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I.
Pre- Cambrian
Defining Characteristics:
o formation of Earths crust and main bombardment
Secondary Characteristics:
o continuing erosion and plate tectonics have destroyed Hadean rocks
The name Hadean Eon comes from Hades, the underworld of Greek
mythology. It refers to the hellish conditions of the Earth during the earliest part of
its history, when much of the Earths surface remained molten. The Hadean Eon of
geologic time began with the birth of the solar system, including our planet, Earth,
and ended with the formation of the oldest rocks that are still preserved on the
surface of Earth.
The Hadean is the first period in Earth history, but one for which we have
little record. The Earth began to form about 4.6 billion years ago through the
condensation of material around the sun. As this material collected, further cosmic
material was drawn to it by gravity from all directions, increasing the size of the
Earth. This process created an enormous amount of heat, which melted these
materials and eventually allowed them to separate into different layers. As the
Earth cooled, it acquired the structure we know todayan iron core, silicate mantle,
and thin outer crust.
Formation of the Earth
The Earth initially formed from the same disk of dust and gas from which the
sun itself coalesced. As the mass of the Earth increased, so did the gravitational
force it exerted, and it was bombarded by objects from space ranging in size from
dust particles to small planets (planetismals). The accretion of this material
increased the size of Earth. The impacts of large bodies and the decay of
radioactive elements generated heat that melted the materials of the young Earth,
creating the hellish conditions for which the Hadean Eon was named.
Although heat was generated through decay of radioactive elements and
continued frequent bombardment by asteroids, Earth lost heat to space and slowly
cooled. The different temperatures at which molten iron and silicate minerals
solidify indicate that as it cooled the Earth eventually segregated into an iron core
and silicate mantle. These two features are still part of the fundamental structure of
the Earth today.
Earths Atmosphere
The composition of Earths earliest atmosphere remains unclear and is a
subject of active investigation by geologists. Some geologists believe that the
earliest atmosphere included large amounts of nitrogen. Others believe that it was
composed largely of carbon dioxide and water vapor, along with some other
volcanic gases. These gases would have come from steaming vents and volcanoes
that were common across the Earths surface. Early in the Hadean, Earths
atmosphere was probably too hot to allow liquid water to condense, but fragments
contained within younger rocks suggest that sedimentary rocks formed later in the
Hadean, which implies the existence of surface water.
Earths Surface
Although continents, oceans and an atmosphere must have existed before
the Acasta Gneiss formed, the size of the continents is unclear. While Earth was still
very hot, mantle convection must have been vigorous. It is not known if plate
tectonics operated on the early Earth. One suggestion is that the earliest surface
crust was thin and unstable, and made of minerals with an extremely high content
of iron and magnesium, which are very dense. This early crust may have been
Defining Characteristics:
o first life appears
Secondary Characteristics:
o plate tectonics established
o oxygen-poor atmosphere
The term Archean means ancient and was originally used to refer to the oldest
known rocks. Rocks of Archean age contain the earliest fossils of life on Earth. Because
these rocks were formed very long agobetween 2.5 to 4.0 billion years agomost
have long since been covered by younger sediments, eroded, or subducted into the
Earth's mantle. Nevertheless, some Archean strata survived in the central parts of
continents. These Archean shields lie in the heart of Canada, Australia, Africa, India,
and Siberia.
By the start of the Archean Eon, the Earth's crust had cooled. The atmosphere was
composed of volcanic gases, including nitrogen, hydrogen, carbon, and possibly low
levels of oxygen. Water vapor was abundant and the first oceans had formed. A
complex set of chemical reactions in these early oceans transformed carbon-containing
molecules into simple, single-celled life forms. Where these chemical reactions
occurred is unclear--hydrothermal vents are one possibility. All life today is descended
from these simple organisms. By the end of the Archean the first photosynthesizing
organisms had evolved and begun to produce oxygen, which was released into the
oceans and atmosphere. This process would dramatically change life on Earth during
the following Proterozoic Eon.
Early Continents and Oceans
Analyses of the approximately 4 billion-year-old Acasta Gneiss suggests that the
first continents and oceans developed before the Archean. The Archean, however, is
the period during which the present continents took shape. Most present continents
have shields at their cores that formed between 3 to 2.5 billion years ago during the
early Archean. Evidence of ancient oceanic crust is often found in today's greenstone
belts. Continental crust gradually grew from selective melting of dark-colored basaltic
igneous rocks within the oceanic crust. Through time, these melts became increasingly
rich in silica, as geologic processes melted more of the lower-temperature, lighterdensity minerals. Much as a helium balloon rises through air, these silica-rich melts
rose from deep within Earth and formed granitic plutons (intrusive bodies) nearer the
surface. Early in the Archean the granitic crust of continents had begun to form from
basaltic crust of the ocean floor. Continental landmasses began forming about 3.7
billion years ago from the horizontal accretion of smaller micro-continents. The Kenoran
orogeny was one such event, which formed what is now the Great Lakes region of
North America during the Late Archean.
The First Life on Earth
Earth was able to support life only after the planet had cooled enough for a rocky
crust to solidify. Once that happened, water vapor from volcanoes condensed in the
atmosphere, fell as rain, and collected on the Earths surface. Besides water vapor,
volcanoes also produced gases rich in the basic ingredients of life: carbon, hydrogen,
oxygen, and nitrogen. Toxic gases such as ammonia and methane were common. At
this point, Earth's early atmosphere consisted entirely of these volcanic gases, and
there was no free oxygen. In the primordial soup of the early seas, organic molecules
concentrated, formed more complex molecules, and became simple cells.
The transition from complex organic molecules to living cells could have occurred in
several environments. Small, warm ponds are one possibility, but recent work has
suggested that deep-sea hydrothermal vents, such as those found along mid-ocean
spreading centers today, may have been the cradle of Earth's life. These environments
contain the chemicals and the source of energy needed to synthesize more complex
organic structures. Although scientists have not succeeded in creating life from organic
molecules in the laboratory, they have reproduced many of the intermediate steps.
