Sie sind auf Seite 1von 43

!

"#$%&%'()*
+,#)-#./01!2&'#%.&)&%3)
45%67.6')-2/63
!"#$%&'()$*&++&,$
!-.' /0$123'-'"453$6$7),&-5/8/9)
:;<=
89:#'.6;#0)
1>-">4-?$/0$@-"'484A&'4/,$6$=B.8&,'&'4/,
C"/.(/28&3'45$+400-"-,'4&'4/,
D8&5-,'&8$/"9&,4A&'4/,E+->-8/.B-,'$
D8&5-,'&8$9"/?'($6$B&'F"&'4/,$
@-'&8$+->-8/.B-,'
@-'&8E*&'-",&8$54"5F8&'4/,$
:B24845&8$5/"+$&,+$&B,4/'45$08F4+#
!"#$%&%'(
C(-$3'&'-$/0$5&"")4,9$&$+->-8/.4,9$0-'F3$?4'(4,$'(-$2/+)#
-#".626<&.67%
<4'(4,$/,-$+&)$B/"F8& 43$5/,>-"'-+$4,'/$28&3'/5)3'
5/,343'4,9$/0$3&B-$'?/$9"/F.3$/0$5-883G$,/?$?4'($
+400-"-,'$,&B-3H
IB2")/28&3' +-"4>-+$0"/B$=,,-"$%-88$*&33
C"/.(/28&3' +-"4>-+$0"/B$1F'-"$%-88$*&33
!"#$%&'()*+'*',%*
'-.%*"/*
-.&0+#'+'-"#1*
2..*3*4*5%%6)*
/7".*89:
;#<*"/*
$"#$%&'()*
)'+=%1*
>0+)'"$?)'*3*
@..1*@*5%%6)*
/7".*89:
A7-=-#*"/*%.>7?"#-$*'-))(%)
B%C%0"&.%#'*"/*',%*%.>7?"*/7".*45%%6)*'"*DE*5%%6)
!2&'#%.&)=)>&.6%)8"6$6%)? @2&.)'&A#B
%(&"&5'-"43'45$/0$B&BB&83#
J/4,3$B/'(-"$&,+$/003."4,9$0/"$,-5-33&")$."/>434/,3$'/$-,3F"-$
3F3'-,&,5-$0/"$+->-8/.B-,'$4,$F'-"/#
=,$(49(-"$/"+-"$."4B&'-3$4'$43$!"#$%!$&'() K B&'-",&8$28//+$5/B-3$4,$
+4"-5'$5/,'&5'$?4'($'(-$0-'&8$5(/"4/,#
C?/$./"'4/,3$H$@-'&8$5/B./,-,'$0"/B$'(-$5(/"4/,45$3&5$L5(/"4/,45$
0"/,+/3FBM
*&'-",&8$5/B./,-,'$0"/B$'(-$-,+/B-'"4FB$L$+-54+F&
2&3&843M
<4'(4,$/,-$+&)$B/"F8& 43$5/,>-"'-+$4,'/$28&3'/5)3'
5/,343'4,9$/0$3&B-$'?/$9"/F.3$/0$5-883G$,/?$?4'($
+400-"-,'$,&B-3H
IB2")/28&3' +-"4>-+$0"/B$=,,-"$%-88$*&33
C"/.(/28&3' +-"4>-+$0"/B$1F'-"$%-88$*&33
IB2")/28&3' 0/"B3$'(-$-B2")/$."/.-"$
C"/.(/28&3' 0/"B3$'(-$.8&5-,'&$&,+$&33/54&'-+$
B-B2"&,-3#
The phases of blastocyst im plantation .

Fulle r W B aze r et a l. Re production 2 00 9;1 38 :19 5-20 9


+,#)C#'63/&
C(-$-,+/B-'"4FB /0$'(-$F'-"F3$&0'-"$4B.8&,'&'4/,$43$5&88-+$+-54+F&#
+-54+F& 2&3&843 K =BB-+4&'-8)$2-,-&'($'(-$-B2")/$'(-$5(/"4/,45$>4884 &,5(/"$4,'/$
'(43$"-94/,#C(43 0/"B3$.&"'$/0$'(-$.8&5-,'&#$
+-54+F& 5&.3F8&"43 K C(-$'(4,$8&)-"$/0$-,+/B-'"4FB '(&'$3F""/F,+3$'(-$4B.8&,'-+$
-B2")/#
+-54+F& .&"4-'&843 K -,+/B-'"4FB 84,4,9$'(-$"-B&4,+-"$/0$'(-$F'-"F3#$NO$?--P3$
'(43$0F3-3$?4'($'(-$+-54+F& 5&.3F8&"43 /284'-"&'4,9$'(-$F'-"4,-$8FB-,#
!-54+F&$ !-54+F&$
5&.3F8"43 2&3&843

