (NMNHS).
Additional material included (studied S. Hristovski). Hotel Edva at Radhime (Vlor
District), 8-15.VII.2006, 1 s., leg. M. Komnenov (cSH).
Notes. In addition to the material studied above, in NMW and ZNHU I have seen more than
130 s. from different localities, which remained unregistered.
Species group pumilus
[Harpalus picipennis (Duftschmid, 1812)]
References. Harpalus picipennis: Apfelbeck, 1904: 200; Hieke & Wrase, 1988: 141; Kataev &
Wrase, 2003: 380.
Notes. Hieke & Wrase (1988) noted the species for Albania following Apfelbecks record
Prisren (Apfelbeck, 1904: 200). In fact this place is situated in the Kosovo Region of Serbia.
Most likely, Kataev & Wrase (2003) have used the same sources. For the time being the species
is treated here as probable but not yet proven for Albania (having in mind recent data from
the north parts of Republic of Macedonia, unpublished information).
56.11. Harpalus pumilus Sturm, 1818 [= vernalis Fabricius, 1801]
References. Harpalus vernalis: Mller, 1931: 69; Hieke & Wrase, 1988: 140; Kataev & Wrase,
2003: 380.
Material studied. Mal i Shenjit / leg. Bischoff Albania 1934, 1 s. (MNHUB).
Notes. First detailed record for Albania.
Species group anxius
56.12. Harpalus anxius (Duftschmid, 1812)
References. Harpalus anxius: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);
Kataev & Wrase, 2003: 373.
Material studied. Latif Cucali, 1 s. (NMW); Merdita Oroshi / Apfelb. Alban. spt.
IV.V.05, 2 s. (NMW); Hotolisht - Librazhd, 300 m, 7.V.1995, 1 m., S. Golovach, B. Petrov, P.
Stoev leg. (NMNHS).
Notes. Hieke & Wrase (1988: 139) noted this species for Albania apparently following the
record Jannina (Apfelbeck, 1904: 199). In fact this locality is situated in Greece.
56.13. Harpalus subcylindricus Dejean, 1829
References. Harpalus subcylindricus: Hieke & Wrase, 1988: 140; Kataev & Wrase, 2003: 382.
Material studied. Skutari / Strasser, 1 s. (NMW); Alban. Exp. 1918 Bicaj 14.15.VI., 1
s. (NMW); Burelle / leg. Bischoff Albania 1934, 1 s. (MNHUB); Rapsh / leg. Bischoff
Albania 1934, 1 s. (MNHUB); Shklzn / Albania leg. Bischoff 1935, 1 s. (MNHUB);
Mal i Dajtit Mt., 10.V.1965, 1 m. (MNST); AL (Berat) Pa ca. 5 km s. Gllav ~ 600 m 27.IV.1993
leg. Heinz u. Sonja Peks, 3 s. (MNHUB); AL (Berat). Pa ca. 5 km S Gllava 600 m. 27.IV.1993
leg. H. + S. Peks, 1 m., 1 f. (MNHUB); North Albanian Alps, Maje Radohimes, 2400-2550
102
m, 29.V.1993, 3 s., P. Beron, B. Petrov leg. (NMNHS); North Albanian Alps, Maje Radohimes,
1000-1100 m, 5.IX.1993, 2 s., P. Beron, B. Petrov leg. (NMNHS).
Notes. First detailed records for Albania.
Species group flavescens
56.14. Harpalus flavescens (Piller et Mitterpacher, 1783)
Material studied. Elbasan, Alb. 11.IX.18 Dr. H. Priesner, 1 s. (NMW).
Notes. New species for the fauna of Albania.
Species group tardus
56.15. Harpalus albanicus Reitter, 1900
References. Harpalus albanicus: Reitter, 1900: 114 (? Shkodr, loc. typ.); Apfelbeck, 1904: 200
(Shkodr); Rambousek, 1912: 86; Horion, 1941: 230; Roudier, 1949: 176 (Shkodr); Magistretti,
1965: 308; Hieke & Wrase, 1988: 134; Kataev & Wrase, 2003: 372.
Material studied. Skutari / coll. Reitter, 1 f. syntype (HNHM).
Notes. Tschitschrine (1901: 154) expressed doubt about the correct interpretation of the
species type locality Scutari. It may as well represent the present city of skdar located on
the Asian shore of the Bosporus right opposite the peninsula of Istanbul, or the present
Albanian City of Shkodr.
56.16. Harpalus tardus (Panzer, 1796)
References. Harpalus tardus: Apfelbeck, 1904: 198 (Vlor); Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top
region, 1500 m); Apfelbeck, 1909: 298 (Vlor); Hieke & Wrase, 1988:134; Kataev & Wrase, 2003: 383.
Material studied. Latif Oroshi, 1 s. (NMW); Maranaj / Apfelb. Alban. spt. IV.V.05, 2
s. (NMW); Alban. Exped. Kula Ljums 18-28.V.1918, 1 s. (NMW).
Species group latus
56.17. Harpalus latus (Linnaeus, 1758)
References. Harpalus latus: Kataev & Wrase, 2003: 378.
Material studied. Rapsa 10-18.V. / Alban. montenegr. Grenze, Penther14, 1 s. (NMW); North
Albanian Alps, Maje Redohimes, 2400-2550 m, 29.V.1993, 1 s., P. Beron & P. Stoev leg. (NMNHS).
Notes. First detailed records for Albania.
56.18. Harpalus solitaris Dejean, 1829
References. Harpalus solitaris: Kataev & Wrase, 2003: 382.
Material studied. North Albanian Alps, Bog, 1800-1900 m, 20-23.VI.1993, 1 f., P. Beron, B.
Petrov leg. (NMNHS).
Notes. First detailed record for Albania.
COLEOPTERA: CARABIDAE: HARPALINAE
103
105
Additional material included (studied S. Hristovski, A. Vigna Taglianti and D.W. Wrase).
Tiran, XI.58, 1 s. (collector illegible) (cWR); Tiran District, Kffaba-Pa, Sdseite, 500 m,
29.IV.1993, 5 s., H. & S. Peks leg. (cWR); Korc District, Mai i That Mt., Zwezde-Sattel, 10001100 m, 4.V.1993, 6 s., Heinz & Peks leg. (cWR); Fier, 4.VII.1993, 1 f., L. Latella leg. (cAVT);
Tiran, botanical garden, 4.VI.2002, 1 f., A. Paparistos leg. (cAVT); Tiran, Mal i Dajtit Mt.,
7.VI.2002, 1 f., A. Paparistos leg. (cAVT); Qesarat, near Memaliaj (Tepelen District), valley of
Vijos River, 300-360 m, 18.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT); Hotel Edva
at Radhime (Vlor District), 8-15.VII.2006, 1 s., leg. M. Komnenov (cSH).
Species group dispar
56.24. Harpalus punctatostriatus Dejean, 1829
References. Harpalus punctatostriatus: Apfelbeck, 1909: 298 (Vlor; Sop); Schatzmayr, 1943b:
103 (Lushnje); Hieke & Wrase, 1988: 128; Kataev & Wrase, 2003: 380.
Material studied. Albanien Avlona v. Oertzen, 1 s. (MNHUB); Albania 1918, 1 s. (HNHM);
Lum i Tiranes / Albania leg. Bischoff 1932, 1 s. (MNHUB); Murthi / Albania leg.
Bischoff 1936, 1 s. (MNHUB).
56.25. Harpalus pygmaeus Dejean, 1829
References. Harpalus pygmaeus: Apfelbeck, 1904: 195 (Vlor; Berat); Apfelbeck, 1909: 298
(Berat); Hieke et al., 1986: 188 (Ib near Krraba); Hieke & Wrase, 1988: 139; Kataev & Wrase,
2003: 380.
Material studied. Albania 1918, 1 s. (HNHM); Skutari, Alb. 4.X.18 Dr. H. Priesner, 10 s.
(NMW); Mal i Dajtit juni / Albania leg. Bischoff 1932, 1 s. (MNHUB); Ura Erzen /
Albania leg. Bischoff 1932, 1 s. (MNHUB); Lum i Tiranes / Albania leg. Bischoff 1933,
1 s. (MNHUB); Tiran / leg. Bischoff Albania 1934, 2 s. (MNHUB); Tiran / Albania
leg. Bischoff 1935, 1 s. (MNHUB); Albania Kamze / 23.III.1958 leg. Xh. Murraj, 1 s.
(HNHM); Albania Dajti / 6.V.1959 leg. Xh. Murraj, 1 s. (HNHM); Sauk, 9.VII.1961, 3 s.
(MNST); Tiran, 10.V.1962, 1 s. (MNST); Tiran Park, 22-31.X.1997, 2 s., R. Ward leg. (CMNH);
Tiran Park, 16-21.II.1998, 2 s., R. Ward leg. (CMNH).
Notes. The specimens from the locality Tiran Park are somewhat bigger (7.1-7.4 mm),
compared with specimens that inhabit Central Europe (cfr. Hrka, 1996).
Species group hospes
56.26. Harpalus hospes Sturm, 1818
56.26.1. Harpalus hospes hospes Sturm, 1818
Material studied. AL (Korc) Qaf e Qarrit 1100 m 4.V.1993, leg. Heinz u. S. Peks, 9 s.
(MNHUB).
Notes. New species for the fauna of Albania.
106
107
Bog, 1000-1100 m, 5-9.VI.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS); Rhesen District,
Merkurth, 11.VI.1993, 3 s., P. Beron & B. Petrov leg. (NMNHS).
Additional material included (studied S. Hristovski and A. Vigna Taglianti). Tropoia, valley
of Valbona River, 900 m, 5.VII.1993, 1 m., L. Latella leg. (cAVT); Durrs, Plepa, 15.VIII.1995,
1 m., 2 f., D. Antonelli leg. (cAVT); Tiran, Mal i Dajtit Mt., 7.VI.2002, 1 f., A. Paparistos leg.
(cAVT); M. Polisit, 8.VII.2004, 1 f., A. Paparistos leg. (cAVT); Malj Cojepit 1300-1400,
17.VII.2004, 1 m., A. Paparistos leg. (cAVT); Hotel Edva at Radhime (Vlor District), 815.VII.2006, 3 s., leg. M. Komnenov (cSH).
Notes. Besides the material studied above, in NMW and MNHUB I have seen more specimens,
which remained unregistered.
109
Sonja Peks, 1 m. (MNHUB); North Albanian Alps, Maje Redohimes, 2400-2550 m, 29.V.1993,
1 s., P. Beron & B. Petrov leg. (NMNHS); Rhesen District, Merkurth, 11.VI.1993, 1 s., P. Beron
& B. Petrov leg. (NMNHS); Saranda District, Lyukove, 5.V.1994, 2 s., P. Stoev leg. (NMNHS);
Saranda District, Qeparo, 4-5.V.1995, 1 s., P. Stoev leg. (NMNHS); Saranda District, Ljukove,
5.V.1995, 1 s., P. Stoev leg. (NMNHS); Dukati Mt., 30 km S Vlor, 450 m, 11.V.1995, 1 s., L.
Penev leg. (NMNHS); Kiari Pass, 1160 m, 1.VI.1995, 5 s., G. Blagoev leg. (NMNHS); Tiran
Park, 15.II.1998, 1 s., from grass and soil around tree trunks, R. Ward leg. (CMNH); Tiran
District, Petrela, 28.XI.2000, 1 s., A. Zhalov leg. (NMNHS).
Notes. In the region of Gllav O. azureus was found to lives sympatrically with O. similis.
57.3. Ophonus cribricollis (Dejean, 1829)
References. Harpalus cribricollis: Apfelbeck, 1904: 184-185 (Vlor; Berat); Csiki, 1940: 219
(Shklzen Mt.; Kula Ljums; Gostil). Ophonus cribricollis: Apfelbeck, 1907a: 499 (Mal i Shenjt
Mt.: top region, 1500 m); Apfelbeck, 1909: 298 (Vlor; Berat); Hieke & Wrase, 1988: 126;
Kataev & Wrase, 2003: 389.
Material studied. Merdita Oroshi, 3 s. (NMW); Skutari Mesi / Apfelb. Alban. spt.
IV.V.05, 3 s. (NMW); Pasha-Liman Albanien V.1908 H. Hopp, 1 s. (MNHUB); Alban.
Exped. Kula Ljums 18-28.V.1918, 3 s. (NMW); Devoll-Tal juni 1932 / Albania leg. Bischoff
1932, 1 s. (MNHUB); Krabe / Albania leg. Bischoff 1932, 1 s. (MNHUB); Tepelene
juni 1932 / Albania leg. Bischoff 1932, 1 s. (MNHUB); Tiran / leg. Bischoff Albania
1934, 1 s. (MNHUB); Mal i Dajtit / Albania leg. Bischoff 1936, 1 s. (MNHUB); Dukati
Mt., 30 km S Vlor, 450 m, 11.V.1995, 1 s., L. Penev leg. (NMNHS).
Additional material included (studied A. Vigna Taglianti). Vasjar (Tepelen District),
18.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT).
Notes. In addition to the material studied above, in HNHM the present author has seen more
specimens from the localities Kula Lums, Delvine, Sklsen and Mont Peklen. Sciaky
(1987: 66) cited the locality timlje for Albania, but actually this place is situated in the
Kosovo Region of Serbia.
57.4. Ophonus similis (Dejean, 1829)
Material studied. Hani Grabom Mai / Alban. montenegr. Grenze Alban. Exped. 1918
Bicaj 14.15.VI., 1 s. (NMW); AL (Berat). Pa ca. 5 km S Gllava 600 m, 27.IV.1993 leg. H. +
S. Peks, 2 f. (MNHUB).
Notes. New species for the fauna of Albania.
57.5. Ophonus subquadratus (Dejean, 1829) [= meridionalis (Dejean, 1829)]
References. Ophonus meridionalis: Apfelbeck, 1909: 298 (Berat; Gramata).
Material examined. Liqen i Butrintes vivari / leg. Bischoff Albania 1934, 1 s. (MNHUB);
Maja e Konispol Seraqin / Albania leg. Bischoff 1936, 1 s. (MNHUB).
Additional material included (studied S. Hristovski). Dhermi, 200 m, 12.VII.2006, 1 s., M.
Komnenov leg. (cSH).
Notes. Kataev & Wrase (2003: 389) have omitted the species for Albania. Here it is confirmed
for the country.
110
111
113
Notes. Hieke & Wrase (1988: 142) noted that the Apfelbecks figures for P. planicollis (Apfelbeck,
1904: 187) refer to P. laeviceps.
115
Notes. All older references and collection records in the museums about A. puncticollis (Coquerel,
1859) from Albania refer to A. paludicola.
60.8. Acupalpus parvulus (Sturm, 1825) [= dorsalis (Fabricius, 1787)]
References. Acupalpus dorsalis: Apfelbeck, 1904: 206 (Velipoja; Vlor); Apfelbeck, 1909: 298
(Vlor); Csiki, 1940: 221 (Kula Ljums); Hieke & Wrase, 1988: 117.
Material studied. Albania Kula Lums / Akad. Balk. Exp. Csiki 1918, 1 s. (HNHM);
Tiran Park, 8.II.1998 / 1.III.1998, 3 s., from grass and soil around tree trunks, R. Ward leg.
(CMNH).
Notes. Jaeger & Kataev (2003: 399) have omitted the species for Albania. Here it is confirmed
for the country. According to B. Jaeger and D.W. Wrase (personal communication) the records
from Albania may concern aberrant specimens of A. maculatus without darkened humeral
macula, but the only certainty for a correct identification is the study of male genitalia.
60.9. Acupalpus planicollis (Schaum, 1857)
References. Acupalpus planicollis: Apfelbeck, 1904: 205 (Velipoja); Apfelbeck, 1909: 298 (VlorMalket); Hieke & Wrase, 1988: 116; Guorguiev & Guorguiev, 1995: 177; Guorguiev et al.,
1997: 20; Jaeger & Kataev, 2003: 399.
Material studied. Liqen i Butrintes vivari / leg. Bischoff Albani 1934, 1 s. (MNHUB).
Additional material included (studied B. Jaeger). Velipoja, Mustjabeg, 1 s. (ZSM).
60.10. Acupalpus suturalis Dejean, 1829
60.10.1. Acupalpus suturalis suturalis Dejean, 1829
References. Acupalpus suturalis: Hieke & Wrase, 1988: 117; Jaeger, 1992: 226. Acupalpus suturalis
suturalis: Jaeger, 1999: 962; Jaeger & Kataev, 2003: 399.