Changes in the Atmosphere
During the Archean and early Proterozoic Eons, the deep oceans had large volumes
of dissolved iron. This combined with oxygen, possibly produced by photosynthetic
microorganisms, to produce iron oxides along the continental margins. This rust was
concentrated in sedimentary deposits known as Stromatolites and other microfossils
provide important evidence for the transformation of the oxygen-poor atmosphere of
the Archean, into one that was oxygen-rich in the Proterozoic, like the atmosphere
today. Some microorganisms that build modern stromatolites are capable
of photosynthesis and release free oxygen into the ocean. Chemical traces of
microorganisms, known as biomarkers, show that photosynthetic organisms had
evolved by 2.7 billion years ago, but they were probably not present during the early
Archean.
Defining Characteristics:
o first multicellular animals at end of interval
o Map of the Proterozoic World
Secondary Characteristics:
o 4 major mountain-building episodes
o oldest known glaciation
The term Proterozoic comes from the Greek words proteros (meaning first) and
zoon (meaning life). Although early fossils are now thought to occur in older, Archean
rocks, for many years the oldest evidence of life was known from Proterozoic strata.
The Proterozoic represents the last stage of what was once called the Precambrianthe
3.7-billion-year-long period before the Paleozoic Era. Rocks of this age are known from
Australia, Canada, and China.
During the Proterozoic, larger continental landmasses continued to form by the
accretion of smaller ones, often leading to extensive mountain-building. As the
continents began to erode, sediments were washed into the oceans, producing shallowwater marine environments where life could flourish and diversify. Many of these life
forms developed the ability to photosynthesize. As a byproduct of photosynthesis, they
created oxygen, and over billions of years this oxygen transformed the Earths
atmosphere. Because oxygen was toxic to some early life forms, many went extinct.
Others thrived and evolved into the first multicellular organisms, which are preserved
as the Ediacaran fauna. Both the beginning and end of the Proterozoic were marked by
widespread glaciation.
Earths Crust as a Platform for Prokaryotic Life
For much of the history of living things, organisms were exclusively primitive, singlecelled forms such as bacteria and cyanobacteria. Then as now, these simple
prokaryotic cells lacked a nucleus and were relatively small, but they already contained
the same basic hereditary materials found in all life todayDNA and RNA. Many of
these early life forms relied on chemosynthesis for their energy, deriving it from the
chemicals around them instead of from sunlight or other organisms. Some inhabited
environments that seem hostile, such as deep-sea hydrothermal vents.
Eukaryotes and the First Multicellular Life Forms
A fundamental biological change occurred with the appearance of eukaryotes.
Eukaryotes differ from prokaryotes in that their cells contain membranous sacs called
organelles, including mitochondria, chloroplasts, and the nucleus. Many scientists think
these organelles are descended from formerly free-living prokaryotic organisms. Thus,
many important functions of eukaryotic cells, such as photosynthesis, and respiration
(the process by which organisms use oxygen to metabolize organic compounds to
produce energy, giving off carbon dioxide) were acquired through a symbiosis of
independent forms of life. Eukaryotes flourished as the environment became richer in
oxygen, perhaps in part because of their more complex intracellular function.
An important evolutionary innovation was multicellularity. The oldest known
possible multicellular eukaryote is Grypania spiralis, a coiled, ribbon-like fossil two
millimeters wide and over ten centimeters long. It looks very much like a coiled
multicellular alga and has been described from banded iron formations in Michigan 2.1
billion years old. Grypania may not be a eukaryote, but another, unrelated colonial
eukaryote, Horodyskia, is known from sedimentary rocks dated at 1.5 billion years in
western North America and from rocks more than 1 billion years old in Western
Australia.
Changes in the Atmosphere
Earths early atmosphere contained only small amounts of free oxygen, probably
produced entirely by the reaction of sunlight with water vapor from volcanoes. The
oxygen-rich atmosphere that evolved later, and upon which oxygen-breathing life now
depends, was a result of the origin of photosynthesis. During the Precambrian, vast
numbers of single-celled algae and cyanobacteria living in the seas eventually released
enough oxygen to transform the environment. The oldest evidence of cyanobacteria
dates to 2.7 billion years ago, although oxygen did not begin to build up in the
environment until about 2.3 billion years ago. During the transition from oxygen-poor to
oxygen-rich atmosphere, the first banded iron formations may have formed.
Thus, the history of Earths early crust also tells the story of its early atmosphere.
Banded iron formations were precipitated from about 3.1 to about 2 billion years ago
most (92%) during the Proterozoic between 2.5 and 2 billion years ago. Until all the
available iron had been deposited in banded iron formations, oxygen could not build up
in the atmosphere. Red beds appeared only after free oxygen was released into the
atmosphere, beginning about 2.0 to 1.8 billion years ago. They are still being formed
today.
Proterozoic Mountains and Glaciers
In the Proterozoic Eon, four major mountain-building episodes occurred, each of
which was followed by an interval of continental erosion. Mountain-building was caused
by converging plates, just as occurs in present-day plate tectonics. It was accompanied
by intrusions of molten granite that welded an additional belt of younger, igneous rock
around the edges of the original microcontinents. In North America, the Proterozoic
episodes of mountain-building greatly expanded the size of the continent.
Widespread continental glaciations evidently occurred at least twice in the
Proterozoic, once near its beginning and again near its end. Several of these glaciations
extended almost to the equator, much farther south than any recent cooling events.
This unusual situation has led a few geologists to propose that the Earth was almost
entirely covered by glaciers for perhaps several million years during the Proterozoic.
During this Snowball Earth phase, life would have been relegated to hydrothermal
vents and other such refuges until the build-up of carbon dioxide released from
volcanoes warmed Earth from its deep-freeze. The final Marinoan Ice Age marked the
transition to the Cambrian, the first period of the Paleozoic Era.
II.
