Sie sind auf Seite 1von 42

Resilience and sustainability of resource-economic

systems
Martin F. Quaas

July 10, 2014

Martin Quaas
University of Kiel Resilience and sustainability of resource-economic systems

1/28

INTERACTIONS IN MULTI-SPECIES FISHERIES


ecological interactions: predator-prey, competition, symbiosis
economic interactions: technical (bycatch), demand-side

Martin Quaas
University of Kiel Resilience and sustainability of resource-economic systems

2/28

INTERACTIONS IN MULTI-SPECIES FISHERIES


ecological interactions: predator-prey, competition, symbiosis
economic interactions: technical (bycatch), demand-side

global trade in fishery products (FAO SOFIA 2014, Fig. 19)


Martin Quaas
University of Kiel Resilience and sustainability of resource-economic systems

2/28

OVERFISHING AND COLLAPSE OF FISH STOCKS

collapse of stocks at world-wide scale


has become focus of scientific interest
and public concern
Worm et al. (2006, Science)

Costello et al. (2008, Science)


Heal and Schlenker (2008, Nature)
empirical definition: a stock is
collapsed if catch is below z% of
historic maximum (z = 3, 6, 10)
Costello et al. (2008)
Quaas/Requate (2013). Sushi or Fish Fingers? Seafood Diversity, Collapsing Fish Stocks,
and Multi-species Fishery Management. Scandinavian Journal of Economics.
3/28

OVERFISHING AND COLLAPSE OF FISH STOCKS


consumer value seafood diversity: love of variety


collapse of fish stocks is economic problem on top of
inefficiently low stocks and yields from individual fisheries

Quaas/Requate (2013). Sushi or Fish Fingers? Seafood Diversity, Collapsing Fish Stocks,
and Multi-species Fishery Management. Scandinavian Journal of Economics.
4/28

CONSUMER PREFERENCES FOR SEAFOOD DIVERSITY

quasi-linear in income y and fish consumption


sub-utility function for fish captures love of variety (Dixit and Stiglitz 1977)
U(y , h1 , . . . , hi , . . . , hn ) = y + ln

n
X

hi

i=1

hi 0

consumption of fish species i

>1

elasticity of substitution between fish species

>0

constant aggregate expenditures for fish

Quaas/Requate (2013). Sushi or Fish Fingers? Seafood Diversity, Collapsing Fish Stocks,
and Multi-species Fishery Management. Scandinavian Journal of Economics.
5/28

MULTI-SPECIES MODEL
consumer demand for species i
hit =

pit
n
P
pjt1

j=1

pjt market price for species j at time t

from estimates of own and cross-price elasticities for


crustaceans and salmon (Asche et al. 1997): = 1.66

Quaas/Requate (2013). Sushi or Fish Fingers? Seafood Diversity, Collapsing Fish Stocks,
and Multi-species Fishery Management. Scandinavian Journal of Economics.
6/28

MULTI-SPECIES MODEL
consumer demand for species i
hit =

pit
n
P
pjt1

j=1

pjt market price for species j at time t

from estimates of own and cross-price elasticities for


crustaceans and salmon (Asche et al. 1997): = 1.66
equation of motion for stock xi of fish species i


xit
hit
x it = i xit 1
i
i intrinsic growth rate of species i
i carrying capacity of species i, normalized to i = 1
hi harvest of species i

note: no ecological interaction between species


Quaas/Requate (2013). Sushi or Fish Fingers? Seafood Diversity, Collapsing Fish Stocks,
and Multi-species Fishery Management. Scandinavian Journal of Economics.
6/28

MULTI-SPECIES MODEL
consumer demand for species i
hit =

pit
n
P
pjt1

j=1

pjt market price for species j at time t

from estimates of own and cross-price elasticities for


crustaceans and salmon (Asche et al. 1997): = 1.66
equation of motion for stock xi of fish species i
x it = i xit (1 xit ) hit
i intrinsic growth rate of species i
i carrying capacity of species i, normalized to i = 1
hi harvest of species i

note: no ecological interaction between species


Quaas/Requate (2013). Sushi or Fish Fingers? Seafood Diversity, Collapsing Fish Stocks,
and Multi-species Fishery Management. Scandinavian Journal of Economics.
6/28