Material studied. Oboti Bojana, 1 s. (HNHM); Tiran Park, 22-31.X.1997, 1 s., R. Ward
leg. (CMNH); Tiran Park, 22.II.1998, 3 s., R. Ward leg. (CMNH).
Additional material included (studied B. Jaeger). Scutari, VIII. 1941, Capra, 1 s. (MHNG).
Notes. First detailed records for Albania.
117
118
119
120
121
122
123
Material studied. Lum i Tiranes / Albania leg. Bischoff 1935, 1 s. (MNHUB); Gjallica
/ Albania leg. Bischoff 1936, 1 s. (MNHUB); Kara Alibey / Albania leg. Bischoff 1936,
4 s. (MNHUB); Librazhd Prenjas, 750 m, 7.V.1995, 1 s., L. Penev leg. (NMNHS).
68.3. Microlestes luctuosus Holdhaus, 1904
68.3.1. Microlestes luctuosus luctuosus Holdhaus, 1904
References. Microlestes exilis luctuosus: Holdhaus, 1904: 331 (Berat). Microlestes luctuosus: Hieke &
Wrase, 1988: 159. Microlestes apterus: Wrase, 1991: 18.
Material studied. Albanien Avlona v. Oertzen, 2 s. (MNHUB); Kjori / Albania leg.
Bischoff 1932, 2 s. (MNHUB); Murrsi - Kara Alibey / leg. Bischoff Albania 1934, 1 s.
(MNHUB); Shklzn / Albania leg. Bischoff 1936, 1 s. (MNHUB).
Notes. According to Wrase (1991) the material cited as M. exilis Schmidt-Gbel, 1846 and M.
exilis luctuosus (Apfelbeck, 1904: 330-331) may concern either M. apterus or M. luctuosus. I refer
it provisionally to the latter species, which was proved to inhabit the country. Kabak (2003:
423) has omitted or M. luctuosus for Albania. Here, the species is confirmed for the country.
68.4. Microlestes maurus (Sturm, 1827)
68.4.1. Microlestes maurus maurus (Sturm, 1827)
References. Microlestes maurus: Holdhaus, 1904: 332 (Velipoja; Vlor); Hieke & Wrase, 1988:
159; Kabak, 2003: 423.
68.5. Microlestes minutulus (Goeze, 1777)
References. Blechrus minutulus: Apfelbeck, 1909: 298 (Vlor; Berat).
Material studied. Tomor-Buranj juli 1932 / Albania leg. Bischoff 1932, 1 s. (MNHUB);
Mal i Krujes / leg. Bischoff Albania 1934, 1 s. (MNHUB); Murthi / Albania leg.
Bischoff 1936, 1 s. (MNHUB).
Notes. Kabak (2003: 423) has omitted the species for Albania. Here it is confirmed for the
country.
68.6. Microlestes negrita (Wollaston, 1854)
68.6.1. Microlestes negrita apfelbecki Holdhaus, 1904
References. Microlestes minutulus apfelbecki: Holdhaus, 1904: 330 (Velipoja). Microlestes negrita:
Hieke & Wrase, 1988: 158.
Material studied. Cafa Liqenit Iballe / Albania leg. Mueller 1939, 1 s. (MNHUB).
Notes. Kabak (2003: 423) has omitted the species for Albania. Here it is confirmed for the
country.
68.7. Microlestes plagiatus (Duftschmid, 1812)
References. Microlestes plagiatus: Apfelbeck, 1907a: 495 (Buen River); Csiki, 1940: 221 (Korab Mt.).
Notes. Kabak (2003: 423) has omitted the species for Albania. Actually it needs further
confirmation for the country.
124
125
Notes. The mention of Hieke & Wrase (1988: 153) for Albania is apparently result of a
following the record Prisren (Apfelbeck, 1904: 323). However, this locality is situated in the
Kosovo Region of Serbia. Kabak (2003: 427) has most likely followed one of these sources,
too. Since there is no previous reliable record from Albania, L. cyanocephala is treated here as
new species for the fauna of Albania.
Subgenus Lebia s.str.
71.2. Lebia cruxminor (Linnaeus, 1758)
71.2.1. Lebia cruxminor cruxminor (Linnaeus, 1758)
References. Lebia cruxminor: Hieke et al., 1986: 190 (North Albanian Alps near Theth; Kula e
Lums near Kuks); Hieke & Wrase, 1988: 153.
Material studied. Skutari Drin, 1 s. (HNHM); Alban. Exp. 1918 Gjalica Ljums 17-26.VI.,
1 s. (NMW); Cucale Geb. / Albania leg. Bischoff 1931, 1 s. (MNHUB); Kjori / Albania
leg. Bischoff 1932, 1 s. (MNHUB); Maja e Tartarit / Albania leg. Bischoff 1932, 1 s.
(MNHUB); Mal i Dajtit / Albania leg. Bischoff 1936, 1 s. (MNHUB); Albanien-Exp.
DEI Nordalban. Thethi, 1.-4.VIII.61, 1 s. (MNHUB).
Notes. Kabak (2003: 428) has omitted the species for Albania. Here L. cruxminor is confirmed
for the country.
71.3. Lebia humeralis Dejean, 1825
References. Lebia humeralis: Hieke & Wrase, 1988: 154.
Material studied. Albania Djakova / Akad. Balk. Exp. Csiki 1917, 1 s. (HNHM); Alban.
Exped. Gjalica Ljums 17-26.VI.18, 1 s. (NMW); Lum i Tirans / Albania leg. Bischoff
1936, 1 s. (MNHUB); Mal i Dajtit / Albania leg. Bischoff 1936, 1 s. (MNHUB).
Notes. Kabak (2003: 429) has omitted the species for Albania. Here it is confirmed for the
country with first detailed records.
71.4. Lebia scapularis (Geoffroy, 1785)
71.4.1. Lebia scapularis scapularis (Geoffroy, 1785)
References. Lebia scapularis: Hieke & Wrase, 1988: 154.
Material studied. Serand Piqeros / Albania leg. Bischoff 1932, 1 s. (MNHUB).
Notes. Kabak (2003: 429) has omitted the species for Albania. Here L. scapularis is confirmed
for the country with a first detailed record.
71.5. Lebia trimaculata (Villers, 1789)
Material studied. 12 km NE Erseka, 900 m, 12.V.1995, 2 s., L. Penev leg. (NMNHS).
Notes. New species for the fauna of Albania.
126
127
129
131
133
is situated in the Kosovo Region of Serbia. Bousquet (2003e: 451) has most likely followed
one of these sources, too. Since there is no previous reliable record from Albania, A. muelleri
is treated here as new species for the fauna of Albania.
81.10. Agonum permoestum Puel, 1938
References. Agonum permoestum: Puel, 1938: 194 (Velipoja); Schmidt, 1994: 25 (Buen River
near Shkodr; Bushnesh; Mamurash; Murrsi-Kara Alibey; Buen River near Oboti; Rushkull;
Liqen i Shkodrs; Sukthi near Gurse); Bousquet, 2003e: 451.
Material studied. Scutari l. See-Ufer, 1 s. (HNHM); Skutari l. See-Ufer / Apfelb. Alban.
spt. IV.V.05, 8 s. (NMW); Oboti Bojana, 1 s. (HNHM).
81.11. Agonum sexpunctatum (Linnaeus, 1758)
References. Agonum sexpunctatum: Csiki, 1940: 218 (Korab Mt.); Hieke et al., 1986: 189 (Lur
e Vjetr near Lura Mt.); Hieke & Wrase, 1988: 84.
Material studied. Alban. Vermosa Penther, 1 s. (NMW); Ricavac 1300 m, 25.VI.-2.VII. /
Alban. montenegr. Grenze, Penther14, 2 s. (NMW); Albania Tirana / 3.VI.1958 leg. Xh.
Murraj, 3 s. (HNHM).
Notes. Bousquet (2003e: 451) has omitted the species for Albania. Here it is confirmed for
the country.
81.12. Agonum sordidum Dejean, 1828
81.12.1. Agonum sordidum sordidum Dejean, 1828
References. Platynus sordidus: Apfelbeck, 1904: 290 (Velipoja; Vlor); Apfelbeck, 1909: 297
(Vlor). Agonum sordidum: Schatzmayr, 1943b: 111 (Stan-Karbunar); Hieke et al., 1986: 189
(Polian near Tomor Mt.); Hieke & Wrase, 1988: 85.
Material studied. Elbasan, Alb. IX Dr. H. Priesner, 1 s. (NMW); Liquen i Butrintes
Vivari / leg. Bischoff Albania 1934, 1 s. (MNHUB); Tiran / Albania leg. Bischoff
1935, 2 s. (MNHUB); Murthi / Albania leg. Bischoff 1936, 1 s. (MNHUB); Sukthi bis
Gurse / Albania leg. Bischoff 1938, 1 s. (MNHUB).
Notes. Bousquet (2003e: 451) has omitted the species for Albania. Here it is confirmed for the country.
81.13. Agonum viridicupreum (Goeze, 1777)
80.13.1. Agonum viridicupreum viridicupreum (Goeze, 1777)
References. Platynus viridicupreus: Apfelbeck, 1904: 290 (Velipoja). Agonum viridicupreum: Hieke
et al., 1986: 189 (Lur e Vjetr near Lura Mt.); Hieke & Wrase, 1988: 85.
Material studied. Murrsi-Kara Alibey / leg. Bischoff Albania 1934, 1 s. (MNHUB);
Tiran / Albania leg. Bischoff 1935, 3 s. (MNHUB); Liquen i Butrintes Vivari / Albania
leg. Bischoff 1936, 1 s. (MNHUB); Bojana Shkoder / Albania leg. Bischoff 1937, 2 s.
(MNHUB); Han-Hot / Albania leg. Bischoff 1937, 1 s. (MNHUB); Sukthi bis Gurse /
Albania leg. Bischoff 1938, 1 s. (MNHUB); Cafa Liqenit Iballe / Albania Mueller leg.
1939, 5 s. (MNHUB); Tiran Park, 11.I.1998 / 1.III.1998 / 8.III.1998, 4 s., shaded lake edge
under leaves and wet, rotten wood, R. Ward leg. (CMNH).
COLEOPTERA: CARABIDAE: HARPALINAE
135
Notes. Bousquet (2003e: 452) has omitted the species for Albania. Here it is confirmed for
the country.
Subgenus Europhilus Chaudoir, 1859
81.14. Agonum antennarium (Duftschmid, 1812)
References. Platynus antennarius: Apfelbeck, 1904: 295 (Velipoja). Agonum antennarium: Hieke
& Wrase, 1988: 87.
Material studied. Mustajbeg Velipoja Albania, 2 s. (MNHUB); Maja Rogamit / Albania
leg. Bischoff 1935, 11 s. (MNHUB); Tiran Park, 22.II.1998 / 1.III.1998 / 8.III.1998, 8 s.,
shaded lake edge under leaves and wet, rotten wood, R. Ward leg. (CMNH).
Notes. Bousquet (2003e: 452) has omitted the species for Albania. Here it is confirmed for the country.
81.15. Agonum thoreyi Dejean, 1828
81.15.1. Agonum thoreyi thoreyi Dejean, 1828
References. Platynus thoreyi: Apfelbeck, 1904: 296 (Velipoja). Agonum thoreyi: Hieke & Wrase,
1988: 87; Bousquet, 2003e: 453.
Crucitti & C. Cavalieri leg. (cAVT); Hotel Edva at Radhime (Vlor District), 8-15.VII.2006,
1 s., leg. M. Komnenov (cSH).
Notes. Bousquet (2003e: 455) has omitted the species for Albania. Here it is confirmed for
the country.
137
139
140
141
Notes. Apfelbeck (1904: 222) described this species basing on a single male from the type
locality Prizren. This place is situated in the Kosovo Region of Serbia. The binomen Molops
transfuga Apfelbeck (Mller, 1930: 124; Mlyn, 1977: 120) is nomen nudum.
87.8. Molops apfelbecki Ganglbauer, 1891
Notes. In the past, M. apfelbecki and M. reiseri have been treated as separate species. Probable
cause for this state is that Apfelbeck (1904: 223), while describing M. reiseri, missed to compare
it with M. apfelbecki. Later, Mlyn (1977) similarly did not compare the aedeagus of both species,
but noticed the resemblance in their female genital structures. The present author has studied
the aedeagus of M. reiseri holdhausi (syntype, NMW), M. reiseri (lectotype, HNHM), M. reiseri
victoris (syntype, NMW; specimens from Koprivnik Mt. and Shklzen Mt., HNHM) and M. apfelbecki
(lectotype, HNHM) and found that M. reiseri holdhausi is different in the shape of the median
lobe from the other three taxa. In fact, the last taxa have identical aedeagus. Therefore, M. reiseri
reiseri and M. apfelbecki apfelbecki are considered as geographical races of single species, and M.
holdhausi as valid species which occurs together with M. apfelbecki s.l.
87.8.1. Molops apfelbecki victoris J. Mller, 1918
References. Molops victoris: Mller, 1918a: 1-3, 6 (Albania / Montenegro border: North Albanian
Alps, 1300-2000 m, loc. typ.). Molops reiseri victoris: Mlyn, 1977: 114-115 (North Albanian
Alps; Shklzen Mt.).
Material studied. Prokletija 1300-2000 m / Alban. montenegr. Grenze, Penther14 /
victoris n. det. J. Mller, 2 m. syntypes of Molops victoris (NMW); Albania Sklsen / Akad.
Balk. Exp. Csiki 1917, 1 m. with extracted penis (HNHM); Maja Peis / Albania leg. Bischoff
1935, 1 s. (MNHUB); Shklzn / Albania leg. Bischoff 1935, 5 s. (MNHUB); Shklzn
/ Albania leg. Bischoff 1936, 22 s. (MNHUB); M. Raba Massiv / Albania leg. Bischoff,
Mueller 1939 / Type / Apfelbecki sbsp. Bischoffi m. det. Mller, 1 s. (MNHUB); M. Raba
Massiv / Albania leg. Bischoff, Mueller 1939 / Type, 5 s. (MNHUB); Maja e Dragomirit
/ Albania leg. Bischoff, Mueller 1939 / M. Apfelbecki sbsp. Bischoffi m. det. G. Mller 1940,
1 s. (MNHUB); Maja e Dragomirit / Albania leg. Bischoff, Mueller 1939, 5 s. (MNHUB).
Additional material studied. Bosnia Herzegovina / Montenegro. Apfelbeck Volujak /
Coll. Apfelbeck, 1 m. lectotype and 3 paralectotypes of Molops apfelbecki (HNHM).
Montenegro. Vunaj, Cafa glava, Alban. Penther / reiseri Apf., 1 m. of Molops apfelbecki
victoris (NMW). Serbia (Kosovo Region). Montes Koprivnik / Akad. Balk. Exp. Csiki 1917,
1 m. of Molops apfelbecki victoris (HNHM). Serbia. Reiser 1898 Kopaonik / Coll. Apfelbeck,
1 m. lectotypus and 2 paralectotypes of Molops reiseri (HNHM); Reiser 1899 Kopaonik /
reiseri Apf. B& typus / type, 1 m. of Molops apfelbecki reiseri (NMW); Reiser 1899 Kopaonik
/ reiseri Apf. Serb. turk. Grence / type, 1 m., 1 f. of Molops apfelbecki reiseri (NMW); Reisser
1899 Kopaonik, 1 m., 1 f. of Molops apfelbecki reiseri (MNHUB).
Additional material included (studied A. Vigna Taglianti). Tropoia, Kollats Mt., Qafa-eRubs, 2000 m, 8.VII.1993, 1 f., L. Latella leg. (cAVT).
Notes. Mller (1930: 126) revised the material of Apfelbeck (1918c: 166 sub M. reiseri from
Vunaj, Cafa glava) and determinated it as M. victoris. However, Bousquet (2003f: 474) has omitted
M. reiseri victoris (currently M. apfelbecki victoris, see above) for Albania in spite of the fact that
Mlyn (1977: 115) gave a detailed record. Here the subspecies is confirmed for the country. The
trinomen M. apfelbecki bischoffi G. Mller 1940, found on labels of two specimens, is nomen nudum.