Phanerozoic
a. Paleozoic
i. Cambrian (542- 488.3 mya)
Defining
Characteristics:
o Cambrian
Explosion
o skeletonized
animals
o early animal
diversification
Secondary
Characteristics:
o diversification
of trilobites
o Burgess Shale
fauna
o warming
climates
o Plate tectonics
The fossil record of Proterozoic life is sparse because it was largely soft
bodied or microbial. The advent of the Paleozoic Era brought an incredible
diversification of multicellular animals having many different body plans,
including most of the major groups alive today, such as mollusks, trilobites,
other arthropods, brachiopods, echinoderms, corals, sponges, and chordates
(our ancestors). Most significantly, the first animals having hard outer
skeletons evolved; as a result, many of these shells became fossilized. Most of
the Early Cambrian animals having exoskeletons were small, measuring one to
five millimeters, and are often called the "Small Shelly Fauna". Another unique
feature of the Cambrian fauna is the sponge-like archaeocyaths that formed
reefs during Early Cambrian time. This sudden appearance of many different
kinds of animals having skeletons (shells) in the fossil record is called the
Cambrian Explosion. The Cambrian Fauna is as different from Vendian life as it
is from the Paleozoic Fauna that occurs above Cambrian-age rocks.
The Burgess Shale Fauna
and biology of the animals better. Some of the soft body plans of Cambrian
animals did not survive beyond that period. To our eyes, some of these body
plans look bizarre and it has taken a long time for paleontologists to
understand them. For example, the first reconstructions of the onycophoran
Hallucigenia were upside-down. The sclerite-bearing Wiwaxia, the arthropodlike predator Anomalocaris, and the five-eyed Opabinia are all strikingly
different from modern animals.
The "Age of Trilobites" and the Cambrian Fauna
The most abundant and diverse animals of Cambrian time were the
trilobites. Trilobites had long antennae, compound eyes, many jointed legs, and
a hard exoskeleton like many of their modern arthropod relatives, such as
lobsters, crabs, and insects. The Cambrian is sometimes called the "Age of
Trilobites" because of their explosive diversification into all marine
environments worldwide. In size, they ranged from a few millimeters (1 mm =
0.25 inches) to 45 centimeters (18 inches).
The Cambrian began after the end of the Marinoan ice age, one of
several instances in the past when ice was widespread on Earth's surface. This
marked a shift from late Proterozoic icehouse conditions, during which much of
the world was cold and ice covered. The Cambrian was quite the opposite, with
global warming leading to greenhouse conditions. This climate change was
probably linked to major changes in the positions of the continents.
Defining
Characteristics:
o diversification
of marine
invertebrate
Paleozoic
Fauna
o endOrdovician
extinction
Secondary
Characteristics:
o deep-water
faunas,
especially
graptolites
o appearance of
conodonts
o development
of diverse
carbonate
platforms
o first major
Paleozoic ice
age
Charles Lapworth in 1879 named the Ordovician for a set of rock strata
that were particularly well exposed in Wales. He took the name from Ordovices,
the Latin name of a Celtic tribe in the Welsh region during Roman times. By
creating the Ordovician, Lapworth helped settled a dispute, because previously
Adam Sedgwick had assigned these rock layers to his Cambrian, whereas
Roderick Murchison called them Silurian. Lapworth recognized them as distinct
from the (older) Cambrian rocks below and the (younger) Silurian rocks above.
This processdetermining the age of a rock by the fossils contained within it
is called biostratigraphy.
Ordovician rock layers were deposited between 443 and 490 million
years ago. During this time period, marine life saw a major shift from the
Cambrian Fauna, which appeared during the Cambrian Explosion, to the
Paleozoic Fauna, which would dominate the marine world for the next 200
million years. Important new animal forms, such as graptolites and conodonts,
first appeared during the Ordovician, along with the development of complex
shallow-water reef ecosystems. The first land organisms, lichens and
bryophytes, appeared. The end of the period was marked by a major ice age
and the second-largest mass extinction of the Paleozoic Era.
Reef Ecosystems and the Paleozoic Fauna
Life in the Ordovician was quite different from that in the Cambrian.
Although many animal groups survived from the Cambrian, some forms
disappeared and other forms appeared for the first time. These changes
marked the Ordovician as the first example of the Paleozoic Fauna. During this
time, bryozoans, mollusks, echinoderms (particularly the fan-like crinoids),
brachiopods, and trilobites diversified; these groups would dominate marine
faunas throughout the Paleozoic. Many of the unusual animal forms seen in the
Burgess Shale had gone extinct, but graptolites, conodonts, nautiloids, and
jawless fishes had their first known appearances.
These new animals radiated alongside the rugose and tabulate corals
that replaced the Cambrian archaeocyaths. In fact, entirely new types of
ecosystems evolved in the Ordovician, associated with the development of
many types of carbonate (limestone) marine shelves (or platforms). These
warm, shallow-water formations were flooded with sunlight, allowing the
establishment of stable reef ecosystems. These included the first coral/algal
reefs, shallow-water assemblages of mollusks, brachiopods, echinoderms and
trilobites, and more-diverse underwater slope and deeper-water faunas. Many
of these faunas would become even more important in later Paleozoic periods.
New Animals as Biostratigraphic Tools
During the Ordovician, tectonic activity occurred in New England and
the British Isles as North America moved toward the equator and began to
collide with Europe. This led to mountain-building along these major plate
margins. As plates continued to shift in position, diverse faunas were brought
into contact with one another. Plate movement may also have contributed to
the formation of the first Paleozoic ice age, as the southern continent of
Gondwana moved over the South Pole. Glaciers began to form in what is now
the Sahara region of Africa. This climate change was possibly associated with
the mass extinction at the end of the Ordovician. These tectonic clues allow
scientists to make sense out of geologic patterns in the Earth, and to map
present-day continents and islands onto these old deposits.
Defining Characteristics:
o distinct estuarine, freshwater, and terrestrial ecosystems develop
the start of significant life on dry land
Secondary Characteristics:
o first vascular plant fossils
o wide and rapid spread of jawless fishes
o first extensive coral reefs
The Silurian physical world was vastly different in many ways from the
world of today. For example, the equator passed through what is now North
America, and a nearly continuous inland sea extended from New York to
Nevada. Life was also quite different. Following the end-Ordovician extinction
event, Silurian marine faunas rebounded in the warm, shallow continental seas.