MULTI-SPECIES MODEL
harvesting cost
ci (xi , hi ) = ci xii hi
ci > 0 and 0 < i 1

note: no technical interactions; perfectly malleable capital


Gordon-Schaefer: i = 1
species/stock
38 cod fisheries (mean)
98 flatfish fisheries (mean)
Baltic cod
bigeye tuna
North Sea herring

value for i
0.75
0.65
0.64
0.60
0.56

reference
Harley, Myers, and Dunn (2001)
Harley, Myers, and Dunn (2001)
Kronbak (2005:472)
Grafton et al. (2007, SI p. 7)
Bjrndal and Conrad (1987),
Nostbakken and Bjorndal (2003:351)
pollock
0.50
Quinn and Deriso (1999:28)
Northern anchovy
0.39
Opsomer and Conrad (1994:29)
Australian Northern prawn 0.40
Grafton et al. (2007)
yellow fin tuna
0.23
Grafton et al. (2007)
Quaas/Requate (2013). Sushi or Fish Fingers? Seafood Diversity, Collapsing Fish Stocks,
and Multi-species Fishery Management. Scandinavian Journal of Economics.
7/28

OPEN ACCESS
zero-profit for fishing firms


pi hi ci xii hi = pi ci xii hi = 0
consumer demand for species i
hi =

pi
n
P
pj1

j=1

Quaas/Requate (2013). Sushi or Fish Fingers? Seafood Diversity, Collapsing Fish Stocks,
and Multi-species Fishery Management. Scandinavian Journal of Economics.
8/28

OPEN ACCESS
zero-profit for fishing firms


pi hi ci xii hi = pi ci xii hi = 0
consumer demand for species i
hi =

pi
n
P
pj1

j=1

market equilibrium: harvest of species i


hi =

ci xi i
n
P
j=1

(1) j

cj1 xj

Quaas/Requate (2013). Sushi or Fish Fingers? Seafood Diversity, Collapsing Fish Stocks,
and Multi-species Fishery Management. Scandinavian Journal of Economics.
8/28

OPEN ACCESS
zero-profit for fishing firms


pi hi ci xii hi = pi ci xii hi = 0
consumer demand for species i
hi =

pi
n
P
pj1

j=1

market equilibrium: harvest of species i


hi =

ci xi i
n
P
j=1

(1) j

cj1 xj

xi

n
c P
(1)
xj
j=1

assume ci = c and i = for all i for theoretical analysis


Quaas/Requate (2013). Sushi or Fish Fingers? Seafood Diversity, Collapsing Fish Stocks,
and Multi-species Fishery Management. Scandinavian Journal of Economics.
8/28

OPEN ACCESS
Lemma
Assuming 1 < 2 < ... < n , steady-state stocks under open
access are ordered according to intrinsic growth rates,
x1oa x2oa . . . xnoa .

stocks xi

0.8
0.7
0.6
0.5
0.4
0.3
species 1
species 2
species 3
0.1
species 4
species 5
0
1 1.5 2 2.5 3 3.5 4 4.5 5
0.2

elasticity of substitution
Quaas/Requate (2013). Sushi or Fish Fingers? Seafood Diversity, Collapsing Fish Stocks,
and Multi-species Fishery Management. Scandinavian Journal of Economics.
9/28

OPEN ACCESS
Lemma
Assuming 1 < 2 < ... < n , steady-state stocks under open
access are ordered according to intrinsic growth rates,
x1oa x2oa . . . xnoa .

Proposition
For all there exists some species i0 with 1 i0 < n such that
such in the neighborhood of we have:
dxioa
>0
d
dxioa
0
d
dxioa
<0
d

for all i < i0


for i = i0
for all i > i0

stocks xi

0.8
0.7
0.6
0.5
0.4
0.3
species 1
species 2
species 3
0.1
species 4
species 5
0
1 1.5 2 2.5 3 3.5 4 4.5 5
0.2

elasticity of substitution
Quaas/Requate (2013). Sushi or Fish Fingers? Seafood Diversity, Collapsing Fish Stocks,
and Multi-species Fishery Management. Scandinavian Journal of Economics.
9/28

OPEN ACCESS

stocks xi

0.8
0.7
0.6
0.5
0.4
0.3
species 1
species 2
species 3
0.1
species 4
species 5
0
1 1.5 2 2.5 3 3.5 4 4.5 5
0.2

elasticity of substitution

Proof:

OPEN ACCESS

Proof:
harvest

differentiate open-access

hi =

xi

n
c P
(1)
xj
j=1

stocks xi

0.8

with respect to :

0.7
0.6

0.5

hioa
= hioa
ln xi

0.4
0.3
species 1
species 2
species 3
0.1
species 4
species 5
0
1 1.5 2 2.5 3 3.5 4 4.5 5
0.2

elasticity of substitution

n
P
j=1

(1)

xj

n
P
j=1

ln xj

(1)

xj

OPEN ACCESS

Proof:
harvest

differentiate open-access

hi =

xi

n
c P
(1)
xj
j=1

stocks xi

0.8

with respect to :

0.7
0.6

0.5

hioa
= hioa
ln xi

0.4
0.3
species 1
species 2
species 3
0.1
species 4
species 5
0
1 1.5 2 2.5 3 3.5 4 4.5 5
0.2

elasticity of substitution

n
P
j=1

(1)

xj

n
P
j=1

ln xj

(1)

xj

differentiate steady-state condition


i xi (1 xi ) = hi implictly w.r.t. :


dhioa dxioa
hioa
oa
i (1 2 xi ) oa
=
dxi
d

{z
}
|
<0 (stability)

PREFERENCES FOR DIVERSITY MAY EXPLAIN CASCADING


COLLAPSE OF FISH STOCKS UNDER OPEN ACCESS

stocks xi

0.8

0.6

0.4
5 = 0.6
4 = 0.4
3 = 0.3
2 = 0.26
1 = 0.25

0.2

0
0

20

40

60

80

100

120

140

160

time t
high elasticity of substitution = 2

low elasticity of substitution = 1.2

Quaas/Requate (2013). Sushi or Fish Fingers? Seafood Diversity, Collapsing Fish Stocks,
and Multi-species Fishery Management. Scandinavian Journal of Economics.
11/28

PREFERENCES FOR DIVERSITY MAY EXPLAIN CASCADING


COLLAPSE OF FISH STOCKS UNDER OPEN ACCESS

stocks xi

5 = 0.6
4 = 0.4
3 = 0.3
2 = 0.26
1 = 0.25

0.8

0.6

0.4
5 = 0.6
4 = 0.4
3 = 0.3
2 = 0.26
1 = 0.25

0.2

0
0

20

40

60

80

100

120

140

time t
high elasticity of substitution = 2

160

20

40

60

80

100

120

140

160

time t

low elasticity of substitution = 1.2

Quaas/Requate (2013). Sushi or Fish Fingers? Seafood Diversity, Collapsing Fish Stocks,
and Multi-species Fishery Management. Scandinavian Journal of Economics.
11/28

OPEN ACCESS
Proposition
If the stock of some species collapses, it will do so in finite time.

Proposition
Assume 1 < 2 < ... < n and consider n species in an initial
open-access equilibrium at some given level of . If
(n + 1 i) i <

(2 )2
c (1 )1

for species i = 1, . . . , n

a threshold value (1 < < ) for the elasticity of substitution


exists such that the stocks of the n (1 n n) least resilient
species will sequentially collapse under open access fishing if the
elasticity of substitution falls from to a level equal to or below .

Quaas/Requate (2013). Sushi or Fish Fingers? Seafood Diversity, Collapsing Fish Stocks,
and Multi-species Fishery Management. Scandinavian Journal of Economics.
12/28

STRONGER PREFERENCES FOR DIVERSITY:


MORE STOCKS COLLAPSE ON AVERAGE
i

open-access harvest of species i: hi =

n
P
j=1

ci xi

(1) j

cj1 xj

percentage of steady-state stocks

open-access equilibrium stocks of all species with i > 1


strictly positive.
100
Monte-Carlo simulation
80

50 samples for 33 stocks

60

randomly drawn intrinsic


growth rates i

40

randomly drawn unit cost


ci of harvesting

randomly drawn stock


optimal # of stocks
lower bound on optimal # of stocks
elasticities of harvesting i
open-access # of stocks
lower bound on open-access # of stocks
0
1
1.4
1.8
2.2
2.6
3
elasticity of substitution
Quaas/Requate (2013). Sushi or Fish Fingers? Seafood Diversity, Collapsing Fish Stocks,
and Multi-species Fishery Management. Scandinavian Journal of Economics.
13/28
20

EMPIRICAL TEST: FIRST STAGE


data: RAM legacy database (Ricard et al. 2012)
use time series of harvest (hit ) and spawning stock biomass
(for xit ) from stock assessments for n = 177 fisheries
assume open-access conditions before 1992
estimation for stock i
ln(hit ) = ln(i ) + i t + i ln(xit ) + it

point estimate
for i

Fricke/Quaas (2014). The Economic Causes of Regime Shifts in Marine Ecosystems.