COLEOPTERA: CARABIDAE: HARPALINAE
143
the first is good species - Molops (Molops) holdhausi J. Mller, 1917, stat. rest. for Molops (Molops)
reiseri holdhausi J. Mller, 1917. Bousquet (2003f: 474) has omitted M. holdhausi for Albania
though Mller (1930) gave a detailed record. Here the species is confirmed for the country.
87.11. Molops plurisetosus J. Mller, 1918 [= malissorum Apfelbeck, 1918]
References. Molops apfelbecki plurisetosus: Mller, 1918a: 3-4, 11 (Albania / Montenegro border:
North Albanian Alps, 1200-2000 m, loc. typ.; Albania / Montenegro border: Ricavac, loc. typ.;
Albania: Vermosh, loc. typ.). Molops malissorum: Apfelbeck, 1918c: 160, 167-168 (Montenegro:
North Albanian Alps, Vusanje, loc. typ.; Albania: pastures over Theth). Molops plurisetosus: Mlyn,
1977: 120-121 (North Albanian Alps; Ricavac; Vermosh).
Material studied. Prokletija 1300-2000 m / Alban. montenegr. Grenze, Penther14, 4
syntypes of Molops plurisetosus (NMW); Ricavac Juni / Alban. montenegr. Grenze, Penther14,
1 syntype of Molops plurisetosus (NMW); Vermosa 20-21.VII. / Alban. montenegr. Grenze,
Penther14, 2 syntypes of Molops plurisetosus (NMW); North Albanian Alps, Theth, 800-1200 m,
28.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Maje Redohimes,
2400-2550 m, 29.V.1993, 3 m., 2 f., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps,
Bog, 1800-1900 m, 20-23.VI.1993, 3 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian
Alps, Bog, cave No 25, 23.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS); Shkodr
District, Kozuja, 1800 m, 27.VIII.1996, 1 m., 1 f., T. Ivanova leg. (NMNHS).
Additional material studied. Montenegro. Vunaj, Prokletija-Geb., Albania Dr. Penther /
Molops malissorum n. sp. Apf., 2 syntypes of Molops malissorum (NMW).
Notes. Bousquet (2003f: 474) has omitted the species for Albania. Here it is confirmed for the
country. Currently, the exact publishing dates of the descriptions of M. plurisetosus and M. malissorum
were not found, so the problem with the priority of one of the two names is pending. Each of
the different localities mentioned in the original descriptions of M. plurisetosus (Mller, 1918a)
could be a potential locus typicus, however lectotype designation still has to be done.
Species group rufipes
87.12. Molops osmanilis Apfelbeck, 1904
[Molops osmanilis osmanilis Apfelbeck, 1904]
Notes. The record from Prisren (Apfelbeck, 1904: 229) refers to the Kosovo Region of Serbia,
so perhaps Bousquet (2003f: 473) has followed this source when listing M. osmanilis osmanilis for
Albania. The nominotypical subspecies is treated here as likely to be found in the country.
87.12.1. Molops osmanilis tomoricensis J. Mller, 1923
References. Molops tomoricensis: Mller, 1923: 109-110 (Tomor Mt., alpine zone, loc. typ.). Molops
osmanilis tomoricensis: Mlyn, 1977: 123 (Tomor Mt.: Tomor, Abbas Ali); Bousquet, 2003f: 473.
Material studied. Tomorica Abbas Ali / Albania 1922 Ravasini, Lona, 5 s. (NMW;
MNHUB); Tomor: Kulmak Alban. mer. / Winkler, Lona, Bischoff, V.1931, 1 s. (NMW);
Tomor: Kulmak Albania mer. / Winkler, Lona, Bischoff, V.1931, 1 s. (NMW); Tomor:
Toik Alban. mer. / Winkler, Lona, Bischoff, V.1931, 3 (NMW); Tomor: Tomor Alban.
mer. / Winkler, Lona, Bischoff, V.1931, 7 s. (NMW); more than 90 s. from Tomor Mt.
(MNHUB); Mal i Dajtit Tirane / Albania leg. Bischoff 1931, 1 s. (MNHUB); Logara
COLEOPTERA: CARABIDAE: HARPALINAE
145
juni / Albania leg. Bischoff 1932, 1 s. (MNHUB); Mal i Dajtit Tirane / leg. Bischoff
Albania 1934, 1 s. (MNHUB); Maja e Papingut / Albania 1936 leg. Ramadan coll. Bischoff ,
13 s. (MNHUB); Maja e Dhembelit / Albania 1936 leg. Ramadan coll. Bischoff , 2 s.
(MNHUB); Mal i Krujes / Albania leg. Bischoff 1939, 2 s. (MNHUB); AL (Tirana) 8 km
N psene E Tirana 1450 m. 30.IV.1993 leg. H. & S. Peks, 2 s. (MNHUB).
Notes. All the specimens examined from Albania are put to this race more or less provisionally.
However, for instance specimens from Southwest Albania (localities Logara and Maja e
Dhembelit) have a shape of the pronotum like spp. osmanilis, whereas these from Central
Albania (localities Mal i Dajtit and Mal i Krujes) like ssp. tomoricensis.
87.13. Molops rufipes Chaudoir, 1843
87.13.1. Molops rufipes rufipes Chaudoir, 1843
References. Molops rufipes: Rambousek, 1912: 93; Mlyn, 1977: 124; Hieke, 1981: 81; Hieke &
Wrase, 1988: 82. Molops rufipes rufipes: Mlyn, 1977: 124 (Mal i Dajtit Mt.).
Material studied. Mal i Daitit b. Tirana, Alb. C. / leg. Bischoff 1931 coll. A. Winkler, 2
s. (NMW); Tomor: Tomor Alban. mer., 1 m. (MNHUB); Tomor-Buranj juli 1932 / Albania
leg. Bischoff 1932, 1 m. (MNHUB); more than 50 s. from locality Iballe (MNHUB); more
than 50 s. from locality Mal i Dajtit (MNHUB); Tiran, 26.VII.1965, 1 f. (MNST); Mal i
Dajtit Mt., 20 km E Tiran, 1000 m, 9.V.1995, 4 m., 3 f., L. Penev leg. (NMNHS).
Notes. Bousquet (2003f: 474) has omitted the subspecies for Albania. Here it is confirmed
for the country. The material of Rambousek (1912) was not revised and therefore it is placed
in the nominotypical race provisionally.
# Molops rufipes klisuranus Apfelbeck, 1902
References. Molops klisuranus: Apfelbeck, 1918c: 166 (North Albanian Alps).
Notes. Two wrong records (Mller, 1930: 125-126, revised as M. curtulus s.l.) for this taxon
have been recorded from the Western Balkan (South Serbia, Sandzak: Raka near Novi Pazar,
Apfelbeck, 1904: 222; North Albanian Alps, Apfelbeck, 1918c: 166). Therefore the taxon is
removed from the list of the Albanian Carabidae.
87.13.2. Molops rufipes munelensis Apfelbeck, 1906
References. Molops sturanyi munelensis: Apfelbeck, 1906: 1665-1666 (Munella Mt., loc. typ.);
Mller, 1918a: 9 (Merdita: Munella Mt.). Molops rufipes munelensis: Mlyn, 1977: 128, 130 (Munella
Mt.; Munella Mt.: Oroshi; Berishe); Bousquet, 2003f: 474.
Material studied. Orosi, 5 s. (HNHM); Oroi (Ptr.) Alban. spt. / Munela / Coll. Apfelbeck,
1 lectotype (HNHM); Merdita Munela, 3 paralectotypes (HNHM); Merdita Munela, 9 s. (DEI;
MNHUB); Merdita Munela Apfb., 4 s. (MNHUB); Merdita / Apfb., 2 s. (DEI); Merdita
Munela / Wgth. 1906 Merdita, 21 s. (NMW); Merdita Zebia, 9 s. (NMW); Alban. Exp.18
Pashtrik, 29.V.-4.VI., 9 s. (NMW); Alban. Exp.18 Pashtrik, 4-17.7, 5 s. (NMW); M. Mulecifit
/ Albania leg. Bischoff 1936, 6 s. (MNHUB); Berishe / Albania leg. Mueller 1939, 6 s.
(MNHUB); AL (Shkodar). Pa SE Puke 850 m 2.V.1993 / leg. H. & S. Peks, 1 m. (MNHUB).
[Molops rufipes pseudoosmanilis Mlyn, 1977]
Notes. Since this subspecies was hitherto recorded only from the Macedonian section of the
Korabi Mt., it is expected to live in Albania, too.
146
147
148
149
151
leg. Bischoff 1935, 18 s. (MNHUB); Maja Pejs / Albania leg. Bischoff 1935, 1 s. (MNHUB);
Maja Rogamit / Albania leg. Bischoff 1936, 1 s. (MNHUB); Boge Aufstieg z. Rab /
Albania Mueller leg. 1939, 1 s. (MNHUB); Boge afa Shtegot / Albania Mueller leg. 1939,
1 s. (MNHUB); Maja e Dragomirit / Albania Mueller leg. 1939, 1 s. (MNHUB).
Additional material included (studied A. Vigna Taglianti). Tropoia, valley of Valbona River,
900 m, 15.VII.1993, 1 f., L. Latella leg. (cAVT).
Notes. Bousquet (2003f: 488) has omitted the species for Albania. Here it is confirmed for the country.
Subgenus Feronidius Jeannel, 1942
91.4. Pterostichus melas (Creutzer, 1799)
91.4.1. Pterostichus melas depressus (Dejean, 1828)
References. Pterostichus melas: Apfelbeck, 1904: 265 (Vlor; Velipoja); Apfelbeck, 1909: 297
(Vlor; Berat); Schatzmayr, 1943b: 109 (Kruja; Dukati; Trpan); Hieke et al., 1986: 189 (Tiran;
Borshi near Vlor; Polian near Tomor Mt.). Pterostichus melas depressus: Hieke & Wrase, 1988:
76; Bousquet, 2003f: 495.
Material studied. Mustajbeg Velipoja Albania, 4 s. (HNHM); Sudalbanien Kopstein, 1 s.
(NMW); Skutari Penther, 4 s. (NMW); Hani Grabom mai / Alban. montenegr. Grenze,
Penther14, 1 s. (NMW); Rapsa 10-18.V. / Alban. montenegr. Grenze, Penther14, 1 s. (NMW);
Elbasan, Alb. VIII.-IX.18 Dr. H. Priesner, 1 s. (NMW); Albania Rapsa Dr. Grabowski, 1 s.
(HNHM); Albania Kamze / 23.III.1958 leg. Xh. Murraj, 3 s. (HNHM); Albania Sauk /
16.VI.1958 leg. Xh. Murraj, 1 s. (HNHM); Albania Terbac / 6.VI.1960 leg. Xh. Murraj, 1 s.
(HNHM); Sauk, 9.III.1961, 1 m. (MNST); Tepelen, 1-3.VI.1961, 1 m. (MNST); Rodenj, 2628.VII.1962, 2 m. (MNST); Kurbneshi, VI.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS); Tiran,
Selit e Vogel, 17.IX.1993, 1 s., S. Beshkov leg. (NMNHS); Saranda District, Ljukove, 5.V.1994, 6 s.,
P. Stoev leg. (NMNHS); Tiran, botanical garden, 8.V.1995, 2 s., S. Golovach, B. Petrov, P. Stoev leg.
(NMNHS); Tiran Park, 22-31.X.1997 / 1-9.XI.1997, 5 s., R. Ward leg. (CMNH).
Additional material included (studied A. Vigna Taglianti). Rapsh, 15.V.2002, 1 f., A. Paparistos
leg. (cAVT); Rapsh, Re, 15.V.2002, 1 f., A. Paparistos leg. (cAVT).
Notes. Besides the material studied above, in MNHUB I have seen other more 63 s. from
Albania, which remained unregistered.
Subgenus Melanius Bonelli, 1810
91.5. Pterostichus atterimus (Herbst, 1784)
91.5.1. Pterostichus atterimus atterimus (Herbst, 1784)
References. Pterostichus atterimus: Apfelbeck, 1904: 258 (Velipoja); Schatzmayr, 1943a: 87.
Pterostichus atterimus atterimus: Bousquet, 2003f: 499.
Material studied. Gjormi / Albania leg. Bischoff 1936, 1 s. (MNHUB).
91.6. Pterostichus elongatus (Duftschmid, 1812)
References. Pterostichus elongatus: Schatzmayr, 1930: 210 (Shkodr); Hieke & Wrase, 1988: 76;
Bousquet, 2003f: 499.
152
153
155
Pterostichus (Pterostichus) latifianus latifianus Apfelbeck, 1906, new assignment from Pterostichus
(Aphaon) latifianus latifianus Apfelbeck, 1906
Pterostichus (Pterostichus) latifianus strupii J. Mller, 1935, new assignment from Pterostichus (Aphaon)
latifianus strupii J. Mller, 1935
Pterostichus (Pterostichus) lumensis lumensis Apfelbeck, 1905, new assignment from Pterostichus
(Aphaon) lumensis lumensis Apfelbeck, 1905
Pterostichus (Pterostichus) lumensis litae Csiki, 1940, new assignment from Pterostichus (Aphaon)
lumensis litae Csiki, 1940
Pterostichus (Pterostichus) lumensis ljubetensis Apfelbeck, 1908, new assignment from Pterostichus
(Aphaon) lumensis ljubetensis Apfelbeck, 1908
Pterostichus (Pterostichus) malissorum Apfelbeck, 1905, new assignment from Pterostichus (Aphaon)
malissorum Apfelbeck, 1905
Pterostichus (Pterostichus) meisteri Reitter, 1885, new assignment from Pterostichus (Aphaon) meisteri
Reitter, 1885
Pterostichus (Pterostichus) merklii (J. Frivaldszky, 1879), new assignment from Pterostichus (Aphaon)
merklii (J. Frivaldszky, 1879)
Pterostichus (Pterostichus) pentheri Apfelbeck, 1918, new assignment from Pterostichus (Aphaon)
pentheri Apfelbeck, 1918
Pterostichus (Pterostichus) reiseri Ganglbauer, 1889, new assignment from Pterostichus (Aphaon)
reiseri Ganglbauer, 1889
Pterostichus (Pterostichus) rhilensis rhilensis Rottenberg, 1874, new assignment from Pterostichus
(Aphaon) rhilensis rhilensis Rottenberg, 1874
Pterostichus (Pterostichus) rhilensis kourili Maan, 1933, new assignment from Pterostichus (Aphaon)
rhilensis kourili Maan, 1933
Pterostichus (Pterostichus) rhilensis vitosensis Maan, 1933, new assignment from Pterostichus (Aphaon)
rhilensis vitosensis Maan, 1933
Pterostichus (Pterostichus) walteri Reitter, 1883, new assignment from Pterostichus (Aphaon) walteri
Reitter, 1883
91.18. Pterostichus bischoffianus Jedlika, 1936
References. Pterostichus bischoffianus: Jedlika, 1936: 167 (Gjalic Mt., loc. typ.); Schatzmayr,
1943a: 109 (Gjalic Mt.); Bousquet, 2003f: 515.
Material studied. Gjallica / Albania leg. Bischoff 1936 / Typus / bischoffianus sp.n.
type det. Ing. Jedlika, 1 m. syntype (MNHUB); Gjallica / Albania leg. Bischoff 1936, 2
m. (MNHUB).
91.19. Pterostichus brucki Schaum, 1859
References. Pterostichus brucki: Apfelbeck, 1918c: 166 (North Albanian Alps, Vermosh, 1100 m);
Jedlika, 1936: 167 (Koritnik Mt.; Gjalic Mt.); Csiki, 1940: 216 (Gjalic Mt.); Schatzmayr, 1943a:
107 (Koritnik Mt.; Gjalic Mt.); Hieke, 1981: 80; Hieke & Wrase, 1988: 78; Bousquet, 2003f: 511.
Material studied. Vermoa 1100 m / Alban. montenegr. Grenze Penther14, 1 s. (NMW);
Albania, Mts. Gyalica Lums / Akad. Balk. Exp. Csiki 1918, 4 s. (HNHM); Alban. Exp.18
Gjalica Ljums 17-26.VI., 1 s. (NMW); Gjalica / Albania leg. Bischoff 1935, 1 s. (NMW);
Koritnik / Albania leg. Bischoff 1935, 8 s. (NMW; MNHUB); Gjallica / Albania leg.