Large coral reefs made their first appearance during this time. Although marine
life was diverse and abundant, there were only a few types of small plants and
terrestrial arthropods on land in the Silurian. However, these represent the first
true terrestrial ecosystems on Earth and include the first fossil records of
vascular plants. These terrestrial ecosystems became more complex, especially
during the Late Silurian, as upright plants evolved and populated the wet parts
of lowland landscapes.
Silurian Marine Life
Marine life thrived and diversified during the Silurian; many forms were
similar to those of the earlier Ordovician Period. Although the later Devonian
Period is often called the Age of Fishes, the Silurian also was an important
time in the evolution of this group. In particular, it was marked by the wide and
rapid spread of jawless fishes (agnathans). Only two of their jawless
descendantslampreys and hagfishessurvive today, but many different kinds
populated the Silurian seas.
Late Silurian. These plants had evolved an internal system of tubular cells
through which water traveled up the body of the plant, allowing them to
maintain cell functions higher above the land surface than would otherwise
have been possible. Simple, small plants such as Cooksonia lacked leaves,
roots, seeds and the capability to grow in diameter, but they were the
dominant early terrestrial plants.
Defining Characteristics:
o Age of Fishes
o diverse land invertebrates and first land vertebrates
o diversification of vascular plants
Secondary Characteristics:
o Late Devonian extinction
o first true trees
The recognition of the Devonian Period was the result of great debate
on the part of many nineteenth-century European geologists. The debate
centered on whether the Old Red Sandstonethe rock layers above the
Silurian (and therefore younger)actually represented a distinct system or
merely a later stage of the Silurian. In 1839, Roderick Murchison and Adam
Sedgwick collaborated to name the Devonian, which they based on rock
exposures in Devonshire, England. Devonian-age rocks are also common in
Scotland, central Pennsylvania, western New York, and Greenland, but they
have been found on all continents.
During the Devonian, most of Earths landmasses formed two
neighboring supercontinents, Gondwana and Euramerica. The rest of the
Earths surface was covered by a vast ocean. The Devonian world was
populated by now-extinct, very primitive plants and animals, so it looked much
different from our world today. In the marine realm, many members of the
Paleozoic Fauna continued to diversify. On land, vascular plants and arthropods
formed diverse terrestrial ecosystems, while the earliest tetrapods appeared in
shallow waters nearby.
Life in the Devonian Seas
Fishes. Although they first appeared in the Silurian, spiny acanthodians and
armored placoderms reached their peak diversity during the Devonian and
began to dwindle in numbers later in the Paleozoic. Some, like the
giant Dunkleosteus, reached nearly three meters in length and were
menacing marine predators. Placoderms had no teeth, instead relying on selfsharpening bony plates in their jaws that performed the function of teeth.
Chondrichthyan (cartilaginous) fishes, such as sharks, were Ordovician
survivors that thrived during the Devonian. A few shark scales have been
dated to the Late Ordovician, but the earliest shark teeth are Early Devonian,
and the first relatively modern-looking sharks had evolved by the Middle
Devonian. The first truebony fishes (osteichthyans) appear as well, as
both lobe-finned (sarcopterygians) and ray-finned (actinopterygians) forms.
The lobe-finned fish are of particular interest because they gave rise to the
first land vertebrates, around 360 million years ago
Terrestrial Habitats Conquered
The Devonian was the time during which life truly conquered land. At
the start of the period, terrestrial life was restricted to the narrow margin along
the waters edge. The first land plants had already evolved but still required a
moist environment to reproduce. By the middle of the Devonian, the first
shrub- and tree-like plants had appeared, some of which reached heights of
five meters or more. By the Late Devonian, the first true trees and forests had
evolved. This change is marked by the arrival of the genus Archaeopteris,
whose species lived on nearly all Devonian landmasses. Like modern forms,
these trees had extensive root systems, broad leaves, and specialized vascular
systems that facilitated the flow of water and nutrients against the pull of
gravity. All of these features helped plants colonize drier areas, which
expanded lifes terrestrial habitat. Broad leaves provided shade and moderated
temperature and humidity levels, whereas advanced root systems assisted
production of the first soils (pedogenesis) by encouraging weathering of the
land. These early forest habitats had important ecological effects on all other
ecosystems. Gymnosperms appeared at the very end of the Devonian. These
were the first plants to bear seeds, which allowed them to move farther into
drier environments.
The Devonian Extinction
Toward the end of the Devonian, nearly 70% of all invertebrate species
vanished during the Late Devonian extinction. Marine species (especially
tropical ones) suffered the most extinctions, followed by freshwater species,
whereas terrestrial species were hardly affected. In particular, tabulate
coral/stromatoporoid reefs vanished entirely, leading to a great decline in
worldwide coral reef-building until the Triassic Period. Many species of
brachiopods, trilobites, and early fishes went extinct, as did the planktonic
graptolites. These extinctions were not the result of a single major extinction
event, but rather smaller extinction events that occurred over a period of more
than 20 million years.
Climate and Tectonics during the Devonian
once called the Old Red Continent because of the color of Devonian-age
rocks in Europe and North America, began to drift northward. Gondwana also
drifted north. The formation of Euramerica and the closure of the oceans
between Euramerica and Gondwana initiated the formation of Pangea, which
continued into the early Mesozoic.
v. Carboniferous
Defining
Characteristics:
o first truly
terrestrial
tetrapods
Secondary
Characteristics:
o formation
of
supercon
tinents
In 1869, American geologist Alexander Winchell coined the
name Mississippian in 1869 for rock outcrops along the drainage basin of
the Mississippi River. He distinguished these limestone-rich Lower
Carboniferous rock layers from the coal-bearing beds of the Upper
Carboniferous (or Pennsylvanian). Because these two sets of rocks are
easily distinguished in North America, the terms Mississippian Epoch and
Pennsylvanian Epoch are commonly used by American geologists and
paleontologists. This distinction was less clearly marked in Europe, and
the names Lower and Upper Carboniferious were widely used. The
European subdivision of the Carboniferous has now been formally replaced
by Mississippian and Pennsylvanian by the International Commission on
Stratigraphy.