!-54+F&$
.&"4'&843
C"/.(/28&3'3 3'&"'$."/840-"&'4,9$"&.4+8)$&,+$+400-"-,'4&'-$
4,'/$'?/$8&)-"3H
%)'/'"/.(/28&3' /"$5-88F8&"$'"/.(/28&3'
Q),5)'4&8 '"/.(/28&3' L3),5)'4/'"/.(/28&3'M
D62267/0 '6"'/2&.67%
R)$-,+$/0$?--P$N$'/$2-94,,4,9$/0$?--P$SG04,9-"3$/0$"&.4+8)$+4>4+4,9$
5)'/'"/.(/28&3' ."/T-5'$4,'/$'(-$3),5)'4/'"/.(/28&3'G$0/"B4,9$>4884$'(&'$&"-$2&'(-+$
4,$'(-$B&'-",&8$28//+#$C(-3-$&"-$'-"B-+$&7-.+7?*C-00-#$

<(-,$'(-$."4B&")$>4884 2-5/B-$0488-+$?4'($B-3-,5()B- 0"/B$'(43$B-3/+-"BG$


'(-)$&"-$5&88-+$)%$"#<+7?*C-00-#$

R)$'(-$-,+$/0$'(-$'(4"+$?--P$L!&)$NUMG$.-,-'"&'4,9$B-3/+-"B$+400-"-,'4&'-$4,'/$
28//+$5-883$&,+$3B&88$28//+$>-33-83$P,/?,$&3$'%7'-+7?*C-00-F
E%;&067%)7@)F&.#"%&2)G2773);#00#20
V$04,&8$5"4'45&8$3'-.$4,$-3'&2843(4,9$'(-$F'-"/.8&5-,'&8 >488/F3$
54"5F8&'4/,$4,>/8>-3$'(-$4,>&34/,$/0$F'-"4,-$>-33-83$L3.4"&8$&"'-"4-3M$2)$
'(-$5)'/'"/.(/28&3'3#$
C(-3-$4,>&34>-$5-883$.-,-'"&'-$'(-$28//+$>-33-83G$"-.8&54,9$'(-$
-,+/'(-84FB$&,+$BF5($/0$'(-$'F,45&$B-+4&#
HI6"&2)4".#"()J#573#226%$
I,+/>&35F8&"$4,>&34/,$2)$
-W'"&>488/F3 '"/.(/28&3' +F"4,9$04"3'$
&,+$3-5/,+$'"4B-3'-"3
X/33$/0$3B//'($BF358-$&,+$
-,+/B-'"4&8$84,4,9
Y-3F8'3$4,$+48&'4/,$/0$'(-$>-33-8$
&,+8/33 /0$>&3/"-&5'4>4')
%/,>-"'3$>-33-8$4,'/$8/?$
."-33F"-L"-343'&,5-MG$$8&"9-"$
+4&B-'-"
=,4'4&88)$&$.8F9$/0$042"4,/4+$43$+-./34'-+$4,$'(-$8FB-,$/0$'(-$28//+$>-33-83G$&88/?4,9$/,8)$B&'-",&8$
28//+$.8&3B&$&,+$,/'$"-+$28//+$5-883$'/$08/?$4,'/$'(-$4,'-">488/F3 3.&5-G$'(F3$P--.4,9$/W)9-,$
8->-83$8/?#$

V,&-"/245$98)5/8)343$."/>4+-3$'(-$-,-"9)$"-ZF4"-B-,'3$0/"$'(-$-&"8)$3'&9-3$4,$/"9&,/9-,-343#$

C(43$B4,4B4A-3$'(-$"43P$/0$+&B &9-$'/$![V$2)$"-&5'4>-$/W)9-,$3.-54-3$9-,-"&'-+$+F"4,9$&-"/245$
B-'&2/843B#$

X/?$/W)9-,$8->-83$&"-$&83/$"-ZF4"-+$0/"$."/840-"&'4/,$/0$5)'/'"/.(/28&3'3 &,+$0/"$'(-4"$4,>&34/,$
&,+$&,5(/"4,9$'/$'(-$+-54+F&#$

R)$UOKUN$?--P3$/0$9-3'&'4/,G$'(-3-$.8F93$2"-&P$+/?,$&,+$B &'-",&8$?(/8-$28//+$'(-,$08/?3$4,'/$
'(-$4,'-">488/F3 3.&5-#$C(43$"-3F8'3$4,$&,$4,5"-&3-$4,$/W)9-,$8->-83$&,+$4,(424'4/,$/0$
5)'/'"/.(/28&3'45 4,>&34>-,-33#
+,#)9&""6#")9#.K##%)5&.#"%&2)&%3)@#.&2)92773
C(-$0F88)$0/"B-+$(FB&,$.8&5-,'&$43$'-"B-+$(-B/5(/"4&8 2-5&F3-$'(-$B&'-",&8$
28//+$43$3-.&"&'-+$0"/B$'(-$0-'&8$28//+$/,8)$2)$-8-B-,'3$/0$'(-$5(/"4/,#$
C(-$5/B./,-,'3$/0$'(-$2&""4-"$2-'?--,
Q),5)'4/'"/.(/28&3'
%)'/'"/.(/28&3'
2&3&8$8&B4,&
0-'&8$B-3-,5()B-G$
2&3&8$8&B4,&$/0$'(-$0-'&8$5&.488&")
-,+/'(-84FB$/0$'(-$0-'&8$5&.488&")#
-/%'.67%)7@).,#)!2&'#%.&