156
157
(MNHUB); Korab / Albania leg. Bischoff 1935, more than 120 s. (MNHUB); Gjallica
/ Albania leg. Bischoff 1936, 14 s. (MNHUB); Korab / Albania leg. Bischoff 1936,
more than 120 s. (MNHUB).
Notes. Apfelbeck published two descriptions of this taxon (Apfelbeck, 1905; Apfelbeck,
1907b), and the older one is correctly treated as valid. Two specimens (1 m., 1 f.) labelled
Latif Alban. sept. and identified as Pterostichus djumanus Apfelbeck were found in DEI.
They are put provisionally here, since both specimens resemble P. lumensis. Pterostichus djumanus
Apfelbeck is nomen nudum.
91.21.2. Pterostichus lumensis litae Csiki, 1940
References. Pterostichus lumensis litae: Csiki, 1940: 216 (Gjalic Mt., loc. typ.; Korab Mt., loc.
typ.); Schatzmayr, 1943a: 108 (Gjalic Mt.; Korab Mt.); Bousquet, 2003f: 515.
Material studied. Albania, Mts. Gyalica Lums / Akad. Balk. Exp. Csiki 1918, 1 m. syntype
(HNHM); Albania Korab / Akad. Balk. Exp. Csiki 1917, 4 syntypes (HNHM); Albania
Mts. Korab / Akad. Balk. Exp. Csiki 1918, 21 syntypes (HNHM).
? Pterostichus lumensis ljubetensis Apfelbeck, 1908 [= chalybaeus Apfelbeck, 1908]
References. Pterostichus lumensis ljubetensis: Schatzmayr, 1930: 255; Bousquet, 2003f: 515.
Notes. Apfelbeck published three descriptions of this taxon (Apfelbeck, 1908; Apfelbeck,
1917; Apfelbeck, 1930), and the oldest one is correctly treated as valid. All data in the literature
and collections about ssp. ljubetensis come from the subalpine and alpine regions of the Shar
Planina Mts. that do not extend into Albania. Here, P. lumensis ljubetensis is treated as doubtful
for the fauna of Albania.
91.22. Pterostichus malissorum Apfelbeck, 1905 [= nigripes Jedlika, 1936]
References. Pterostichus malissorum: Apfelbeck, 1905: 241-242 (ukall Mt., loc. typ.); Apfelbeck,
1907a: 497 (Maranajt Mt.: top region, 1500-1576 m); Apfelbeck, 1907b: 637 (ukall Mt., loc.
typ.); Schatzmayr, 1930: 255; Jedlika, 1936: 169 (Malissora; ukall Mt.; Cafa Valbones; Cafa
Pejs; Maja Pejs); Schatzmayr, 1943a: 108 (ukall Mt.; Malissora; Cafa Valbones; Cafa Pejs;
Maja Pejs); Bousquet, 2003f: 515. Pterostichus latifianus ab. nigripes: Jedlika, 1936: 168 (North
Albanian Alps: Quf e Valbone, loc. typ.); Jedlika, 1939: 53. Pterostichus latifianus strupii (=
nigripes): Schatzmayr, 1943a: 109 (Quf e Valbone).
Material studied. Latif Alban. sept., 4 s. (DEI; HNHM); Latif Cukali / Coll.
Apfelbeck, 1 m. syntype (HNHM); Latif Cukali, 2 syntypes (NMW); Maranai Apfelb.,
1 s. (HNHM); Maranai Apf., 1 s. (HNHM); Apfelb. Albanien, 1 m. syntype (MNHUB);
Apfelb. Albanien / Alban. spt. Maranaj, 1 f. probably syntype (MNHUB); Apfelb. Alban
spt / Maranaj Geb., 2 s. (HNHM); Apfelb. Alban. spt. IV.V.05 / Maranaj, 6 s. (NMW);
Kunor Vukai Zukali Albanien Lona Giugno 1914, 6 s. (NMW; MNHUB); Cukale Albania
Bischoff , 12 s. (MNHUB); Cukale-Geb. / Albania leg. Bischoff 1931, 1 s. (MNHUB);
afa Stams / leg. Strupi Albania 1934 / Cotype / latifianus a. nigripes n. ab. det. Ing.
Jedlika, 1 f. designated as syntype of Pterostichus latifianus ab. nigripes Jedlika, 1936
(MNHUB); Cukale / Albania leg. Bischoff 1935, 7 s. (MNHUB); afa Valbons /
Albania leg. Bischoff 1935, 1 s. (MNHUB); Maja Pejs / Albania leg. Bischoff 1935, 3
s. (MNHUB); M. Mulecifit / Albania leg. Bischoff 1936, 14 s. (MNHUB); Maranaj /
Albania leg. Bischoff 1937, 23 s. (MNHUB); Boge ober D. Quelle / Albania leg. Mueller
158
159
Material studied. Mal i Dajtit / leg. Bischoff Albania 1934, 4 s. (MNHUB); Bojana
Shkoder / Albania leg. Bischoff 1937, 1 s. (MNHUB).
93.2. Stomis rostratus (Duftschmid, 1812)
93.2.1. Stomis rostratus rostratus (Duftschmid, 1812)
References. Stomis rostratus: Apfelbeck, 1918c: 166 (North Albanian Alps, Grebeni near
Vermosh, 1100-1800 m); Magistretti, 1965: 362; Bousquet, 2003f: 518.
Material studied. Grebeni 16.VI. / Alban. montenegr. Grenze, Penther14, 3 s. (NMW);
Alban. Vermosa Penther, 2 s. (NMW); Vermosa 1200 m 3.VI.-5.VII. / Alban. montenegr.
Grenze, Penther14, 4 s. (NMW); Ricavac 1300 m 25.VI.-2.VII. / Alban. montenegr. Grenze,
Penther14, 4 s. (NMW); Shklzn / Albania leg. Bischoff 1936, 2 s. (MNHUB).
161
Jedl.: Maan, 1939b: 295-299, 302-303 (Llogora). Tapinopterus extensus extensoides Jedlika:
Bousquet, 2003f: 519.
Material studied. Logara Juni / Albania leg. Bischoff 1932, 1 m. syntype (NMW); Maja
e Tartarit / Albania leg. Bischoff 1932, 1 s. (MNHUB); Reza e Kanalit Logara, Alban. m.
/ Albania leg. Bischoff 1932, 1 f. syntype (NMW); Logara juni / Albania leg. Bischoff
1932, 3 s. (MNHUB); Logara / leg. Bischoff Albania 1934, 7 s. (MNHUB); Llogora,
Alb. Mer. / Albania leg. Bischoff 1935, 1 f. syntype of extensoides Jedlika, 1936 (MNHUB);
Llogora / leg. Bischoff Albania 1936, 17 s. (MNHUB); Shen Ili / leg. Bischoff Albania
1936, 1 s. (MNHUB); Borshe / Albania leg. Bischoff 1940, 1 s. (MNHUB); Shpella Buz
i Lugit Mal i Piqs / Borshe / Albania leg. Bischoff 1940, 2 s. (MNHUB); Albania,
Periferi Tepelen, a gorge 1.5 km W of Progonat, / 785 m, 9.X.2004, Z. Fehr, J. Kontschn
& D. Murnyi (No. 133), 2 f. (HNHM).
Notes. In 1936 Jedlika (1936) and Straneo (1936) described T. extensoides having at disposal
identical specimens collected by Bischoff from the region of Llogora. Later, Mller (1937:
132, note 1) stated that the Straneos description has priority over that of Jedlika. This opinion
is correct in spite of Maans statement that both are authors of this taxon (Maan, 1939b:
302). It is Maan who first regarded extensoises as subspecies of T. extensus. However, in my
view T. extensoides Straneo is clearly distinct from all the other subspecies of T. extensus by an
evident apomorphic character 2 (instead 1) posterior supraorbital setiferous pores (see also
Straneo, 1936). The check of all specimens at hand demonstrated that this character is constant
(as only a single female from HNHM has a single left and two right posterior supraorbital
pores) so that it may define a separate species: Tapinopterus (Tapinopterus) extensoides (Straneo,
1936), stat. rest. from Tapinopterus (Tapinopterus) extensus extensoides (Straneo, 1936).
Species group protensus
94.2. Tapinopterus protensus (Schaum, 1857)
Notes. Bousquet (2003f: 520) noted that T. protensus aetolicus Ganglbauer, 1889 inhabits
Montenegro, and not Albania. In spite of that Schatzmayr (1943: 122) cited it for the studied
country (see below). Here, the species records and reference data from Albania are referred to
two defined and two undefined subspecies. The occurrence of T. protensus in Montenegro
needs serious confirmations.
94.2.1. Tapinopterus protensus peristericus Apfelbeck, 1901
References. Tapinopterus aetolicus peristericus: Mller, 1935: 153 (Shndelli Mt. near Tepelen).
Material studied. Maja e Papingut / Albania 1936 leg. Ramadan coll. Bischoff , 2 s.
(MNHUB); Maja e Policanit / Albania 1936 leg. Ramadan coll. Bischoff , 6 s. (MNHUB);
Shen Ili / leg. Bischoff Albania 1936, 3 s. (MNHUB); Maja e Policanit / Albania leg.
Bischoff 1937, 31 s. (MNHUB).
Notes. This subspecies is confined to the area between the valleys of Drina River and Vijos
River and lives also in Northwestern Greece. Mller (1935: 153-154) noted the identity of
syntypes of T. peristericus Apfelbeck, 1901 with specimens from the Shndelli Mt. In MNHUB,
I found specimens from Shndelli Mt. labelled aetolicus sbsp. peristericus det. J. Mller, and
others - aetolicus subsp. nova m. det. J. Mller.
162
163
165
167
Albania leg. Bischoff 1935, 2 s. (MNHUB); Gjallica / Albania leg. Bischoff 1935, 23 s.
(MNHUB); Teja / Albania leg. Bischoff 1935, 1 s. (MNHUB); Shkalla Harushe / Korab
/ Albania leg. Bischoff 1936, 1 s. (MNHUB).
Notes. First detailed records for Albania.
Subgenus Neocalathus Ball & Ngre, 1972
96.8. Calathus albanicus Apfelbeck, 1906
References. Calathus albanicus: Apfelbeck, 1906: 1666 (Merdita: Zep Mt., loc. typ.); Apfelbeck,
1907a: 500, 503; Schatzmayr, 1937: 2, 8; 36 (Zep Mt.); Csiki, 1940: 218 (Gjalic Mt.; Korab
Mt.); Hieke, 1981: 82; Hovorka & Sciaky, 2003b: 526.
Material studied. More than 160 s. with three kinds of labeling: Albania Galica Lums /
Akad. Balk. Exp. Csiki 1917 (HNHM); Albania, Mts. Gyalica Lums / Akad. Balk. Exp.
Csiki 1918 (HNHM); Albania Mts. Korab / Akad. Balk. Exp. Csiki 1918 (HNHM); more
than 150 s. from the massifs of Shar Planina Mt., Merdita, Munella Mt., Korab Mt., Gjalica
Ljums and others (NMW); Bjeshka Matroxh / Albania leg. Bischoff 1935, 9 s. (MNHUB);
Derz / Albania leg. Bischoff 1935, 1 s. (MNHUB); Gjallica / Albania leg. Bischoff
1935, 22 s. (MNHUB); Korab / Albania leg. Bischoff 1935, 8 s. (MNHUB); Mal i
Dajtit / Albania leg. Bischoff 1935, 1 s. (MNHUB); Fusha Korabit / Albania leg. Bischoff
1936, 2 s. (MNHUB); Gjallica / Albania leg. Bischoff 1936, 12 s. (MNHUB); Berishe
/ Albania leg. Mueller 1939, 2 s. (MNHUB); Maja Shalles i Vogel-Ibale / Albania leg.
Mueller 1939, 1 s. (MNHUB); Kunora Drunit Iballe / Albania leg. Bischoff 1939, 1 s.
(MNHUB); Mal i Krujes / Albania leg. Bischoff 1939, 1 s. (MNHUB).
Notes. This species inhabits the mountain chain Shar Planina Mt. - Korabi Mt. and massifs in
the area of Mirdita, but not the North Albanian Alps, where C. albanicus most probably is
replaced by C. metallicus.
96.9. Calathus ambiguus (Paykull, 1790)
96.9.1. Calathus ambiguus ambiguus (Paykull, 1790)
Material studied. AL (Berat) Pa ca. 5 km s. Gllav-600 m 27.IV.1993 leg. Heinz u. S. Peks,
3 s. (MNHUB).
Notes. The mention of Hieke & Wrase (1988: 90) for Albania is apparently result of following
the record Prisren (Apfelbeck, 1904: 285). Currently this locality is situated in the Kosovo
Region of Serbia. Hovorka & Sciaky (2003b: 526) have most likely followed one of these
sources too. Since there is no previous reliable record from Albania, C. ambiguus is treated here
as new species for the fauna of the country.
96.10. Calathus cinctus Motschulsky, 1850 [= erythroderus Gemminger & Harold, 1868]
References. Calathus mollis f. erythroderus: Schatzmayr, 1943b: 111 (Lushnje). Calathus cinctus:
Aukema, 1990: 138.
Material studied. Mal i Daitit / Albania leg. Bischoff 1932, 3 s. (MNHUB); Mal i Krujes
/ Albania leg. Bischoff 1935, 7 s. (MNHUB); Okol Shkalla / Albania leg. Bischoff 1935,
7 s. (MNHUB); Mal i Krujes / Albania leg. Bischoff 1936, 3 s. (MNHUB); Mal i Daitit
COLEOPTERA: CARABIDAE: HARPALINAE
169
/ Albania 1936 leg. Ramadan coll. Bischoff , 4 s. (MNHUB); Kruja Castle, 16.XI.1997, 1 s.,
R. Ward leg. (CMNH).
Additional material included (studied A. Vigna Taglianti). Rapsh, 15.V.2002, 1 m., A.
Paparistos leg. (cAVT).
Notes. Hovorka & Sciaky (2003b: 527) have omitted C. cinctus for Albania. Here the species is
confirmed for the country.
96.11. Calathus erratus (C.R. Sahlberg, 1827)
96.11.1. Calathus erratus erratus (C.R. Sahlberg, 1827)
References. Calathus erratus: Hieke & Wrase, 1988: 90. Calathus erratus erratus: Hovorka &
Sciaky, 2003b: 527.
Material studied. Maja e Jezerco / Albania leg. Bischoff 1935, 4 s. (MNHUB); Shklzn
/ Albania leg. Bischoff 1935, 2 s. (MNHUB).
Additional material included (studied A. Vigna Taglianti). Tropoia, valley of Valbona River,
900 m, 5.VII.1993, 1 m., L. Latella leg. (cAVT).
Notes. First detailed records for Albania. In addition to the material studied above, the present
author has seen several further specimens in NMW, which remained unregistered.
96.12. Calathus melanocephalus (Linnaeus, 1758)
96.12.1. Calathus melanocephalus melanocephalus (Linnaeus, 1758)
References. Calathus melanocephalus: Csiki, 1940: 218 (Shklzen Mt.; Gjalic Mt.; Korab Mt.);
Hieke et al., 1986: 189 (Biz near Shngjergji; Lur e Vjetr near Lura Mt.); Aukema, 1990:
138. Calathus melanocephalus melanocephalus: Hovorka & Sciaky, 2003b: 527.
Material studied. Albania Mts. Korab / Akad. Balk. Exp. Csiki 1918, 5 s. (HNHM); 30 s.
from localities Shklzn, Bjeshka Matroxh, Tomor, Korab, Koritnik, Gjallica,
Mal i Shenjit, Boge, Cucale, Taja, Llogora, Cafa e Sherit Iballe, afa Markofes
(MNHUB); Tomor Mt., 21.IX.1961, 1 m., 1 f. (MNST).
Additional material included (studied A. Vigna Taglianti). Tropoia, valley of Valbona River,
900 m, 5.VII.1993, 1 f., L. Latella leg. (cAVT).
Notes. The mention of Hieke & Wrase (1988: 91) for Albania refers apparently to the record
Prisren (Apfelbeck, 1904: 286). Actually this locality is presently situated in the Kosovo
Region of Serbia. Besides the specimens above, I have seen several further specimens in
NMW, which remained unregistered.
96.13. Calathus metallicus Dejean, 1828
96.13.1. Calathus metallicus prokletijensis Maan, 1938
References. Calathus metallicus: Hieke & Wrase, 1988: 90. Calathus metallicus metallicus: Hovorka
& Sciaky, 2003b: 527.