The uplifting of the continents in the Mississippian reduced the
area covered by inland seas and resulted in less available space for
marine life. As Gondwana continued to move across the South Pole during
the Late Mississippian, temperatures cooled and ice began to form in
great sheets. As more water was frozen into these ice sheets, sea levels
dropped and terrestrial habitats in turn increased. The globally warm
climate became cooler later in the Mississippian.
Defining Characteristics:
o Coal Age, very well known for plant fossils that make up
the worlds major coal seams
Secondary Characteristics:
o Age of Amphibians, including the ancestors of modern
terrestrial vertebrates
o diversification of terrestrial and aquatic insects.
o evolution of amniotic egg in reptiles
and tropical for much of the period. Mid-latitudes were seasonally dry, and
some of the changes in climate and vegetation recorded by fossils and
rocks in North America and Europe probably reflect the movement of
these areas from the warm, wet tropical belt into the seasonally dry
subtropics.
Defining Characteristics:
o the end of the Permian and the Paleozoic is based on the most
extensive mass extinction in the past 600 million years, leading to
the advent of dinosaurs and modern terrestrial and marine biotas
Secondary Characteristics:
o extensive glaciation at the start gives way to general global
warming
o great diversity of amphibians, diapsids, and synapsids
o the greatest high-level taxonomic diversity of insects of all time
o seed plants become more dominant
The Permian Period marked the end of the Paleozoic Era. The land was
inhabited by a wide diversity of terrestrial insects and vertebrates. Insects
included the earliest representatives of the two most dominant groups of
insectsthe hemipteroids, represented by extant cicadas and lice, and the
highly diverse holometabola, represented by beetles, flies, wasps, and moths.
Also important were the first vertebrate herbivores. Marine faunas were still
dominantly composed of groups belonging to the Paleozoic Fauna. The Earths
continents were coalescing into a single supercontinent, Pangea. The rise of
conifers as dominant elements of tropical vegetation reflects increased
seasonality of rainfall in tropical areas. The Permian is most remarkable for its
conclusion, however: the end-Permian extinction was the largest of the past
600 million years, and it heralded the end of the dominance of the Paleozoic
Fauna. In its aftermath, archosaurs rose to dominate the land, and more
modern groups filled the marine realm.
Terrestrial Animal Life and Evolution of Herbivores
The most important terrestrial herbivores in the Early Permian were the
insects. They represented a mixture of holdovers from the earlier
Pennsylvanian, such as the dragonfly-like palaeodictyopterans and various
forms related to modern grasshoppers and cockroaches, as well as newly
evolved forms such as beetles and scorpionflies that are the earliest known
members of insect groups that undergo true metamorphosis. (Insects that
metamorphose develop from egg to larva to pupa to adult, and generally make
their living in a very different way in the larval and adult phases.) The advent
of insects that could metamorphose had a profound effect on the ecology and
evolution of insects, because immature life stages could use different resources
than adults of the same species. Evidence for these new herbivores is seen in
Permian plant fossils with new types of damage to leaves and seeds.
Sedimentary rocks of Permian age also reveal fossil burrows made by the first
insects to colonize quiet-water aquatic ecosystems.
have been more seasonal. Climatic zonation was pronounced, however, and
floras from different climatic belts were markedly distinct.
The end of the Permian was marked by the greatest mass extinction of
the last 600 million years of Earth history, during which perhaps 90% of marine
animal species disappeared. Major groups such as trilobites, fusulinid
foraminiferans, rugose and tabulate corals, acanthodian and placoderm fishes,
and blastoid echinoderms vanished entirely, Although they did survive,
brachiopods, bryozoans, ammonoids, sharks, bony fishes, crinoids, eurypterids,
ostracods, and many echinoderms lost the majority of their species. Finally,
insects suffered their greatest mass extinction in Earth history.
The southern ice cap melted off permanently during the earliest
Permian. The supercontinent of Pangea was nearly fully assembled by the end
of the period, with only North and South China and Cimmeria remaining
unattached to the east. An open gulf called the Paleo-Tethys Ocean formed
along Pangeas eastern margin. The total length of shorelines continued to be
reduced as the major landmasses coalesced, and shallow seas decreased as
sea level dropped. The salty, inland Zechstein Sea occupied much of Europe.
b. Mesozoic
Defining Characteristics:
o Early Triassic: the start of lifes remarkable recovery after the endPermian extinction
o the origin and rise of dinosaurs and the first mammals in the
Middle to Late Triassic
Secondary Characteristics:
o the emergence of giant marine reptiles (ichthyosaurs) and the
rapid diversification of cephalopod ammonites in the oceans
o the first truly modern coral reefs in the Middle Triassic
At the start of the Triassic, 252 million years ago, the equatorial
supercontinent of Pangea completed its formation as the final continents
collided with the mainland. Polar ice caps were absent and sea level remained
fairly low and stable. The climate was generally warm and arid, but moist river
and lake environments sustained gymnosperm forests and large amphibians.
Nevertheless, life on Earth was sparse. Only a few types of animals and plants
were left after the great mass extinction at the end of the Permian, and sea life
in particular was severely depleted. The repopulation of Pangea had meager
beginnings, and the rediversification of life took 46 million years. The Early
Triassic is characterized by low diversity in both marine and terrestrial habitats,
but by the Middle or Late Triassic diversity had rebounded in most
environments.