Manuscipt, University of Kiel.
14/28

EMPIRICAL TEST: SECOND STAGE


are stocks with i < 1 more likely to collapse?
define dummy

D0<i <1 =

1 if 0 < i < 1
0 else

Fricke/Quaas (2014). The Economic Causes of Regime Shifts in Marine Ecosystems.


Manuscipt, University of Kiel.
15/28

EMPIRICAL TEST: SECOND STAGE


are stocks with i < 1 more likely to collapse?
define dummy

D0<i <1 =

1 if 0 < i < 1
0 else

second stage: probit


P(Dcollapse = 1) = F (1 + 2 D0<i <1 + . . .)
collapse dummy Dcollapse = Dz%
Worm et al. (2006), Costello et al. (2008): catch hit below z%
of historic maximum
here: stock xit below z% of historic maximum at any t 2002

Fricke/Quaas (2014). The Economic Causes of Regime Shifts in Marine Ecosystems.


Manuscipt, University of Kiel.
15/28

EMPIRICAL TEST: SECOND STAGE


P(Dz% = 1)

D10%

D6%

D3%

Marginal Effects (at means)


D0<i <1
ln
i
i

0.12**
-0.02
-0.49

regressor #
log likelihood
n
ncollapsed
bootstrap
replications

4
-85.41
177
36
1000

0.08

2
-88.53
177
36
1000

0.08***
-0.01**
-0.70**
4
-49.05
177
16
900

0.08***

2
-52.06
177
16
900

0.06***
-0.01***
-0.58***
4
-37.09
177
11
683

0.06***

2
-39.59
177
11
683

Significance *** 1% ** 5% * 10%. OLS in first, probit in second stage.

Fricke/Quaas (2014). The Economic Causes of Regime Shifts in Marine Ecosystems.


Manuscipt, University of Kiel.
16/28

FIRST-BEST management of all species

SECOND-BEST management of one species k


other species i 6= k harvested under open access
effects of management on other stocks taken into account

MYOPIC management of one species


other species harvested under open access
effects of management on other stocks ignored

Quaas/Requate (2013). Sushi or Fish Fingers? Seafood Diversity, Collapsing Fish Stocks,
and Multi-species Fishery Management. Scandinavian Journal of Economics.
17/28

FIRST-BEST management of all species


maximize present-value of consumer well-being
Z
max

y + ln

y ,{hi }
t=0

X

hj

!
e t dt

taking stock dynamics of all species into account


> 0: social discount rate
2

SECOND-BEST management of one species k


other species i 6= k harvested under open access
effects of management on other stocks taken into account
constraint to optimization:

1
X 1
1
1

hk
= ci xii
hi
+
hj

for all species i 6= k

j6=k

MYOPIC management of one species


other species harvested under open access
effects of management on other stocks ignored
prices of other species assumed to remain at current levels

Quaas/Requate (2013). Sushi or Fish Fingers? Seafood Diversity, Collapsing Fish Stocks,
and Multi-species Fishery Management. Scandinavian Journal of Economics.
17/28

stocks xi

MANAGEMENT SCENARIOS
second-best management of one fish species must be less
strict than first-best managment of the same species
myopic management of one fish species is too strict compared
to second-best managment of the same species
myopic management may lead to collapse of fish stocks that
would not collapse under open access
first-best

second-best

myopic

0.8
0.7
0.6
0.5
0.4
0.3
species
species
species
species
species

0.2
0.1
0

species
species
species
species
species

1
2
3
4
5

50

100

time t

150

1
2
3
4
5

50

species
species
species
species
species

100

time t

150

1
2
3
4
5

100

200

300

time t

400

500

Model of a natural-resource-dependent economy


two resources, identical growth and harvesting cost functions
dynamically optimal harvest of natural resource
Z 
X
 1 
1
e t dt
max
y + ln
hj1
0