Material studied. Prokletija 1300-2000 m / Alban. montenegr. Grenze, Penther14, 15 s.
(NMW); nearly 100 s. from localities Shklzn, afa Markofes, Maja e Jezerze
(MNHUB); North Albanian Alps, Bog, 1400-1800 m, 1.VI.1993, 1 m., 1 f., P. Beron & B.
Petrov leg. (NMNHS); North Albanian Alps, Bog, 1000-1100 m, 5-9.VI.1993, 1 m., P. Beron
& B. Petrov leg. (NMNHS).
170
Additional material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollats Mt.,
2250 m, 11.VII.1993, 2 m., 1 f., L. Latella leg. (cAVT).
Notes. First detailed records for Albania. The absence of C. metallicus in the massifs of Shar
Planina and Korab Mt. may be a result of its replacement by C. albanicus there.
96.14. Calathus mollis (Marsham, 1802)
96.14.1. Calathus mollis mollis (Marsham, 1802)
References.Calathus mollis: Apfelbeck, 1904: 285 (Vlor; Berat); Apfelbeck, 1909: 297 (Vlor;
Berat); Hieke & Wrase, 1988: 91.
Material studied. Albania Avlona v. Oertzen, 1 s. (MNHUB).
Notes. Hovorka & Sciaky (2003b: 528) have omitted C. mollis for Albania. Here the species is
confirmed for the country. C. mollis is exclusively a littoral species and most probably the record
from Berat (Apfelbeck, 1904; 1909) concerns C. cinctus (A. Vigna Taglianti, personal communication).
171
173
Material studied. Shklzn / Albania leg. Bischoff 1936, 1 s. (MNHUB); Albania Theth
/ 23.VII.1960 leg. Xh. Murraj, 1 s. (HNHM).
Notes. First detailed records for Albania.
175
176
177
Material studied. Tomor Albanien, 1 s. (MNHUB); Shen Ili / Albania leg. Bischoff
1932, 1 s. (MNHUB); Tomor-Buranj juli 1932 / Albania leg. Bischoff 1932, 1 s. (MNHUB).
Notes. First detailed records for Albania.
Subgenus Paracelia Bedel, 1899
100.21. Amara dalmatina Dejean, 1828
References. Amara dalmatina: Hieke (in press).
Material included (studied F. Hieke). SE of Medua, Albania, 1 s. (MNHUB); Shendeli,
Albania mer., 1 s. (MNHUB).
Notes. First detailed records for Albania.
100.22. Amara quenseli (Schnherr, 1806)
100.22.1. Amara quenseli quenseli (Schnherr, 1806)
References. Amara quenseli: Apfelbeck, 1907a: 497, 499 (Maranajt Mt.: top region, 1500-1576
m; Mal i Shenjt Mt.: top region, 1500 m); Csiki, 1940: 216 (Korab Mt.); Holdhaus & Lindroth,
1940: 146-147, 250 (Koritnik Mt.; Korab Mt.); Hieke & Wrase, 1988: 106. Amara quenseli quenseli:
Hieke, 2003: 564.
Material studied. Latif Koritnik / Coll. Apfelbeck, 2 s. (HNHM); Latif Koritnik, 8 s.
(NMW); Merdita Munela / Apfelb. Alban. spt. IV.V.05, 1 s. (NMW); Prokletija 13002000 m / Alban. montenegr. Grenze, Penther14, 2 s. (NMW); Prokletija 1300-2000 m /
Alban. montenegr. Grenze, Penther14, 1 s. (NMW); Albania Galica Lums / Akad. Balk.
Exp. Csiki 1917, 2 s. (HNHM); Albania, Mts. Gyalica Lums / Akad. Balk. Exp. Csiki
1918, 3 s. (HNHM); Korab / Albania leg. Bischoff 1935, 16 s. (MNHUB); Korab /
Albania leg. Bischoff 1936, 19 s. (MNHUB); Shklzn / Albania leg. Bischoff 1936, 23
s. (MNHUB).
Subgenus Parapercosia Tschitschrine, 1899
100.23. Amara taurica (Motschulsky, 1844) [= fallaciosa (J. Mller, 1946)]
References. Zabrus fallaciosus: Mller, 1946: 107-110 (South Albania: Shndelli Mt. near Tepelen,
loc. typ.); Turin, 1981: 148. Amara taurica: Hieke, 2003: 565.
Notes. Hieke (1995: 35-36) synonymized Zabrus fallaciosus Mller, 1946 with Amara taurica,
noting the type locality of the former as Patria falsa!, but later the same has cited A. taurica
for Albania (Hieke, 2003: 565). According to F. Hieke (personal communication) A. taurica is
known from different parts of Turkey and from two localities in Greece, so that the type
locality of Zabrus fallaciosus in Albania should not be considered as doubtful (Hieke, 2005).
Subgenus Percosia C. Zimmermann, 1832
100.24. Amara equestris (Duftschmid, 1812)
100.24.1. Amara equestris equestris (Duftschmid, 1812) [= tomoricensis J. Mller, 1923]
COLEOPTERA: CARABIDAE: HARPALINAE
179
References. Amara tomoricensis: Mller, 1923: 108-109 (Tomor Mt.: Kulmak, loc. typ.). Amara
equestris: Csiki, 1940: 216 (Korab Mt.); Hieke, 1978: 229, 257 (Tomor Mt.: Kulmak; Koritnik
Mt.; Korab Mt.); Hieke & Wrase, 1988: 108. Amara equestris equestris: Hieke, 2003: 565.
Material studied. Albania Mts. Korab / Akad. Balk. Exp. Csiki 1918, 1 s. (HNHM);
Shen Ili / Albania leg. Bischoff 1932, 1 s. (MNHUB); Korab / Albania leg. Bischoff
1935, 1 s. (MNHUB); Koritnik / Albania leg. Bischoff 1935, 1 s. (MNHUB); Albania
Zalle-Dardha / 24.VII.1958 leg. Xh. Murraj, 1 s. (HNHM).
Subgenus Xenocelia Hieke, 2001
100.25. Amara bischoffi Jedlika, 1946
References. Amara municipalis: Csiki, 1940: 215 (Gjalic Mt.; Korab Mt.); Jedlika, 1946: 7
Peshtrik Mt. (loc. typ.); Hieke & Wrase, 1988: 104. Amara bischoffi: Turin, 1981: 145; Hieke,
1983: 328-329, 334 (Peshtrik Mt.; Taja); Hieke, 2001: 33 (two localities). Amara municipalis
bischoffi: Hieke, 2003: 557.
Material studied. Taja / Albania leg. Bischoff 1935, 1 s. (MNHUB); Bashtrik / Albania
leg. Bischoff 1936, 1 syntype (MNHUB).
Notes. According to F. Hieke (personal communication) the records of Csiki (1940) are
problematic since they may concern A. bischoffi, as well as A. messae.
100.26. Amara cursitans C. Zimmermann, 1832 [= properans C. Zimmermann, 1832]
References. Amara properans: Csiki, 1940: 215 (Gjalic Mt.; Korab Mt.). Amara cursitans: Hieke
& Wrase, 1988: 104; Hieke, 2001: 50 (five localities); Hieke, 2003: 556.
Material studied. Prokletija 1300-2000 m / Alban. montenegr. Grenze, Penther14, 2 s.
(NMW); Sklzn / Albania leg. Bischoff 1935, 2 s. (MNHUB); Gjallica / Albania leg.
Bischoff 1936, 5 s. (MNHUB); Llogora / Albania leg. Bischoff 1936, 1 s. (MNHUB);
Maja e Dragomit / Albania leg. Bischoff 1939, 1 s. (MNHUB).
100.27. Amara fusca Dejean, 1828 [= complanata Dejean, 1828]
References. Amara complanata: Csiki, 1940: 215 (Gjalic Mt.). Amara fusca: Hieke & Wrase,
1988: 104; Hieke, 2001: 42 (Koritza-Ebene, 1 s. in MNHN); Hieke, 2003: 556.
100.28. Amara messae Baliani, 1924
References. Amara messae: Hieke & Wrase, 1988: 104; Hieke, 2001: 65; Hieke, 2003: 557.
Material studied. Giallica / Albania leg. Bischoff 1935, 1 s. (MNHUB); Maja e Jezerco
/ Albania leg. Bischoff 1935, 1 s. (MNHUB); Cotype / Bashtrik / Albania leg. Bischoff
1936, 1 s. (MNHUB); North Albanian Alps, Bog, 1400-1800 m, 1.VI.1993, 1 s., P. Beron, B.
Petrov leg. (NMNHS).
Notes. First detailed records for Albania.
180
181
from collections, and may bases of supposition. The subspecies most probably inhabits the
Albanian part of the Jablanitsa Mt., too.
101.2.2. Zabrus albanicus latifianus Ganglbauer, 1915
References. Zabrus albanicus: Apfelbeck, 1907a: 503 (Zep Mt.). Zabrus albanicus latifianus:
Ganglbauer, 1931: 39 (Zep Mt.); Maan, 1939d: 145-146 (Zep Mt.); Schatzmayr, 1943b: 108
(Zep Mt.); Freude, 1989: 83, 88, 97 (Zep Mt.); Serrano & Andjar, 2003: 570.
Material studied. Merdita Munela, 1 s. (NMW); Merdita Zebia, 2 s. (NMW).
Notes. In my opinion, only the populations from the region of Merdita have to be considered
as belonging to spp. latifianus.
! Zabrus albanicus ssp.
Zabrus albanicus: Schatzmayr, 1943b: 108 (Tomor Mt.: Abbas Ali and Toik).
Material studied. Tomorica Abbas Ali / Albania 1922 Ravasini, Lona, 4 s. (NMW;
MNHUB); Maja e Dhembelit / Albania 1936 leg. Ramadan coll. Bischoff , 8 s. (MNHUB);
Albania 1936 leg. Ramadan coll. Bischoff / Maja e Papingut, 21 s. (MNHUB).
Notes. The specimens from the massifs of Tomor Mt., Maja e Dhembelit Mt. and Maja e
Papingut Mt. (all in South Albania) may concern another subspecies, since the latter are far
removed in the south from the massifs of the Zep Mt. and Koritnik Mt.
# Zabrus apfelbecki Ganglbauer, 1915
References. Zabrus apfelbecki: Magistretti, 1965: 360.
Notes. Magistretti (1965) referred older records of Z. ignavus Csiki, 1907 to Z. apfelbecki,
but I failed to find any records in the literature. The latter species lives only in Bosnia
Herzegovina (Serrano & Andjar, 2003: 570), so that it is removed from the list of the
Albanian ground beetles.
101.3. Zabrus ganglbaueri Apfelbeck, 1906
101.3.1. Zabrus ganglbaueri ganglbaueri Apfelbeck, 1906
References. Zabrus ganglbaueri: Apfelbeck, 1906: 1667 (Merdita: Mal i Shenjt Mt. near Oroshi,
loc. typ.); Apfelbeck, 1907a: 499-500, 503 (Mal i Shenjt Mt.: top region, 1500 m); Mller, 1936:
108 (Mal i Shenjt Mt. near Oroshi); Schatzmayr, 1943b: 108 (Mal i Shenjt Mt. near Oroshi);
Hieke, 1981: 75. Zabrus ganglbaueri ganglbaueri: Freude, 1989: 82, 110 (Mal i Shenjt Mt. near
Oroshi; Mal i That Mt. near Gjirokastr); Serrano & Andjar, 2003: 570.
Material studied. Merdita M. Scheit, 1 s. (NMW); Merdita 18.19. n. 20/5 1934, 2 s. (NMW).
Notes. Freude (1989) stated that the populations from the Mal i Shenjt Mt. (Merdita District)
and the Mal i That Mt. (Gjirokastr District) belong to the nominotypical race. In fact both
populations inhabit opposite outskirts of the country, so the specimens from the latter massif
may belong either to the following subspecies or to a separate geographical race.
101.3.2. Zabrus ganglbaueri lonae J. Mller, 1923
References. Zabrus lonae: Mller, 1923: 107-108 (Tomor Mt.: Toik, loc. typ.); Mller, 1935:
154 (Tomor Mt.: Toik); Maan, 1940a: 30-31, 33, 35. Zabrus lonai: Schatzmayr, 1943b: 108
(Tomor Mt.: Toik). Zabrus ganglbaueri lonae: Freude, 1989: 83, 111 (Tomor Mt.: Toik); Serrano
& Andjar, 2003: 570.
COLEOPTERA: CARABIDAE: HARPALINAE
183
Material studied. Albania 1922 Ravasini, Lona / Tomor Monte Toik, 1 s. (DEI); Tomor:
Toik Alban. mer. / Winkler, Lona, Bischoff, V.1931, 1 s. (NMW); Tomor: Tojk / Albania
leg. Bischoff 1932, 9 s. (MNHUB).
Species group incrassatus
101.4. Zabrus bischoffi J. Mller, 1936
References. Zabrus bischoffi: Mller, 1936: 107-108 (Mal i Dajtit Mt. near Tiran, loc. typ.;
Llogora); Schatzmayr, 1943b: 106, 109 (Kruja; Mal i Dajtit Mt.; Llogora); Turin, 1981: 148.
Zabrus incrassatus bischoffi: Freude, 1989: 86, 120 (Mal i Dajtit Mt. near Tiran; Llogara, south of
Vlor); Serrano & Andjar, 2003: 571.
Material studied. 19 s. from Mal i Dajtit (MNHUB); 5 s. from Mal i Krues (MNHUB);
3 s. from Mal i Shenjit (MNHUB); 37 s. from Llogora (MNHUB); Llogora Pass, 1025 m,
6.6.1995, 1 m., G. Blagoev leg. (NMNHS).
Notes. This species has a quite different apex of the median lobe and sides of pronotum
narrower anteriorly, which is in contrast with the respective characteristics in Z. incrassatus s.l.
In my opinion, the former is separate species not subspecies of the latter: Zabrus (Pelor) bischoffi
J. Mller, 1936, stat. rest. of Zabrus (Pelor) incrassatus bischoffi J. Mller, 1936.
101.5. Zabrus boldorii Schatzmayr, 1943
References. Zabrus boldori: Schatzmayr, 1943: 106, 108 (Dukati, south of Vlor, loc. typ.);
Serrano & Andjar, 2003: 570. Zabrus baldori sic!: Turin, 1981: 148. Zabrus incrassatus boldorii:
Freude, 1989: 86, 119 (Dukati near Vlor).
Material studied. Sdalbanien Kopstein / coll. Prof. F. Werner / Paaliman, 1 m.
(NMW); Sdalbanien Kopstein / Valona, 2 f. (NMW).
101.6. Zabrus incrassatus (Ahrens, 1814)
101.6.1. Zabrus incrassatus incrassatus (Ahrens, 1814)
References. Zabrus incrassatus: Apfelbeck, 1904: 310 (Berat; Vlor); Anonymous, 1907: 285;
Apfelbeck, 1907a: 495-496 (northeast of Shkodr; Maranajt Mt.: between Vorfai Postme, 300
m, and Vorfai Siperme, 700 m); Apfelbeck, 1909: 297 (Vlor; Berat); Mller, 1936: 108 (Shndelli
Mt.; Tepelen; Maja e Tartarit Mt.; Tiran; Gjirokastr; Berat; Vlor); Schatzmayr, 1943b: 106,
108 (Poggio Boschetto; Lushnje; Berat; Vlor; Shen Ili; Tepelen; Maja e Tartarit Mt.; Tiran;
Shkodr); Hieke & Wrase, 1988: 110.
Material studied. Sdalbanien Kopstein, 2 s. (NMW); Albanien Ochrida, 1 s. (NMW);
Hani Grabom Mai / Alban. montenegr. Grenze, Penther14, 2 s. (NMW); Rapsa 1018.V. / Alban. montenegr. Grenze, Penther14, 4 s. (NMW); Alban. Exp. 1918 Bicaj
14.15.VI., 3 s. (NMW); Ib, 2-11.VII. 1962, 1 m. (MNST); Shkodr, 15.V.1963, 3 m., 1 f.
(MNST); 2 km N Durrs, 24.V.1993, 1 m., P. Stoev leg. (NMNHS); Vlor, 1.V.1994, 2 m., 3 f.,
P. Stoev leg. (NMNHS); Saranda District, Himar, 3-4.V.1994, 1 f., P. Stoev leg. (NMNHS);
Saranda District, Borsh, 5.V.1994, 1 f., P. Stoev leg. (NMNHS); Saranda District, Ljukova, 34.VI.1995, 1 f., G. Blagoev leg. (NMNHS); Vlor District, Llogora Mt., 1027 m, 4.VI.1995, 1
f., G. Blagoev leg. (NMNHS).