Extinction and Recovery
The first third of the Triassic was a recovery period from the endPermian extinction. This greatest extinction in Earths history, eliminating
approximately 70% of the species of land vertebrates and 90% of marine
animal species, sharply reduced the number of different ways in which plants
and animals made a living. During the Triassic life re-evolved many strategies
for living, and added new ones not seen during the Paleozoic. Newly evolved
scleractinian corals formed small reefs, beginning the recovery of reef
ecosystems. Mollusks such as ammonoids (relatives of the modern chambered
nautilus) were severely reduced in diversity by the extinction but evolved
rapidly afterward to become more diverse than ever before and to dominate
the open-ocean marine invertebrate world.
New types of animals and plants continued to evolve throughout the
Triassic. The first mammals and dinosaurs originated almost simultaneously, in
the late Middle or early Late Triassic. The oldest known fossil of an amniote egg
is from the Early Triassic. In the seas, ichthyosaurs (dolphin-shaped reptiles),
nothosaurs, and placodonts (mollusk-eating reptiles) appeared and thrived.
Some ichthyosaurs reached lengths of 23 meters (75 feet). Turtles,
crocodyliforms, and pterosaurs all made their debuts, along with frogs and
sphenodontians. In fact, by the end of the Triassic, many of the animal groups
we see today had made their first appearances on Earth.
Origin of Mammals
evolved into the very first mammals. Although many different types of
mammals evolved later in the Mesozoic, for the most part they remained small,
probably because of the dinosaurs' success. However, later in the Mesozoic
some mammals achieved the size of a 30 lb. (13 kg) dog despite the dominant
success of the dinosaurs. Many types of mammals appeared in the Mesozoic,
and evidence from Early Cretaceous outcrops (130 million-years ago) in China
shows that at least one carnivorous mammal, Repenomamus robustus, feasted
on small dinosaurs.
Origin of the Dinosaurs
The earliest dinosaurs are known from Argentina and include predatory
theropods (Herrerasaurus, Eoraptor) and herbivorous ornithischians
(Pisanosaurus). By the end of the Triassic, dinosaurs were widespread and
dominated most terrestrial ecosystems. The most abundant were small
theropods such as Coelophysis (from North America), and large, herbivorous
prosauropods such as Plateosaurus (from Europe). Rare evidence also exists of
other groups, including the first sauropods and armored dinosaurs. During the
following Jurassic Period, dinosaurs and other archosaurs would become even
more diverse and spectacular, evolving into gigantic sauropods, large
theropods, and birds. Their dominance would continue until the end of the
Mesozoic.
Climate and Plate Tectonics
Early Triassic climate was quite similar to that at the end of the
Permian. Much of Pangea was warm and dry, and the interior of this
supercontinent was particularly arid. These environments were often
dominated by conifers and other gymnosperms. However, the Triassic also saw
an increase in seasonality, as well as prominent monsoon weather cycles.
Provincial biotas developed as well. In the north (Laurasia), these ecosystems
included ginkgoes, bennettitalians, cycads, and tree ferns. In contrast,
southern (Gondwanan) environments were dominated by seed ferns, most
prominently one called Dicroidium.
Defining Characteristics:
o The Age of Dinosaurs: dinosaurs become very diverse, evolving
into stegosaurs, theropods, and huge sauropods
Secondary Characteristics:
o origin of birds
o origin of the parasitic feeding guilds in terrestrial ecosystems
possibly replacing predators as top insect carnivores
o Pangea continues breaking apart, and is better separated toward
the end of the Jurassic, with a lush, warm tropical climate
The name Jurassic comes from the Jura Mountains, an extension of the
Swiss Alps into eastern France, where rocks of this age were first studied. They
were first identified as the Jura Kalkstein (Jura Limestone) by Alexander von
Pangea was centered on the equator for most of the Jurassic Period,
and Earths climate was decidedly tropical. During the Early Jurassic, some
regions of the world were still arid, but by the Late Jurassic much of the planet
was lush. Great rivers covered North America, and the land was green with
ferns, seed ferns, cycads, ginkgos, and conifers. It was the ideal environment
for the largest animals ever to walk on Earthgiant sauropods such as
Diplodocus, as well as many other types of large dinosaurs. Like today, much of
the land was covered with trees, although flowering plants had not yet evolved.
And, like today, there were many understory plants such as ferns, cycads, and
horsetails, but there was no grass for smaller animals to hide in. Because of the
dinosaurs dominance, mammals were still no larger than rats but had begun
to diversify by the Early Jurassic. The Jurassic seas were filled with many types
of sharks, bony fishes, marine crocodiles, and other marine reptiles of all sizes.
Cephalopods such as ammonites propelled themselves through the oceans.
Birds evolved and began to diversify during the Late Jurassic, but the most
common flying vertebrates were the reptilian pterosaurs. By the end of the
Jurassic, major parts of Europe and North America had become flooded as sea
level rose, and Pangea continued to break apart. As North America and Eurasia
drifted away from Africa and South America, the Atlantic Ocean was born,
creating a barrier for land travel between these regions.
Jurassic Life
Rising sea levels flooded many of the continental interiors during the
Jurassic, creating warm, shallow-water environments where marine animals
and plants could thrive. These regions saw an increase in diversity of
microscopic phytoplankton such as coccolithophores, dinoflagellates, and
foraminiferans. Reef ecosystems continued to flourish, thanks to many species
of corals and sponges. Among these sessile (stationary) organisms lived
gastropods (snails), along with the now-rarer brachiopods and crinoids. In the
waters above swam predatory cephalopods such as ammonoids and
belemnites. Sharks and bony fishes remained common and shared the seas
with ichthyosaurs, plesiosaurs, and other marine reptiles. The first true marine
crocodiles had appeared, alongside the first true teleost fishes (which today are
the most diverse vertebrates on Earth).
By the Late Jurassic nearly every major kind of dinosaur had appeared,
and they are found on every continent. Although prosauropods had gone
extinct, the world was now home to armored stegosaurs (Stegosaurus) and
ankylosaurs, herbivorous ornithopods of all sizes (Camptosaurus), and large
(Allosaurus) and small (Ornitholestes) predatory theropods. Many of these
dinosaurs were collected from the Morrison Formation, a thick formation of
mud, silt and sand that was deposited in western North America about 150
million years ago by a large braided river system running across much of the
central part of the continent. These Late Jurassic dinosaurs had close relatives
in Africa and Europe, indicating that these areas were still connected. In the
Cretaceous, however, these connections would finally be severed.
