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Muitidisciplinary data indicate that southwestern Amazonia was a savanna environment during Last Glacial times. Interpretation of
biogeographical and geological records, as compared with the present climatic conditions, permit us to postulate a dry season more
pronounced and prolonged than the present one; city northerly winds (vinodatog] with S O U ~ I~,misphcre trade winds) dominated during
the dry season, and the southerly cold air masses (locally termed 'friagem' or 'surazo') would have been more frequent and intensive than
today.
INTRODUCTION
The large Amazon rainforest, occupying 5,000,000 lan2,
has been a center of attention of the natural sciences in the
last centuries. Although the Amazon rainforest extends into
various countries, 80% of the total area is located in Brazil.
The Amazon ecosystem houses more living species than
any other single system on Earth. Today, for example, some
80,000 vascular plant species could be identified, and as
many as thirty million animal species may exist (Colinvaux,
1989).
In the last two decades, the most frequent explanation for
this biodiversity has been that of the forest refuges (Haffer,
1969, 1982; Prance, 1982). According to this theory, during
periods of glacial activity Amazonia had a much drier
climate, resulting in a rainforest reduction and in an increase
in the savanna areas. No refuges were available in the
southwestern Amazonia; however, some important
biogeographical changes occurred. Two important papers
give a review about the geomorphological and paleobotanical changes in Amazonia during the Quaternary, with
emphasis on the Last Glacial Maximum (van tier Hammen,
1991; Clapperton, 1993). In this work, we include both a
review and new data, explaining a probable circulation
model, specific for the southwestern Amazonia; most data
are included in Latmbesse's Doctoral Thesis (1992).
PRESENT CLIMATE IN AMAZONIA
In the Amazonia rainforest a humid tropical climate
prevails. Rainfall, averaging 2000 mm in the whole basin,
increases in the northwest to 5000-7000 mm (in Ecuador)
and to 10,000 ram in Choco (Colombia), outside the basin.
The area south from the centerline of the Solimoes--Amazon
River, known as southwestern Amazonia, shows a
pronounced dry season with high- and low-peaks of rainfall,
which does not hold true for the area north of the Solimoes
River. The shifting of the Intertropical Convergence Zone
(ITCZ) influences this region.
During summer, in the southern hemisphere, the ITCZ is
located from 10 to 15 S. In this period, southern Amazonia
163
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FIG. la. Normal rainfall in January (isoyets in ram). The shaded areas are above 500 meters above sea level (from Salati
FIG. lb. Normal rainfall in JaliLla/-y (isoyets in ram). The shaded areas are 500 meters above sea level (from Salati
et a L ,
et al.,
1978).
1978).
60"W
I
I
165
E_cua_dor
Geology
:"
Paleontology of Vertebrates
Rancy (1991) found large m a m m a l s of Lujanense
Mammal Age in southwestern Amazonia and proposed a
savanna environment during the Last Glaciation (Table 1).
Habitat
Diet
Eremotherium
Ocnopus
Glossotherium
Forestedge/savanna
Forestedge/savanna
Savanna
Grass/browse
Grass/browse
Ca'ass/b~wse
Lestodon
Scelidotherium
Mylodon
Megalonyx
Savanna
Savanna
Savanna
Savanna
Grass/browse
Grass/browse
C_a'ass/bfowse
Grass/browse
Edentata-Cingulata
Propraopus
Forestedge/savanna
Omnivo~
Dasypus
Euphractus
Pampatherium
Hoplophortts
Neuryurus
Panocthus
Glyptodon
Forest/savanna
Savanna
Savanna
Savanna
Savanna
Savanna
Savanna
Insectivore
Omnivore
Grass
Grass
Grass
Grass
Grass
Savanna
Savanna
Grass/low browse
Grass/lowbrowse
Savanna
Savanna
Grass/browse)fruit
Grass/browse/fruit
Forest/savanna
Browse/fruit
Savanna
Savanna
Forest/savanna
Grass/lowbrowse
Grass/lowbrowse
OnmivoreHrugivore
Forest
Carnivore
Edentata-Pilosa
Notoungulata
Toxodon
Mixotoxodon
Proboscidea
Cuvieronius
Haplomastodon
Perissodactyla
Tapirus
Attiodactyla
Vicugna
Palaeolama
Tayassu
Carnivora
Eira
166
65
. O*
CO
cation Hap
ECUADOR
,/
6~
B R A SjI
pu~
PERU
$
l0
Boundorl e s
"~
'
~.