184
Notes. Serrano & Andjar (2003: 571) have omitted the species for Albania, so it is here
confirmed for the country. Besides the material studied above, in MNHUB I have seen other
more than 100 s. from different localities (Shen Ili, Tirane, Murthi, Tepelene, Munela,
Burelle, Vlor, Mal i Gjer, Myzege, Bose, Maja e Dhembellit, Tarabosh, Mal
i Tartarit, etc.).
Species group spinipes
101.7. Zabrus spinipes (Fabricius, 1798)
101.7.1. Zabrus spinipes insignis J. Mller, 1932
References. Zabrus spinipes: Burmeister, 1939: 127; Hieke et al., 1986: 190 (Tomor Mt.: Abbas
Ali Monastery); Hieke & Wrase, 1988: 110; Freude, 1989: 147.
Material studied. Shen-Ili Juni / Albania leg. Bischoff 1932, 11 s. (MNHUB); Mal i
That / Albania leg. Bischoff 1937, 22 s. (MNHUB).
Notes. Serrano & Andjar (2003: 572) have omitted the species for Albania, so it is here
confirmed for the country.
Subgenus Zabrus s.str.
101.8. Zabrus tenebrioides (Goeze, 1777)
101.8.1. Zabrus tenebrioides tenebrioides (Goeze, 1777)
References. Zabrus tenebrioides: Schatzmayr, 1943b: 105-106 (Dukati; Lushnje). Zabrus tenebrioides
tenebrioides: Serrano & Andjar, 2003: 573.
Material studied. Myzeqe / Albania leg. Bischoff 1937, 1 s. (MNHUB); 11 km west of
Tiran, 20.VI.1994, 1 s., B. Guorguiev leg. (NMNHS); Radona, Shkiela, 21.VI.1994, 5 s., B.
Guorguiev leg. (NMNHS).
Additional material included (studied A. Vigna Taglianti). Durrs, Currila, 8.V.2002, 1 f., A.
Paparistos leg. (cAVT); Qesarat near Memaliaj (Tepelen District), valley of Vijos River, 300360 m, 18.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT).
185
186
Results
When omitting doubtful species and taxa still not well defined, the Albanian Carabidae
listed in this catalogue represent 543 species. They are distributed among 11 subfamilies
(Nebriinae 25, Cicindelinae 10, Carabinae 20, Elaphrinae 1, Omophroninae
1, Brachininae 17, Siagoninae 1, Scaritinae 28, Apotominae 2, Trechinae 115,
and Harpalinae 323), 31 tribes and 103 genera (Table 1).
One tribe, 10 genera, 11 subgenera and 55 species are new for the fauna of Albania,
and 1 subgenus and 3 species are new for the fauna of the Balkan Peninsula (Appendix
A.1-2). From these, 1 tribe, 1 subtribe, 7 genera, 20 subgenera and 72 species are
reported firstly with detailed record/s for the country (Appendix A.3). Altogether
119 species, omitted in the recent Palaearctic Catalogue (Lbl & Smetana, 2003), are
here confirmed for Albania (Appendix A.4).
Twelve taxa from the species group remain still not well defined, as likely most of
them represent new for the science species or subspecies (Appendix A.5). Two
subgenera and 22 species and subspecies are doubtful for the country (Appendix
A.6); generally the published records about them concern adjacent regions in
Montenegro, the Kosovo Region of Serbia, Macedonia or Northwestern Greece. Most
of the species and subspecies from the last category may inhabit Albania, but until
now we do not have reliable data to confirm this. Two subgenera and 29 species and
subspecies are deleted from the list (Appendix A.7). The systematic list includes also
some endemics described from massifs shared between Albania and neighbour
countries (but not recorded yet from the former), as well as several more widespread
species found from localities very close to Albania.
A moderate taxonomic contribution is also achieved (Appendix B). Principally, it is
based on study of types, topotypes or more extensive series of previously problematic
species and subspecies from Albania and neighbouring countries. In result of that 38
various taxonomic innovations (new synonyms, new status or revalidations) are
introduced. Further, nine taxa from the species group (5 valid species and 4 synonyms),
all described by V. Apfelbeck, have been found to have assigned an incorrect year of
description. Accordingly, the relevant earlier years of their descriptions are fixed. Finally,
eight nomina nuda were also traced out, as one of them may be a result of mislabeling,
and one is supplied with year of description in the literature.
RESULTS
187
Conclusions
Undoubtedly the Balkan Peninsula contains the richest ground beetle diversity
compared to other diverse European regions like the Iberian Peninsula, Italy or the
Caucasus Major (Turin, 1981; Kryzhanovskij et al., 1995; Serrano, 2003; Vigna Taglianti,
1993; 2004; 2005a; 2005b). Currently an impressive total of at least 1487 species of
Carabidae, following the classification used in Lbl & Smetana, 2003, is reported
from this territory (unpublished data). This figure is provisional while the systematics,
diversity and distribution of the Carabidae in the Balkan Peninsula are better known.
Until now, the carabid fauna of Bulgaria is probably the best investigated one in the
region (Hieke & Wrase, 1988; Guorguiev & Guorguiev, 1995). The species
composition of no other Balkan regions has been examined in details; only that of
ex-Yugoslavia was subject of uncritical registration (Drovenik & Peks, 1994; Drovenik
& Peks, 1999). Therefore, the Carabidae of Slovenia, Croatia, Bosnia Herzegovina,
Montenegro, Serbia, Macedonia, Romania, Greece and European Turkey need scrutiny
and writing of up-to-date checklists.
Bearing in mind the surface of Albania, the number of the carabid species and
subspecies there seems relatively high (543), if compared it with that of other Balkan
or Mediterranean regions. In Bulgaria there are known more than 720 species
(Guorguiev & Guorguiev, 1995), from Macedonia there are reported about 420
species (Guorguiev, 1998; Hristovski, personal communication), and for ex-Yugoslavia
more than 1500 species and subspecies have been cited (Drovenik & Peks, 1999).
Both the Iberian Peninsula and Italy, having territories from 10 to 20 times bigger
than Albania, have a number of species only 2.13-2.42 times higher than is that in the
last country (Serrano, 2003; Vigna Taglianti, 2005a; 2005b).
188
Tribe
Nebriinae
Nebriini
Cicindelinae
Carabinae
Subtribe
Apotominae
Trechinae
Nr.
species
Eurynebria
Leistus
1
7
Nebria
Notiophilus
Calomera
Cicindela
Cylindera
8
2
4
3
Myriochila
Calosoma
1
4
Carabus
15
Cychrus
Elaphrus
Omophron
Aptinus
Brachinus
1
1
1
3
14
Clivinina
Reicheiina
Dyschiriini
Siagona
Clivina
Reicheadella
Dyschirius
1
3
1
21
Scaritini
Scaritina
Scarites
Apotomini
Bembidiini
Bembidiina
Apotomus
Asaphidion
Bembidion
2
4
63
Notiophilini
Cicindelini
Carabini
Cychrini
Elaphrinae
Elaphrini
Omophroninae Omophronini
Brachininae
Brachinini
Siagoninae
Scaritinae
Genus
Siagonini
Clivinini
Subgenus
Nr.
species
Leistus
Pogonophorus
Alpaeus
Boreonebria
Eunebria
Nebria
2
5
5
1
1
2
Cicindela
Cylindera
Eugrapha
Myriochila
Calosoma
Microcallisthenes
Archicarabus
Carabus
Chaetocarabus
Megodontus
Oreocarabus
Pachystus
Procerus
Procrustes
Tachypus
Tomocarabus
4
1
2
1
2
2
1
1
1
3
3
2
1
1
1
1
Neoelaphrus
Omophron
Aptinus
Brachinus
Brachynidius
Brachynolomus
Cnecostolus
insertae sedis
1
1
3
5
4
1
3
1
Reicheadella
Chiridysus
Dyschiriodes
Dyschirius
Eudyschirius
Paradyschirius
Parallelomorphus
Scallophorites
1
1
14
1
3
2
2
1
Bembidion
189
Subfamily
Tribe
Subtribe
Tachyina
Patrobini
Pogonini
Trechini
Harpalinae
Chlaeniini
Cyclosomini
Dryptini
190
Perileptina
Trechina
Trechodina
Callistina
Chlaeniina
Masoreina
Genus
Nr.
species
Ocys
Sinechostictus
1
3
Lymnastis
Paratachys
Polyderis
Sphaerotachys
Tachys
Tachyta
Tachyura
Deltomerus
1
3
1
1
1
1
2
2
Pogonistes
Pogonus
Perileptus
Albanotrechus
Duvalius
2
2
1
1
10
Neotrechus
Trechus
Thalassophilus
Callistus
Chlaenius
3
12
1
1
13
Masoreus
Drypta
1
1
Subgenus
Nr.
species
Bembidionetolitzkya
Chlorodium
Diplocampa
Emphanes
Euperyphus
Metallina
Microserrullula
Nepha
Notaphemphanes
Notaphus
Ocydromus
Ocyturanes
Odontium
Omoperyphus
Peryphanes
Peryphus
Philochthus
Phyla
Princidium
Talanes
Terminophanes
Testedium
Trepanes
9
1
2
5
3
2
1
3
1
1
3
6
2
1
4
4
6
1
1
1
1
1
3
Pseudolimnaeum
Sinechostictus
1
2
Tachyta
Tachyura
Deltomerus
Paradeltomerus
1
2
1
1
Pogonus
Perileptus
2
1
Duvalius
Euduvalius
9
1
Trechus
12
Chlaeniellus
Chlaenites
Chlaenius
Dinodes
Epomis
Trichochlaenius
Masoreus
Drypta
6
1
1
2
2
1
1
1
Subfamily
Tribe
Subtribe
Harpalini
Amblystomina Amblystomus
Anisodactylina Anisodactylus
Harpalus
32
Ophonus
20
Parophonus
Trichotichnus
Acupalpus
Anthracus
Bradycellus
Dicheirotrichus
Loxoncus
Stenolophus
1
10
2
4
1
1
7
Cymindidina
Cymindis
Dromiina
Dromius
Microlestes
Paradromius
Philorhizus
Lebia
1
7
1
2
5
Apristus
Lionychus
Syntomus
Badister
2
2
4
4
Licinus
Odacantha
Oodes
Omphreus
Panagaeus
Agonum
1
2
3
1
15
Stenolophina
Lebiina
Lionychina
Odacanthini
Oodini
Omphreini
Panagaeini
Platynini
3
4
1
1
3
2
3
1
1
4
Harpalina
Licinini
Nr.
species
Diachromus
Gynandromorphus
Carterus
Ditomus
Dixus
Graniger
Odontocarus
Acinopus
Ditomina
Lebiini
Genus
Licinina
Subgenus
Nr.
species
Anisodactylus
Pseudodichirius
3
1
Carterus
Acinopus
Oedematicus
Osimus
Cryptophonus
Harpalus
Pseudophonus
Brachyophonus
Hesperophonus
Incisophonus
Metophonus
Ophonus
Ophonomimus
Parophonus
Trichotichnus
Acupalpus
2
1
1
1
29
2
1
4
1
10
4
1
5
1
10
Bradycellus
Dicheirotrichus
4
1
Egadroma
Stenolophus
Cymindis
Menas
Tarulus
Dromius
1
6
7
1
1
1
Manodromius
Philorhizus
Lamprias
Lebia
1
2
1
4
Lionychus
Badister
Baudia
Licinus
Neorescius
Odacantha
Oodes
Omphreus
Panadaeus
Agonum
Europhilus
2
2
3
1
1
2
3
1
13
2
191
Subfamily
Tribe
Subtribe
Zabrini
Zuphiini
11 subfamilies 31 tribes
192
Nr.
species
Anchomenus
Olisthopus
Paranchus
Platynus
1
2
1
2
Abax
Molops
15
Myas
Pedius
Poecilus
1
2
8
Pterostichus
24
Speluncarius
Atranopsina
Calathina
Stomis
Tapinopterus
Platyderus
Calathus
2
6
4
14
Sphodrina
Synuchidius
Laemostenus
1
7
Synuchina
Amarina
Synuchus
Amara
1
32
Zabrina
Zabrus
Pterostichini
Sphodrini
Genus
Polystichus
Zuphium
103 genera
Subgenus
Nr.