Flight and the Origin of Birds
One of the most important paleontological finds came in 1861, with the
discovery of Archaeopteryx in Late Jurassic (around 146 million years ago)
limestones near Solnhofen, Germany. Although the skeleton of Archaeopteryx
was nearly identical to that of the small theropod dinosaur Compsognathus,
this fossil also bore the unmistakable imprints of feathers. For over 100 years,
Archaeopteryx was the strongest evidence that birds had evolved from
theropod dinosaurs, and it therefore deserved a place on the dinosaur family
tree. Since then, many other feathered dinosaurs have been found in China,
further supporting this hypothesis, and future discoveries will help us
understand exactly how flight evolved in this unique group of theropods.
Pangea, which had begun forming in the Devonian (400 million years
ago) and lasted through the Triassic, finally began to split apart in the Late
Triassic. By the Middle and Late Jurassic, enough plate movement had occurred
to separate South America from southern Africa. Laurasia (which consisted of
North America and Eurasia) also moved away from Africa and South America,
helping to create the Atlantic Ocean and the Gulf of Mexico. Volcanic activity
was common along these rifting continental margins. At the same time, Eurasia
(Europe and Asia) moved to the south and started to close off the Tethys
Ocean. Sea level gradually rose during the Jurassic, creating epicontinental
seaways in North America and Europe. The end result was a world with many
more separate land-masses, and a great deal more coastline, than the world of
the Triassic.
Much of the Jurassic world was warm and moist, with a greenhouse
climate. Coal deposits formed under forests in Australia and Antarctica.
Although some arid regions remained, much of the rest of Pangea was lush and
green. Northern (Laurasian) and southern (Gondwanan) biotas were still
distinct in many ways, but by the Jurassic, faunas had acquired a more
intercontinental character. Some animals and plants were now found nearly
worldwide, instead of being restricted to particular regions.
Defining Characteristics:
o extinction of the dinosaurs
o first appearance and diversification of flowering plants
o extreme global warming
Secondary Characteristics:
o oldest known specimens of termites, ants, and bees
o diversification of birds and many new insect groups
o new kinds of dinosaurs
Cretaceous, including the oldest known ants and bees as well as newly evolved
groups of pollinating species such as flies, beetles, wasps, and moths. Some
paleontologists think that the coincident evolution of these insect groups and
the diversification of flowering plants is an example of the process of
coevolution, in which two different types of organisms (such as an insect and
plant) become specifically adapted to one another. Insects also evolved more
types of feeding behavior both in quiet-water habitats such as lakes and in
flowing-water habitats.
Life in the Cretaceous Seas
Reptiles were not the only marine giants of the Cretaceous. Strangelooking, often gigantic rudistid clams, reminiscent of Paleozoic horn corals,
reached up to one meter in length and formed extensive reefs in shallow
tropical oceans. Inoceramid clams over three meters long occurred in shallow,
warm seas, including environments that were nearly devoid of oxygen.
Ammonite cephalopods continued to diversify into amazing sizes and shapes,
with some coiled forms over two meters across, and other forms that
resembled an extended hook over two meters long.
Extinction of the Dinosaurs
Perhaps the most notable event of the Cretaceous was its conclusion.
About 65 million years ago the second greatest mass extinction in Earth history
occurred, resulting in the loss of the dinosaurs as well as nearly 50% of all the
worlds species. Though not nearly as severe as the end-Permian mass
extinction, the end-Cretaceous extinction is the most famous mass extinction
in Earth history. Other great animals also went extinct at that time, including
flying reptiles (pterosaurs) and the last mosasaurs and plesiosaurs. Many
mollusks, including rudistid and inoceramid clams, ammonites, and belemnites,
also became extinct, as did many species of microscopic marine plankton.
Terrestrial plants also suffered a major extinction at this time; in some regions
up to 60% of latest Cretaceous plant species were absent in the subsequent
Paleocene. Terrestrial insects also suffered a high level of extinction, especially
those that were highly specialized to feed on one or a few types of plants. In
fact, the level of insect herbivoryboth generalized and specializeddid not
recover to latest Cretaceous levels until the Paleocene-Eocene boundary,
approximately 9 million years later. In spite of the severity of extinctions at the
end of the Cretaceous, many types of animals and plants survived and gave
rise to new groups of organisms in the Paleocene.
dinosaurs were already in decline before the asteroid impact, so that its
environmental effects merely hastened their extinction. Alternatively, others
point to the high abundance and variety of dinosaur species recorded even in
the sediments deposited just below the asteroid impact layer in the Hell Creek
Formation of western North America.
Shifting Continents and Greenhouse Climates
Defining Characteristics:
o Age of Mammals begins
o flowering plants and conifers abundant
Secondary Characteristics:
o global climate warming
o abundant plants, fish, crocodiles, and mammals
vertebrates. Although Paleocene bird fossils are rare, mammals are well
represented in Paleocene sediments. Insectivorans, early relatives of true
primates (plesiadapiforms), carnivorans, creodonts, and primitive
herbivores (such as condylarths and early uintatheres) inhabited the
forests. The largest mammal, Pantolambda (a primitive plant eater), was
about the size of a small pony. The multituberculates, small mammals with
chisel-like front teeth that had evolved in the Mesozoic, remained common
in the Paleocene; rodents appeared late in the Paleocene.
Through most of the Paleocene, fossil leaves show low amounts
and few types of damage caused by herbivorous insects. This suggests
that insect herbivores were slow to recover diversity and abundance
following the end-Cretaceous extinction. The diversity of insect feeding
marks on plants did not recover to Cretaceous levels until the late
Paleocene.