j=1,2

1 > 0: degree of complementarity


> 0: social discount rate
> 1: complements
consumption res. 2, h2

consumption res. 2, h2

< 1: substitutes

consumption res. 1, h1

consumption res. 1, h1

Quaas/van Soest/Baumg
artner (2013). Complementarity, impatience, and the resilience
of natural-resource-dependent economies. J. Environmental Economics and Management.19/28

pictures: http://en.wikipedia.org/wiki/Easter island

(higher-level) ecosystem services are often complements for well-being

SUBSTITUTES < 1
LOW DISCOUNT RATE < 0
1

< 1; > 0

stock of resource 2, x2

< 1; < 0

stock of resource 2, x2

HIGH DISCOUNT RATE > 0


1

A
O
0

A0

O
1

stock of resource 1, x1

stock of resource 1, x1

Quaas/van Soest/Baumg
artner (2013). Complementarity, impatience, and the resilience
of natural-resource-dependent economies. J. Environmental Economics and Management.21/28

SUBSTITUTES < 1
LOW DISCOUNT RATE < 0
1

< 1; > 0

stock of resource 2, x2

< 1; < 0

stock of resource 2, x2

HIGH DISCOUNT RATE > 0


1

A
O
0

A0

O
1

stock of resource 1, x1

stock of resource 1, x1

> 0 : A = A0 = (0, 0) for < 1

Quaas/van Soest/Baumg
artner (2013). Complementarity, impatience, and the resilience
of natural-resource-dependent economies. J. Environmental Economics and Management.21/28

degree of complementarity
()

III

IV

II

0
social discount rate

>
() > 1: symmetric steady state is locally unstable

Quaas/van Soest/Baumg
artner (2013). Complementarity, impatience, and the resilience
of natural-resource-dependent economies. J. Environmental Economics and Management.22/28

stock of resource 2, x2

stock of resource 2, x2

>
(); < 0
A

III

>
(); > 0

IV
S

A0
0

O
0

O
0

1<<
(); < 0

stock of resource 1, x1
stock of resource 2, x2

stock of resource 2, x2

stock of resource 1, x1

O
0

stock of resource 1, x1

1
C

1<<
(); > 0

II
S

A0

C0
1

stock of resource 1, x1

RESILIENCE MANAGEMENT
resources hit by one-time random shock = (1 , 2 ) with
bounded support at unkown time T (constant hazard rate )
(x1T , x2T ) (x1T + 1 , x2T + 2 )

Quaas/van Soest/Baumg
artner (2013). Complementarity, impatience, and the resilience
of natural-resource-dependent economies. J. Environmental Economics and Management.24/28

RESILIENCE MANAGEMENT
resources hit by one-time random shock = (1 , 2 ) with
bounded support at unkown time T (constant hazard rate )
(x1T , x2T ) (x1T + 1 , x2T + 2 )
optimal resilience management:

max E

 ZT 
0

y + ln

X

hj1

1
1

e t dt + V (x1T + 1 , x2T + 2 )

j=1,2

Quaas/van Soest/Baumg
artner (2013). Complementarity, impatience, and the resilience
of natural-resource-dependent economies. J. Environmental Economics and Management.24/28

RESILIENCE MANAGEMENT
resources hit by one-time random shock = (1 , 2 ) with
bounded support at unkown time T (constant hazard rate )
(x1T , x2T ) (x1T + 1 , x2T + 2 )
optimal resilience management:

max E

 ZT 
0

y + ln

X

hj1

1
1

e t dt + V (x1T + 1 , x2T + 2 )

j=1,2

value function of post-shock optimization problem


V (x1T + 1 , x2T + 2 )
Z 
X
 1 
1
= max
y + ln
hj1
e t dt
T +dt

j=1,2

Quaas/van Soest/Baumg
artner (2013). Complementarity, impatience, and the resilience
of natural-resource-dependent economies. J. Environmental Economics and Management.24/28

resource stocks xi

resource stocks xi

0.33

x1? (t) = x2? (t)

0.32

0.31

0.3

0.33

x1? (t) = x2? (t)

0.32

0.31

0.3

xS
0.29

xS

= 1.5, = 0.17
0

100

200

300

400

0.29

500

= 1.8, = 0.17
0

100

time t

200

300

400

stock of resource 2, x2

stock of resource 2, x2

1
C

C0

C0

500

time t

1
stock of resource 1, x1

1
stock of resource 1, x1

RESILIENCE MANAGEMENT
0.18

probability of regime shift

0.16
0.14
0.12
0.1
0.08
0.06
0.04
0.02
0
1.5

1.55

1.6
1.65
1.7
degree of complementarity,

1.75

1.8

Quaas/van Soest/Baumg
artner (2013). Complementarity, impatience, and the resilience
of natural-resource-dependent economies. J. Environmental Economics and Management.26/28