species
Anchomenus
Batenus
Platynus
Abacopercus
Abax
Molops
Stenochoromus
Myas
1
1
1
1
14
1
1
Angoleus
Poecilus
Argutor
Bothriopterus
Cheporus
Feronidius
Melanius
Morphnosoma
Parahaptoderus
Phonias
Platypterinus
Platysma
Pseudomaseus
Pterostichus
Hypogium
Speluncarius
Stomis
Tapinopterus
Platyderus
Bedelinus
Calathus
Neocalathus
1
7
1
1
1
1
2
1
1
3
1
1
4
7
1
1
2
6
4
1
6
7
Antisphodrus
Laemostenus
Pristonychus
Synuchus
Amara
Amarocelia
Bradytus
Celia
Curtonotus
Paracelia
Parapercosia
Percosia
Xenocelia
Zezea
Pelor
Zabrus
2
3
2
1
13
1
2
3
1
2
1
1
4
4
7
1
1
1
Zuphium
543 species
Appendix
A. Faunistic contributions
A.1. New taxa for the fauna of the Balkan Peninsula
Subgenera
1. Terminophanes Mller-Motzfeld, 1998
Species
1. Bembidion (Terminophanes) terminale Heer, 1841
2. Cymindis (Cymindis) angularis Gyllenhal, 1810
3. Lionychus (Lionychus) maritimus Fairmaire, 1862
A.2. Taxa new for the fauna of Albania
Tribes
1. ODACANTHINI Laporte, 1834
Genera
1. Eurynebria Ganglbauer, 1891
2. Myriochila Motschulsky, 1858
3. Lymnastis Motschulsky, 1862
4. Polyderis Motschulsky, 1862
5. Pogonistes Chaudoir, 1871
6. Odontocarus Solier, 1835
7. Dromius Bonelli, 1810
8. Lionychus Wissmann, 1846
9. Odacantha Paykull, 1798
10. Paranchus Lindroth, 1974
Subgenera
1. Myriochila Motschulsky, 1858 s.str.
2. Dyschirius Bonelli, 1810 s.str.
3. Talanes Motschulsky, 1864
4. Terminophanes Mller-Motzfld, 1998
5. Pseudolimnaeum Kraatz, 1888
6. Menas Motschulsky, 1864
7. Dromius Bonelli, 1810 s.str.
8. Lamprias Bonelli, 1810
9. Lionychus Wissmann, 1846 s.str.
10. Odacanta Paykull, 1798 s.str.
11. Parahaptoderus Jeanne, 1969
Species (subspecies not considered)
1. Eurynebria complanata (Linnaeus, 1767)
2. Leistus (Pogonophorus) apfelbecki Ganglbauer, 1891
APPENDIX
193
195
197
199
APPENDIX
201
B. Taxonomic contributions
B.1. Nomenclatorial acts
Nebriini
Nebria (Alpaeus) macedonica Maan, 1938, stat. rest. for Nebria (Alpaeus) aetolica macedonica Maan, 1938
Nebria (Alpaeus) macedonica galicica Horvatovich, 1974a, new assignment of Nebria (Alpaeus)
aetolica galicica Horvatovich, 1974a
Nebria (Alpaeus) macedonica papingutica Horvatovich, 1974a, new assignment of Nebria (Alpaeus)
aetolica papingutica Horvatovich, 1974a
Nebria (Alpaeus) macedonica rambouseki Maan, 1938, new assignment of Nebria (Alpaeus) aetolica
rambouseki Maan, 1938
Carabini
Carabus (Oreocarabus) neumeyeri Schaum, 1856, stat. nov. for Carabus (Oreocarabus) hortensis neumeyeri
Schaum, 1856
Carabus (Oreocarabus) preslii Dejean, 1830, stat. rest. for Carabus (Oreocarabus) hortensis preslii Dejean, 1830
Cychrini
Cychrus semigranosus albanicus Hopp, 1929, syn. rest. of Cychrus semigranosus montenegrinus Apfelbeck,
1904
Cychrus semigranosus dinaricus Mandl, 1989, syn. nov. of Cychrus semigranosus montenegrinus Apfelbeck,
1904
Licinini
Licinus (Licinus) submarginatus J.R. Sahlberg, 1903, syn. nov. of Licinus (Licinus) graecus Apfelbeck, 1901
Platynini
Platynus (Batenus) scrobiculatus serbicus Csiki, 1904, stat. nov. for Platynus (Batenus) serbicus Csiki, 1904
Platynus (Batenus) scrobiculatus turcicus Apfelbeck, 1904, stat. rest. for Platynus (Batenus) turcicus
Apfelbeck, 1904
Pterostichini
Molops (Molops) maderi J. Mller, 1933, syn. nov. of Molops (Molops) malshentianus Apfelbeck, 1917
Molops (Molops) apfelbecki reiseri Apfelbeck, 1904, new assignment of Molops (Molops) reiseri
reiseri Apfelbeck, 1904
Molops (Molops) apfelbecki corallipes J. Mller, 1936, new assignment of Molops (Molops) reiseri
corallipes J. Mller, 1936
Molops (Molops) apfelbecki pseudoalpestris Mlyn, 1977, new assignment of Molops (Molops) reiseri
pseudoalpestris Mlyn, 1977
Molops (Molops) apfelbecki victoris J. Mller, 1918, new assignment of Molops (Molops) reiseri
victoris J. Mller, 1918
Molops (Molops) holdhausi J. Mller, 1917, stat. rest. for Molops (Molops) reiseri holdhausi J. Mller, 1917
Aphaon Heyden, 1906, syn. nov. of Pterostichus s. str. Bonelli, 1810
Pterostichus (Pterostichus) bischoffianus Jedlika, 1936, new assignment of Pterostichus (Aphaon)
bischoffianus Jedlika, 1936
202
Pterostichus (Pterostichus) brucki Schaum, 1859, new assignment of Pterostichus (Aphaon) brucki
Schaum, 1859
Pterostichus (Pterostichus) latifianus latifianus Apfelbeck, 1906, new assignment of Pterostichus
(Aphaon) latifianus latifianus Apfelbeck, 1906
Pterostichus (Pterostichus) latifianus strupii J. Mller, 1935, new assignment of Pterostichus (Aphaon)
latifianus strupii J. Mller, 1935
Pterostichus (Pterostichus) lumensis lumensis Apfelbeck, 1905, new assignment of Pterostichus
(Aphaon) lumensis lumensis Apfelbeck, 1905
Pterostichus (Pterostichus) lumensis litae Csiki, 1940, new assignment of Pterostichus (Aphaon) lumensis
litae Csiki, 1940
Pterostichus (Pterostichus) lumensis ljubetensis Apfelbeck, 1908, new assignment of Pterostichus
(Aphaon) lumensis ljubetensis Apfelbeck, 1908
Pterostichus (Pterostichus) malissorum Apfelbeck, 1905, new assignment of Pterostichus (Aphaon)
malissorum Apfelbeck, 1905
Pterostichus (Pterostichus) latifianus ab. nigripes Jedlika, 1936, syn. rest. for Pterostichus (Pterostichus)
malissorum Apfelbeck, 1905
Pterostichus (Pterostichus) meisteri Reitter, 1885, new assignment of Pterostichus (Aphaon) meisteri
Reitter, 1885
Pterostichus (Pterostichus) merklii (J. Frivaldszky, 1879), new assignment of Pterostichus (Aphaon)
merklii (J. Frivaldszky, 1879)
Pterostichus (Pterostichus) pentheri Apfelbeck, 1918, new assignment of Pterostichus (Aphaon) pentheri
Apfelbeck, 1918
Pterostichus (Pterostichus) reiseri Ganglbauer, 1889, new assignment of Pterostichus (Aphaon) reiseri
Ganglbauer, 1889
Pterostichus (Pterostichus) rhilensis rhilensis Rottenberg, 1874, new assignment of Pterostichus
(Aphaon) rhilensis rhilensis Rottenberg, 1874
Pterostichus (Pterostichus) rhilensis kourili Maan, 1933, new assignment of Pterostichus (Aphaon)
rhilensis kourili Maan, 1933
Pterostichus (Pterostichus) rhilensis vitosensis Maan, 1933, new assignment of Pterostichus (Aphaon)
rhilensis vitosensis Maan, 1933
Pterostichus (Pterostichus) walteri Reitter, 1883, new assignment of Pterostichus (Aphaon) walteri
Reitter, 1883
Tapinopterus (Tapinopterus) extensoides (Straneo, 1936), stat. rest. for Tapinopterus (Tapinopterus)
extensus extensoides (Straneo, 1936).
Sphodrini
Calathus (Calathus) ravasinii macedonicus Maan, 1935, stat. nov. for Calathus (Calathus) macedonicus
Maan, 1935
Zabrini
Zabrus (Pelor) bischoffi J. Mller, 1936, stat. rest. for Zabrus (Pelor) incrassatus bischoffi J. Mller, 1936.
B.2. Year of description
The following taxa (valid or synonyms) have long time been accredited to have a correct year
of description. However, it was found out that this is incorrect. Below, the relevant earlier
APPENDIX
203
years of publication dates of the original descriptions are given. All the descriptions considered
fulfil the conditions of the ICZN (1999).
Carabus croaticus ljubetensis Apfelbeck, 1917 (synonym of Carabus croaticus durmitorensis Apfelbeck,
1904) instead Carabus croaticus ljubetensis Apfelbeck, 1918a
Carabus violaceus merditanus Apfelbeck, 1917 (synonym of Carabus violaceus dryas Gistl, 1857)
instead Carabus violaceus merditanus Apfelbeck, 1918a
Carabus violaceus shardagensis Apfelbeck, 1917 (synonym of Carabus violaceus dryas Gistl, 1857)
instead Carabus violaceus shardagensis Apfelbeck, 1918a
Aptinus merditanus Apfelbeck, 1917 (valid species) instead Aptinus merditanus Apfelbeck, 1918b
Aptinus merditanus var. granulipennis Apfelbeck, 1917 (synonym of Aptinus merditanus Apfelbeck,
1917) instead Aptinus merditanus var. granulipennis Apfelbeck, 1918b
Trechus albanicus Apfelbeck, 1905 (valid species) instead Trechus albanicus Apfelbeck, 1907b
Ophonus krueperi Apfelbeck, 1902 (valid species) instead Ophonus krueperi (Apfelbeck, 1904)
Licinus graecus Apfelbeck, 1901 (valid species) instead Licinus cassideus graecus Apfelbeck, 1904
Molops malshentianus Apfelbeck, 1917 (valid species) instead Molops malshentianus Apfelbeck, 1918b
B.3. Nomina nuda found in both collections and literature
Eurynebria complanata orientalis Apfelbeck
Eurynebria complanata albanica Apfelbeck
Eurynebria complanata var. albanica Apfelbeck
? Nebria ganglbaueri ditana Apfelbeck (or incorrect spelling of Nebria ganglbaueri merditana
Apfelbeck, 1906)
Aptinus lugubris rufipes Apfelbeck (according to Apfelbeck, 1907a: 499, 501)
Bembidion merditanum Apfelbeck
Harpalus bischoffi Schauberger
Molops transfuga Apfelbeck (according to Mller, 1930: 124; Mlyn, 1977: 120)
Molops apfelbecki bischoffi J. Mller
Pterostichus djumanus Apfelbeck
204
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223
Index
(valid taxa are given in bold)
Abacopercus 138
Abax 138
Abax s.str. 139
abdominalis (Stenolophus) 119
abdominalis persicus (Stenolophus) 119
Acinopus 97
Acinopus s.str. 97
Acupalpus 12, 115
Acupalpus s.str. 115
acutangulus (Aptinus) 39
Adriaphaenops 79
aenea (Amara) 174
aeneocephalus (Chlaenius) 92
aeneocephalus aeneocephalus (Chlaenius) 92
aeneus (Dyschirius) 46
aeneus (Harpalus) 107
aeneus aeneus (Dyschirius) 46
aestuans (Notiophilus) 23
aetolica (Nebria) 18
aetolica galicica (Nebria) 18, 21
aetolica korabica (Nebria) 21
aetolica macedonica (Nebria) 18
aetolica papingutica (Nebria) 18, 21, 22
aetolica rambouseki (Nebria) 18
aetolica tomoricensis (Nebria) 21, 22
aetolicum (Bembidion) 63
aetolicum aetolicum (Bembidion) 63
aetolicus (Carabus) 30
aetolicus (Tapinopterus) 163
aetolicus peristericus (Tapinopterus) 162
aetolicus tomoricensis (Tapinopterus) 163
aetolus (Zabrus) 182
aetolus kodymi (Zabrus) 182
aetolus winkleri (Zabrus) 182
affinis (Harpalus) 107
affinis ab. limbopunctatus (Harpalus) 107
affinis ab. viridulus (Harpalus) 107
afrum (Agonum) 133
agnatus (Dyschirius) 46
224
Agonum 133
Agonum s.str. 133
albanensis (Carabus) 31
albanensis (Harpalus) 111
albanensis (Ophonus) 111
albanicola (Carabus) 34
albanicum (Bembidion) 68
albanicum (Hypogeobium) 160
albanicum (Platysma) 160
albanicus (Calathus) 169, 171
albanicus (Carabus) 35
albanicus (Cychrus) 37
albanicus (Harpalus) 103
albanicus (Molops) 142
albanicus (Omphreus) 132
albanicus (Ophonus) 111
albanicus (Pterostichus) 160
albanicus (Speluncarius) 160
albanicus (Tapinopterus) 160
albanicus (Trechus) 87
albanicus (Zabrus) 182, 183
albanicus albanicus (Zabrus) 182
albanicus latifianus (Zabrus) 182, 183
albanicus jablanicensis (Zabrus) 182
Albanotrechus 79
albipes (Paranchus) 137
alexandri (Brachinus) 40
alinae (Molops) 140
Alpaeus 18
alpestre (Agonum) 133
alpestris (Molops) 141
alpestris imitator (Molops) 141
Amara 12, 174
Amara s.str. 174
AMARINA 174
Amarocelia 177
ambiguus (Calathus) 169
ambiguus (Chlaenius) 91
ambiguus ambiguus (Calathus) 169
AMBLYSTOMINA 93
Amblystomus 93
ammophilus (Acinopus) 98
anatolicus (Harpalus) 104
anatolicus anatolicus (Harpalus) 104
Ancholeus 149
Anchomenus 136
Anchomenus s.str. 136
andreae (Bembidion) 68
andreae albanicum (Bembidion) 68
andreae bualei (Bembidion) 68
angularis (Cymindis) 121
angustatum (Agonum) 134
ANISODACTYLINA 94
Anisodactylus 12, 94
Anisodactylus s.str. 94
antennarium (Agonum) 136
antennarius (Platynus) 136
anthobia (Amara) 174
anthracinus (Pterostichus) 154
anthracinus anthracinus (Pterostichus) 154
anthracinus biimpressus (Pterostichus) 155
Anthracus 117
Antisphodrus 171
anxius (Harpalus) 102
apfelbecki (Leistus) 17
apfelbecki (Molops) 143
apfelbecki (Nebria) 19
apfelbecki (Pterostichus) 153
apfelbecki (Zabrus) 183
apfelbecki apfelbecki (Molops) 143
apfelbecki bischoffi (Molops) 143
apfelbecki corallipes (Molops) 142
apfelbecki kulti (Nebria) 19
apfelbecki plurisetosus (Molops) 145
apfelbecki pseudoalpestris (Molops) 142
apfelbecki reiseri (Molops) 142, 143
apfelbecki victoris (Molops) 142, 143
Aphaenopsis 79
Aphaon 155
apicalis (Dyschirius) 47
APOTOMINAE 51, 187
APOTOMINI 51
Apotomus 51
apricaria (Amara) 177
Apristus 127
225
227
circumscriptus (Chlaenius) 92
circumseptus (Calathus) 166
clathratus (Carabus) 31
clathratus auraniensis (Carabus) 31
Clivina 44
CLIVININA 44
CLIVININI 44
clypeatus (Aristus) 96
clypeatus (Ditomus) 96
clypeatus (Dixus) 96
Cnecostolus 43
coadunata (Cymindis) 121
coadunata coadunata (Cymindis) 121
coerulescens (Poecilus) 151
coerulescens (Pterostichus) 151
coeruleum (Bembidion) 57
coeruleum concoeruleum (Bembidion) 57
collaris (Bradycellus) 118
collaris (Clivina) 44
columbinus (Ophonus) 113
combustum (Bembidion) 60, 78
combustum combustum (Bembidion) 60
comita (Tapinopterus) 164
complanata (Amara) 180
complanata (Eurynebria) 16
complanata albanica (Eurynebria) 16
complanata orientalis (Eurynebria) 16
complanata var. albanica (Eurynebria) 16
complanatum (Bembidion) 55
concoeruleum (Bembidion) 57
concoeruleus (Ocydromus) 57
conforme (Bembidion) 55
connexus (Polystichus) 185
consularis (Amara) 178
convexicollis (Pogonistes) 78
convexior (Amara) 175
convexiusculus (Argutor) 153
convexiusculus (Pterostichus) 153
convexiusculus (Tapinopterus) 161
convexus (Carabus) 36
convexus dilatatus (Carabus) 36
convexus dilatatus n. miliusi (Carabus) 36
convexus dilatatus var. miliusi (Carabus) 36
convexus miliusi (Carabus) 36
convexus var. chionophilus (Carabus) 36
corax (Pterostichus) 160
228
decolor (Bembidion) 69
decorum (Bembidion) 63
decorum decorum (Bembidion) 63
dejeani (Chlaenius) 92
dejeani (Epomis) 92
dejeani (Parophonus) 114
deletum (Bembidion) 67
deletum deletum (Bembidion) 67
Deltomerus 76
Deltomerus s.str. 76
dentata (Drypta) 93
depressa (Siagona) 44
detonii (Tapinopterus) 164
diabrachys (Tachys) 76
diabrachys (Tachyura) 76
Diachromus 94
Dicheirotrichus 118
Dicheirotrichus s.str. 118