Rivers
"~
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/'~
FIG. 3. Paleontological sites (black circles) with Upper Pleistocene mammals (Lujanean Mammal age) in southwestern Amazonia
(adapted from Rancy, 1991).
SouthwesternAmazoninin LastGlacialTimes
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REFERENCES:
~,C
SAND SEA
. . . . . . - - -
LOESS
AEOLIAN SILTY SANDS
--'~ ICE FIELD OF PATAGONIAN
ANDES
m
<,
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TRANSPORT OF SEDIMENT
BY MELT WATER
WINDS
168
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TRADE WIN ]S
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~AEOLIANSILTYSANDS
ICEFIELDS
t,0
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dP NORTHERLY WINDS
o-I-ES
ff
DOe
90
80
70"
60
50
FRIAGEHon SURAZO
40
30
SO*i
FIG, 5, Wind circulation model for southwestern Amazonia at Last Glacial times.
FINAL REMARKS
(1) Southwestern Amazonia supported a savanna
environment during the Last Glacial Maximum, with a dry
season more pronounced and prolonged than the present one.
(2) The dry northerly winds, originated in the anticyclonic
circulation of southern hemisphere trade winds, dominated
during the dry season.
(3) The 'friagem' or 'surazo' should have been more
frequent and intensive than today, having great importance
as a biD-regulation factor because of the temperature
decrease.
ACKNOWLEDGEMENTS
We wish to express our thanks to the Laboratorio de Paleontologia,
Universi__de,~__do Acre, for its professional aid; especially, to A. Rancy, J.
REFERENCES
Absy, M.L. (1993). Mndancas da ve~_L~c_~e clima da Amazoaia ~anm o
Quatemario. In: ~
E.G., dos Santos, G.M., LeaD, E.L. and
Oliveira, L. (eds), Bases Cientiflcas para Esn'osegias de Preservacao e
Desenvolmimenm da Anmzoni~" Fo~os e Perspectivas, M M A / U N A MAZ. Vol. 11, p. 310. Manaus.
Absy, M.L and van der I-lammen, T. (1976). Some pa]eoecological data
from Rondonia, southern part of the Amazon Basin. Acre Amaz6nica.
6(3), 293-299.
Absy, M.L., Cleef, A., Foornier, M.. Ma~dn, L.. Servant M., Siffeddine, A..
Ferreira Da Silva, M.) Soubies. F., Suguio, K., Turq, B. and van Der
H ~ , T. (1991). Mise en evideace de quM_~-phases d'ouvemu~ de
la foret dense clans le sudest de l'Amazonie au cours des 00.000
demieres ana~. Premiere compareimn avec d'autres regions tropic,ales.
CR. Acad. Sci. Paris, t. 312, Set. If, 673.-678.
Brinkman, W.LF.. Weinman, J.A. and Goes Ribeiro. M.N. (1971). Air
temperatures in cenuai Anmzoaia. Acre Anm~n/ca, 2, 5t-56.
Cant~, M. (1992). Provincia de C6rdobL In: Iriondo, M. (ed.), El Holoceno
en/a Argembm, Vol. 1, pp. 1-I6. CADINQUA.
Clapporton. Ch.M. (1986). Glacial leomorphology. Quaternary glacial
sequence and l~aeoclimatic iafemaces in the Ecuadm'ian Andes. In:
Garcfiner. V. (ed.). International G-eomorphology, Part I], pp. 843-870.
Wiley, London.
Clapperton, Ch.M. (1993), Nature of environmental changes in South
169