The marine world of the Paleocene was much more like the
modern marine realm than that of the Cretaceous. In particular, some of
the largest and most common Mesozoic animals had vanished. Although
many Cretaceous species of invertebrates and fishes survived the
Cretaceous extinction event, among aquatic reptiles only turtles,
crocodilians, and champsosaurs (a freshwater, crocodile-like fish eater)
persisted into the Paleocene. Sharks are represented by mackerel sharks,
several genera of sand tiger sharks, and the first small-toothed white
shark. Bony fishes, especially teleosts, became more common as well.
New forms of sea urchins and foraminiferans appeared, alongside more
modern forms of gastropods and bivalves
Defining Characteristics:
o first appearances of many modern mammal orders
o maximum extent of warm climate and tropical vegetation
Secondary Characteristics:
o evolution of marine mammals
o cooling climate
In 1833, Charles Lyell derived the name Eocene from the Greek
words Eos (meaning dawn) and Kainos (meaning recent). At the time,
the Paleocene had not yet been named, so the Eocene became the dawn
of the recent (Cenozoic Era). Lyell chose this term because only about
3.5% of fossil mollusks from sediments of this age were recent species.
During the Eocene, volcanoes were active in the Rocky Mountains as the
uplift of this region was completed. The rising mountains were eroded into
sediment that filled the adjacent basins, which (along with nearby large
lakes) became important fossil sites in Wyoming and Colorado.
Spectacular Eocene fossils come from lake deposits at Messel, in
Germany, and from the Green River Formation in southwestern Wyoming.
During the warmest part of the early Eocene, palm trees grew as
far north as Alaska and Spitsbergen Island in the North Atlantic.
Crocodilians lived above the Arctic Circle, and forests of dawn redwoods
grew at 80 N latitude. As climate cooled and became more seasonal
during the middle and late Eocene, forests gave way to dry woodlands,
perhaps with open patches of grasses and herbs. The Eocene saw changes
in the distribution of plants, as some species occupied new geographic
regions. Most of the modern bird orders were present by the Eocene, as
well as several unusual, now-extinct species. One, the North American
Diatryma, was a flightless, six-foot-tall predator.
Among the cartilaginous fishes, modern forms such as requiem
sharks increased, while sand tiger sharks decreased. By the Middle
Eocene, the giant sand tiger shark Otodus obliquus was extinct. Mako and
giant-toothed white sharks first appeared at this time as well. However,
bony fishes continued to dominate the seas, as they do today. Marine
invertebrates were also more modern. Paleozoic Fauna animals such as
brachiopods were uncommon, while cephalopods, echinoderms, snails,
and bivalves thrived.
Defining Characteristics:
o transition of woodlands to open grasslands
o appearance of most living families of mammals
o appearance of the two orders (toothed and baleen) of living
whales
o initiation of North Atlantic deep-water formation
Secondary Characteristics:
o development of the Antarctic circumpolar current
o erosion of the Rockies
o decline of browsing mammals
eastern opening to the Indian Ocean. The Arabian peninsula and Iran
pushed into Asia Minor. The Atlantic continued to widen.
ii. Neogene
Defining Characteristics:
o half of marine invertebrates species are extant forms
o uplift of land separates Tethys Ocean from Indian Ocean
o ocean circulation changes to form major gyres (circles) in N &
S hemispheres
o more seasonal climates in N. Hemisphere
Secondary Characteristics:
o horses and even-toed herbivorous mammals diversify
o whales, seals, sea lions, and walruses diversify
ungulates, also appeared, as did our own near ancestors in Africa, the first
anthropoid apes.
In the seas, marine crocodiles were still common and lived as far
north as Maryland. Many species of marine mammals also inhabited the
seas, including baleen and toothed whales, seals, sea lions, walruses, and
sea cows. Both toothed and baleen whales were particularly diverse
during this period. One odd marine mammal that had appeared in the
Oligocene and was common in the North Pacific Ocean during the Miocene
was called Desmostylus. It had four legs that were well adapted both for
living on land and for swimming, and its teeth were shaped like bundles of
cylinders. Distant relatives of elephants and sea cows, Desmostylus and
its kin were entirely extinct by the end of the Miocene.
Defining Characteristics:
o continuation of Age of mammals
o appearance of early bipedal ancestors of humans
o Great American Interchange
Secondary Characteristics:
o expansion of grasslands
o beginning of separation between Atlantic and Pacific marine
organisms
beginning of the familiar modern faunas; many of the plants and animals
from that time have continued to live and evolve to the present day.
Indeed, Lyell devised the name because he noticed that of all the
Cenozoic marine fossils, those from Pliocene strata were the most similar
to modern forms. This characteristic distinguished these layers from
earlier Miocene or Eocene strata.
During the Pliocene, the Panamanian land bridge linked North and
South America, allowing terrestrial species to migrate between the two
continents. This event is called the Great American Faunal Interchange, a
time when two long-isolated faunas came into contact. Sixteen native
southern genera moved to the north, including armadillos, giant ground
sloths, flightless predatory birds, marsupials (including opossums), and
porcupines. At the same time, 23 native northern genera moved south,
including cats, dogs, bears, tapirs, camels, and certain rodents. The
exchange was not a balanced one, however. Many more South American
immigrant species became extinct, perhaps as a result of competition and
the inability to adapt to new conditions. As a result, many North American
species now live in South America, but few South American imports still
survive in the north.
Defining Characteristics:
o significant human geographic expansion and cultural
development
o marked climatic fluctuations and glacial events
Secondary Characteristics:
o plants and animals very similar to surviving modern forms
o first major human-influenced extinctions
Because so much water was taken up as ice, global sea level dropped
approximately 140 meters.
Defining Characteristics:
o impact of Homo sapiens and technology
Secondary Characteristics:
o climate warming following the last ice age
o continents drying out, polar areas contract
o plant communities shifting with the climate
The term Holocene was first proposed at the third International
Geological Congress in 1885. It comes from the Greek words holos
(meaning whole) and kainos (meaning recent), referring to fact that
this epoch is the most recent division of Earth history. However, many
scientists also used the term Recent or Postglacial for this epoch until
1967, when the U.S. Geological Survey formally adopted the term
Holocene and discontinued the use of Recent.