didonnai (Duvalius) 80
diffinis (Harpalus) 113
diffinis (Ophonus) 113
dilatatus (Badister) 129
diligens (Pterostichus) 153
dimidiatus (Harpalus) 105
dinaricus (Cychrus) 37
Dinodes 91
Diplocampa 58
discophorus (Acupalpus) 119
discophorus (Stenolophus) 119
distinctissimus (Trechus) 87
distinctus (Bradycellus) 118
distinguendus (Calathus) 167
distinguendus (Harpalus) 107
distinguendus distinguendus (Harpalus) 107
DITOMINA 95
Ditomus 95
Dixus 96
djumanus (Pterostichus) 158
dochii (Pterostichus) 164
dochii (Tapinopterus) 164
doderoi (Sinechostictus) 73
dorsale (Platynus) 136
dorsalis (Acupalpus) 117
dorsalis (Anchomenus) 136
dorsalis (Platynus) 136
INDEX
229
DROMIINA 123
Dromius 123
Dromius s.str. 123
droveniki (Duvalius) 79
Drypta 93
Drypta s.str. 93
DRYPTINI 93
duftschmidi (Agonum) 133
Duvalius 79
Duvalius s.str. 80
dyrrhachii (Bembidion) 63
DYSCHIRIINI 46
Dyschiriodes 46
Dyschirius 12, 46
Dyschirius s.str. 49
effluviorus (Sinechostictus) 73
Egadroma 119
egregius (Trechus) 87
ejaculans (Brachinus) 41
ELAPHRINAE 38, 187
ELAPHRINI 38
Elaphrus 38
elegans (Acupalpus) 115
elegans (Bembidion) 52, 53, 57, 58
elegans (Brachinus) 41
ellipticus (Calathus) 167
ellipticus ellipticus (Calathus) 167
elongatum (Bembidion) 73
elongatum tarsicum (Bembidion) 73
elongatum var. effluviorum (Bembidion) 73
elongatus (Laemostenus) 172
elongatus (Pterostichus) 152
elongatus (Sinechostictus) 73
elongatus pentheri (Laemostenus) 172
emarginatus (Acinopus) 97
Emphanes 58
eomita sic! (Tapinopterus) 164
ephippium (Bembidion) 62
epirensis (Carabus) 31
epiroticus (Pterostichus) 160
Epomis 92
equestris (Amara) 179, 180
equestris equestris (Amara) 179, 180
eremita (Ditomus) 96
eremita (Dixus) 96
erratica (Amara) 177
230
femoratum (Bembidion) 68
femoratum femoratum (Bembidion) 68
Feronidius 152
festivus (Chlaenius) 91
festivus festivus (Chlaenius) 91
fischeri (Calomera) 25
fischeri (Cicindela) 25
fischeri fischeri (Calomera) 25
fissuralis (Microlestes) 123
flavescens (Harpalus) 103
flavicornis (Harpalus) 101
flavicornis flavicornis (Harpalus) 101
flavipes (Asaphidion) 52, 53
flavipes (Chlaenius) 89
fodori (Duvalius) 82
foraminosum (Bembidion) 66
fossor (Clivina) 45
fossor fossor (Clivina) 45
franziniorum (Ophonus) 113
fuchsi (Duvalius) 81
fugax (Harpalus) 108
fulvibarbis (Leistus) 16
fulvibarbis fulvibarbis (Leistus) 16
fulvicollis (Paratachys) 74
fulvicollis (Tachys) 74
fulvipes (Amara) 181
fumigatum (Bembidion) 58
fusca (Amara) 180
fuscatus (Olisthopus) 137
fuscipes (Calathus) 167
fuscipes fuscipes (Calathus) 167
fuscipes graecus (Calathus) 167
fuscipes var. graecus (Calathus) 167
fuscipes var. latus (Calathus) 167
galilaeus (Lymnastis) 73
ganglbaueri (Brachinus) 41
ganglbaueri (Bradycellus) 118
ganglbaueri (Nebria) 20
ganglbaueri (Synuchidius) 171
ganglbaueri (Zabrus) 183
ganglbaueri ditana (Nebria) 21
ganglbaueri ganglbaueri (Nebria) 20
ganglbaueri ganglbaueri (Zabrus) 183
ganglbaueri gyalicaensis (Nebria) 20
ganglbaueri korabensis (Nebria) 20
ganglbaueri lonae (Zabrus) 183
231
intricatus (Carabus) 30
intricatus chinobatus (Carabus) 30
intricatus intricatus (Carabus) 30
intricatus intricatus m. montenegrinus (Carabus) 30
intricatus intricatus n. weiratheri (Carabus) 30
intricatus intricatus var. weiratheri (Carabus) 30
intricatus montenegrinus (Carabus) 30
intricatus var. montenegrinus (Carabus) 30
irenis (Trechus) 86
iricolor (Bembidion) 70
irregularis (Carabus) 35
irregularis bucephalus (Carabus) 35
irregularis ramanus (Carabus) 35
janthinus (Laemostenus) 172
janthinus janthinus (Laemostenus) 172
javurkovae (Bembidion) 68
jockischi (Nebria) 22
jockischi jockischi (Nebria) 22
jonicus (Carabus) 34
joniensis (Carabus) 35
justinae (Bembidion) 55
kaufmanni (Tapinopterus) 161
klisuranus (Molops) 144, 146
kobingeri (Trechus) 87
kobingeri bulgaricus (Trechus) 88
kobingeri kobingeri (Trechus) 87, 88
kobingeri pawlowskianus (Trechus) 88
korabensis (Carabus) 33
korabica (Nebria) 21
korax (Calathus) 167, 168
koyi (Poecilus) 150
koyi (Pterostichus) 150
koyi dinaricus (Pterostichus) 150
koyi koyi (Poecilus) 150
kratteri (Nebria) 23
kratteri kratteri (Nebria) 23
kratteri valonensis (Nebria) 23
krueperi (Harpalus) 109
krueperi (Ophonus) 109
Laemostenus 171
Laemostenus s.str. 172
laeviceps (Parophonus) 114, 115
laevicollis (Trichotichnus) 115
laevicollis laevicollis (Trichotichnus) 115
laevifrons (Clivina) 45
laevigatus (Scarites) 50
233
Lymnastis 73
macedonica (Nebria) 18, 21
macedonica galicica (Nebria) 18, 21
macedonica papingutica (Nebria) 18, 21
macedonica rambouseki (Nebria) 18, 22
macedonicus (Calathus) 168
macedonicus (Pachycarus) 97
macroderus (Dyschirius) 48
macroderus macroderus (Dyschirius) 48
maculatum (Bembidion) 72
maculatum maculatum (Bembidion) 72
maculatus (Acupalpus) 116, 117
maculicornis (Harpalus) 114
maculicornis (Ophonus) 114
maculicornis (Parophonus) 114
maculicornis (Trichotichnus) 114
maderi (Molops) 139
maglicensis (Duvalius) 79
magnicollis (Leistus) 17
malissorum (Carabus) 31, 32
malissorum (Deltomerus) 76, 77
malissorum (Neotrechus) 83, 84
malissorum (Molops) 145
malissorum (Pterostichus) 156, 158, 159
malissorum (Trechus) 84
malissorum ab. munella (Pterostichus) 157
malshentianus (Molops) 139
mannerheimii (Bembidion) 70
Manodromius 125
marginata (Egadroma) 119
marginatum (Agonum) 134
marginatus (Stenolophus) 119
maritimus (Lionychus) 127
MASOREINA 92
Masoreus 92
Masoreus s.str. 92
maurus (Microlestes) 124
maurus maurus (Microlestes) 124
megacephalus (Acinopus) 98
Megodontus 29, 31
meisteri (Pterostichus) 156
melanarius (Pterostichus) 153
melanarius melanarius (Pterostichus) 153
melancholica (Myriochila) 28
melancholica melancholica (Myriochila) 28
Melanius 152
235
Oodes 131
OODINI 131
Ophonomimus 113
Ophonus 12, 109
Ophonus s.str. 112
Oreocarabus 33
Oreoplatysma 153
Osimus 98
osmanilis (Molops) 145
osmanilis osmanilis (Molops) 145, 146
osmanilis tomoricensis (Molops) 145, 146
ottomanus (Pterostichus) 154
ottomanus ottomanus (Pterostichus) 154
ovalis (Abax) 139
ovata (Amara) 176
Pachycarus 97
Pachystus 34
pallipes (Metabletus) 128
pallipes (Syntomus) 128
paludicola (Acupalpus) 116, 117
palustris (Notiophilus) 24
PANAGAEINI 133
Panagaeus 133
Panagaeus s.str. 133
Paracelia 179
Paradeltomerus 77
paradoxus (Deltomerus) 77
paradoxus korabensis (Deltomerus) 77
paradoxus paradoxus (Deltomerus) 77
Paradromius 125
Paradyschirius 50
Parahaptoderus 153
Parallelomorphus 50
parallelus (Dyschirius) 50
parallelus (Ophonus) 111
parallelus albanensis (Harpalus) 111
parallelus albanicus (Ophonus) 111
parallelus ruficornis (Dyschiriodes) 50
parallelus ruficornis (Dyschirius) 50
Paranchus 137
Parapercosia 179
Paratachys 74
parnassicola (Molops) 140
parnassicola alinae (Molops) 140
parnassicola valonensis (Molops) 140
parnassicus (Carabus) 35
237
robustus (Odontocarus) 97
rossii (Asaphidion) 53
rostratus (Stomis) 161
rostratus rostratus (Stomis) 161
rotundatus (Olisthopus) 137
rotundatus rotundatus (Olisthopus) 137
rotundicollis (Carterus) 95
rubripes (Harpalus) 99
rufescens (Nebria) 22
rufescens rufescens (Nebria) 22
rufibarbis (Harpalus) 112
rufibarbis (Ophonus) 112
ruficornis (Dyschirius) 50
rufipalpis (Harpalus) 100
rufipalpis rufipalpis (Harpalus) 100
rufipes (Harpalus) 108
rufipes (Molops) 146
rufipes (Notiophilus) 25
rufipes (Pseudophonus) 108
rufipes klisuranus (Molops) 146
rufipes munelensis (Molops) 146
rufipes pseudoosmanilis (Molops) 146
rufipes rufipes (Molops) 146
rufipes steindachneri (Molops) 147
rufipes sturanyi (Molops) 147
rufitarsis (Harpalus) 100
rufithorax (Apotomus) 51
rufomarginatus (Leistus) 18
rufus (Apotomus) 51
rufus (Platyderus) 166
rufus rufus (Platyderus) 166
rugosicollis (Myas) 148
rugosus (Carabus) 35
rumelicum (Bembidion) 63
rupicola (Harpalus) 112
rupicola (Ophonus) 112
sabulicola (Harpalus) 113
sabulicola (Ophonus) 113
sabulicola columbinus (Harpalus) 113
sabulicola columbinus (Ophonus) 113
sahlbergi albanica (Cicindela) 27
salina (Nebria) 23
salinus (Dyschirius) 49
salinus striatopunctatus (Dyschirius) 49
saphyrea (Amara) 177
sarajevoensis (Carabus) 31
INDEX
239
sterbai (Carabus) 34
stictus (Ophonus) 113
stierlini (Asaphidion) 53
stolfai (Bembidion) 65, 66
Stomis 160
Stomis s.str. 160
strenuus (Argutor) 154
strenuus (Pterostichus) 154
strepens (Brachinus) 42
striatopunctatus (Poecilus) 150
striatul sic! (Bembidion) 66
striatum (Bembidion) 66
strumosus (Dyschirius) 46
strupii (Duvalius) 82
sturanyi (Duvalius) 81, 82
sturanyi (Molops) 147
sturanyi (Nebria) 19
sturanyi matejkai (Duvalius) 82
sturanyi munelensis (Molops) 146
sturanyi stenocephalus (Duvalius) 81, 82
sturanyi sturanyi (Duvalius) 82
subaeneus (Apristus) 127
subchalybaeus (Harpalus) 101
subcostatum (Bembidion) 68
subcostatum javurkovae (Bembidion) 68, 69
subcostatum spartanum (Bembidion) 69
subcostatum vau (Bembidion) 68, 69
subcylindricus (Harpalus) 102
subfasciatum (Bembidion) 71
submarginatus (Licinus) 129, 130
subnotatus (Trechus) 85
subnotatus ljubetensis (Trechus) 86
subnotatus subnotatus (Trechus) 85
subquadratus (Acinopus) 97, 98
subquadratus (Ophonus) 110
subrugosus (Carabus) 35
subsinuatus (Harpalus) 100
substriatum (Bembidion) 62
substriatus (Dyschiriodes) 50
substriatus (Dyschirius) 50
substriatus (Notiophilus) 25
substriatus priscus (Dyschirius) 50
sulphuripes (Harpalus) 100
sulphuripes sulphuripes (Harpalus) 100
sutomorensis (Carabus) 30
suturalis (Acupalpus) 117
suturalis (Neotrechus) 84
suturalis (Trechus) 84
suturalis pentheri (Neotrechus) 84
suturalis pretneri (Neotrechus) 83
suturalis suturalis (Acupalpus) 117
sycophanta (Calosoma) 29
sylvatica (Cicindela) 27
sylvatica sylvatica (Cicindela) 27
Syntomus 12, 127
Synuchidius 171
SYNUCHINA 173
Synuchus 173
Synuchus s.str. 173
TACHYINA 73
TACHYINI 10
Tachypus 36
Tachys 75
Tachyta 75
Tachyta s.str. 75
Tachyura 76
Tachyura s.str. 76
Taenidia 26
Talanes 71
Tapinopterus 12, 161
Tapinopterus s.str. 161
tardus (Harpalus) 103
tarsicus (Sinechostictus) 73
Tarulus 123
taurica (Amara) 179
tenebrioides (Zabrus) 185
tenebrioides tenebrioides (Zabrus) 185
tenebrosus (Harpalus) 98
tenebrosus var. centralis (Harpalus) 98
tenellum (Bembidion) 59
tenellum tenellum (Bembidion) 59
terminale (Bembidion) 71
terminatus (Chlaenius) 90
Terminophanes 71
terricola (Laemostenus) 173
terricola (Pristonychus) 173
terricola (Scarites) 51
terricola punctatus (Laemostenus) 173
terricola terricola (Scarites) 51
testaceum (Bembidion) 60
testaceum testaceum (Bembidion) 60
Testedium 71
INDEX
241
tethys (Bembidion) 70
tetracolum (Bembidion) 69
tetracolum tetracolum (Bembidion) 69
teutonus (Acupalpus) 120
teutonus (Stenolophus) 120
Thalassophilus 88
thessalicus tomoricensis (Tapinopterus) 163
thoreyi (Agonum) 136
thoreyi (Platynus) 136
thoreyi thoreyi (Agonum) 136
tibiale (Bembidion) 54
tibialis (Amara) 177
Tomocarabus 36
tomoricensis (Amara) 179, 180
tomoricensis (Molops) 145
tomoricensis (Nebria) 21
Trachycarabus 37
transfuga (Molops) 143
transversalis (Cicindela) 27
transversalis transversalis (Cicindela) 27
TRECHINA 79
TRECHINAE 7, 52, 187
TRECHINI 79
TRECHINITAE 10
TRECHODINA 88
Trechus 85
Trechus s.str. 85
Trepanes 72
Trichochlaenius 92
Trichotichnus 12, 115
Trichotichnus s.str. 115
tricolor (Bembidion) 56
tricuspidata (Amara) 181
tricuspidatus (Carterus) 95
tricuspidatus (Ditomus) 95
trimaculata (Lebia) 126
triseriatus (Harpalus) 105
trisignata (Cylindera) 28
trisignata trisignata (Cylindera) 28
tristis (Chlaenius) 90
tristis (Harpalus) 104
tristis (Trechus) 87
tristis tristis (Chlaenius) 90
truncatellus (Metabletus) 128
truncatellus (Syntomus) 128
turcicus (Platynus) 138
242
uliginosus (Elaphrus) 38
unipustulatus (Badister) 128
ustulatum (Bembidion) 69
valbonensis (Trechus) 85
valonensis (Nebria) 23
vaporariorum (Cymindis) 123
varicolor (Bembidion) 56
varicolor varicolor (Bembidion) 56
variolosus (Carabus) 31
variolosus nodulosus (Carabus) 31
varium (Bembidion) 63
varium dyrrhachii (Bembidion) 63
velutinus (Chlaenius) 91
venustus (Laemostenus) 172
verbasci (Bradycellus) 118
vernalis (Harpalus) 102
vernalis cursor (Pterostichus) 151
versicolor (Poecilus) 151
vestitus (Chlaenius) 90
victoris (Molops) 143, 144
viduus moestus (Platynus) 133
viduus var. moestus (Platynus) 133
vignai (Duvalius) 82
violaceus (Carabus) 32
violaceus azurescens (Carabus) 32
violaceus azurescens m. merditana (Carabus) 33
violaceus azurescens m. merditanus (Carabus) 33
violaceus azurescens m. zabljakensis (Carabus) 32
violaceus azurescens n. zabljakensis (Carabus) 32
violaceus azurescens var. shljebensis (Carabus) 32
violaceus dryas (Carabus) 33
violaceus dryas var. korabensis (Carabus) 33
violaceus dryas var. merditanus (Carabus) 33
violaceus exasperatus (Carabus) 33
violaceus merditanus (Carabus) 33
violaceus shardagensis (Carabus) 33
violaceus var. korabensis (Carabus) 33
violaceus var. shardagensis (Carabus) 33
violaceus vlasuljensis (Carabus) 33
violaceus zabljakensis v. merditanus (Carabus) 33
viridicupreum (Agonum) 135
viridicupreum viridicupreum (Agonum) 135
viridicupreus (Platynus) 135
viridulus (Harpalus) 107
vivalis (Synuchus) 173
vivalis vivalis (Synuchus) 173
vlasuljensis (Carabus) 33
vseteckai (Bembidion) 62
vseteckai dissimile (Bembidion) 62
walteri (Pterostichus) 156
weiratheri (Carabus) 30
weiratheri (Molops) 140
weiratherianus (Carabus) 35
wetterhallii (Masoreus) 92
wetterhallii wetterhallii (Masoreus) 92
winkleri (Carabus) 35
winkleri (Zabrus) 182
winklerianus (Molops) 144
Xenocelia 180
zabljakensis (Carabus) 32
ZABRINA 181
ZABRINI 174
Zabrus 181
Zabrus s.str. 185
Zezea 181
zhalovi (Duvalius) 82
ZUPHIINA 185
ZUPHIINI 185
Zuphium 186
Zuphium s.str. 186
INDEX
243
244