SUMMARY AND CONCLUSIONS


preferences for seafood diversity and collapse of fish stocks
preferences for seafood diversity may explain cascading
collapse of fish stocks under open access
strong consumer preference for seafood diversity may lead to
low fish biodiversity
the difference between the equilibrium and optimal number of
stocks incerases with preferences for diversity

complementarity, limited resilience and collapse of economies


optimality < resilience
resilience < sustainability
(Derissen, Quaas, and Baumgartner 2011)
the coupled resource-economic system may exhibit limited
resilience, although objective is concave and ecological
dynamics are logistic

ignoring demand-side interactions may lead to false optimism


with regard to sustainability of natural resource use
Martin Quaas
University of Kiel Resilience and sustainability of resource-economic systems

27/28

Asche, F., F. Steen, and K. G. Salvanes (1997). Market Delineation and Demand Structure. American Journal of Agricultural Economics,
79(1): 139150.

Bjrndal, T. and J. M. Conrad (1987). The Dynamics of an Open Access Fishery. The Canadian Journal of Economics, 20(1): 7485.
Clark, C. W. (1990). Mathematical Bioeconomics. Wiley, New York, 2nd edition.
Costello, C., S. D. Gaines, and J. Lynham (2008). Can Catch Shares Prevent Fisheries Collapse? Science, 321(5896): 16781681.

Derissen, S., M. F. Quaas, and S. Baumg


artner (2011). The relationship between resilience and sustainable development of ecologicaleconomic systems. Ecological Economics, 70: 11211128.

Dixit, A. K. and J. E. Stiglitz (1977). Monopolistic Competition and Optimum Product Diversity. American Economic Review, 67(3):
297308.

Fricke, L. and M. F. Quaas (2014). The Economic Causes of Regime Shifts in Marine Ecosystems. Manuscript, Department of Economics,
University of Kiel.

Grafton, R. Q., T. Kompas, and R. Hilborn (2007). Economics of Overexploitation Revisited. Science, 318: 1601.

Harley, S., R. Myers, and A. Dunn (2001). Is catch-per-unit-effort proportional to abundance? Can. J. Fish. Aquat. Sci., 58(9): 17601772.
Heal, G. and W. Schlenker (2008). Sustainable fisheries. Nature, 455: 10441045.

Kronbak, L. G. (2005). The Dynamics of an Open-access Fishery: Baltic Sea Cod. Marine Resource Economics, 19: 459479.

Nostbakken, L. and T. Bjorndal (2003). Supply Functions for North Sea Herring. Marine Resource Economics, 18(4): 345361.

Opsomer, J.-D. and J. Conrad (1994). An Open-Access Analysis of the Northern Anchovy Fishery. Journal of Environmental Economics
and Management, 27(1): 2137.

Quaas, M. F. and T. Requate (2013). Sushi or Fish Fingers? Seafood Diversity, Collapsing Fish Stocks and Multi-species Fishery
Management. Scandinavian Journal of Economics, 115(2): 381422.

Quaas, M. F., D. van Soest, and S. Baumg


artner (2013). Complementarity, impatience, and the resilience of natural-resource-dependent
economies. Journal of Environmental Economics and Management, 66(1): 1532.

Quinn, T. and R. Deriso (1999). Quantitative fishery dynamics. Oxford University Press, Oxford.

Ricard, D., C. Minto, O. P. Jensen, and J. K. Baum (2012). Examining the knowledge base and status of commercially exploited marine
species with the RAM Legacy Stock Assessment Database. Fish and Fisheries, 13(4): 380398.

Worm, B., E. Barbier, N. Beaumont, J. Duffy, C. Folke, B. Halpern, J. Jackson, H. Lotze, F. Micheli, S. Ralumbi, E. Sala, K. Selkoe,
J. Stachowicz, and R. Watson (2006). Impacts of biodiversity loss on ocean ecosystem services. Science, 314: 787790.
Martin Quaas
University of Kiel Resilience and sustainability of resource-economic systems

28/28

Das könnte Ihnen auch gefallen