C"&,30-"$/0$,F'"4-,'3$6$?&3'-$."/+F5'3L$"-3.4"&'/")G$-W5"-'4>-$&,+$
,F'"4'4>-M

D"/+F5'4/,$/0$&,+$B-'&2/843B$/0$(/"B/,-3$,--+-+$'/$B&4,'&4,$
."-9,&,5)

D"/'-5'4>-$2&""4-"$&9&4,3'$,/W4/F3$&9-,'3$4,$B&'-",&8$54"5F8&'4/,

=BBF,/8/945&8$."/'-5'4/,K 042"4,/4+$&,+$34&8/BF54, 5/&'-+$3F"0&5-G$


>-")$84''8-$;XV$&,+$28//+$9"/F.$&,'49-,$/,$'"/.(/28&3'45$3F"0&5-#
H."/'./"#
V'$C- "BH$
Q(&.- H$+435/4+#$

!4&B-'-"H$NOKN\$5B#$

<-49('H$\OOK]OO$$9B#

C(45P,-33H$SK^$5B$&'$4'3$5- ,'-"$&,+$9"&+F&88)$'&.- "3$'/?&"+3$'(-$.-"4.(-")#

_488/F3$3F"0&5-$&"-&$U`$BN

D/34'4/,H$4,$'(- $F..-"$F'-"4,-$3-9B-,'$Laa#\bMG$-4'(-"$4,$'(-$./3'-"4/"$3F"0&5-$LNESM$/"$'(-$&,'-"4/"$3F"0&5-$
LUESM#
H."/'./"#
QF"0&5-3H$$
@/-'&8 3F"0&5-H$3B//'(G$9843'-,4,9$&,+$43$5/>-"-+$2)$'(-$&B,4/,$
?(45($43$"-08-5'-+$/,$'(-$FB 24845&8$5/"+#$C(-$FB24845&8$5/"+$43$
4,3-"'-+$,-&"$/"$&'$'(-$5-,'-"$/0$'(43$3F"0&5-$&,+$4'3$"&+4&'4,9$
2"&,5(-3$5&,$2-$3--,$2-,-&'($'(-$&B,4/,#

*&'-",&8$3F"0&5-H$+F88$9"-)43(E28F-$"-+$4,$5/8/F" &,+$43$+4>4+-+$
4,'/$U\KNO$5/')8-+/,3#$I&5($5/')8-+/,$43$0/"B-+$/0$'(-$2"&,5(-3$
/0$/,-$B&4,$>488F3 3'-B$5/>-"-+$2)$+-54+F& 2&3&843#
+,#)L59626'&2)M7"3)
C<1$QCVXcQ$&"-$3--,$&'$'(-$2-94,,4,9$/0$+->-8/.B-,'
V$>-,'"&8$/,-G$'(-$)/8P$3&5$3'&8PG$5/,'&4,4,9$'(-$>4'-884,- +F5'$&,+$'(-$
>4'-884,- >-33-83
V$5&F+&8$/,-G$'(-$5/,,-5'4,9$3'&8PG$5/,'&4,4,9$'(-$&88&,'/43$&,+$'(-$
FB24845&8$>-33-83
C(-$5/,,-5'4,9$3'&8P$."/9"-33-3$>-,'"&88)$&3$&$"-3F8'$/0$'(-$
5-.(&8/5&F+&8 08-W4/,$/0$'(-$-B2")/$&,+$0F3-3$?4'($'(-$)/8P$3&5$3'&8P$
'/$0/"B$'(-$FB24845&8$5/"+
+,#)L59626'&2)M7"3)

C;I$:*R=X=%VX$%1Y!$43$3(/"'$&,+$'(45P$4,$'(-$)/F,9$-B2")/$&,+$43$
4,3-"'-+$4,$'(-$8/?-"$./"'4/,$/0$'(-$>-,'"&8$"-94/,$/0$'(-$-B2")/
<4'($'(-$+->-8/.B-,'$/0$'(-$&,'-"4/"$&2+/B4,&8$?&88G$'(-$"-94/,$/0$
FB24845&8$4B.8&,'&'4/,$5/,'"&5'3G$'(-$5/"+$-8/,9&'-3G$&,+$&83/$
2-5/B-3$38-,+-"
V'$'-"BG$'(-$5/"+$5/,'&4,3$/,8)$'(-$FB24845&8$>-33-83$3F""/F,+-+$2)$
<(&"'/,d3$T-88)$L&$3B//'($B-3-,5()B/F3 B&'-"4&8G$&$BF5/F3$
+400-"-,'4&'4/,$."/+F5'$/0$B-3/+-"BM$&,+$43$&2/F'$\OK`O$5B$8/,9$&,+$
&2/F'$N$5B$4,$+4&B-'-"
N#"#)60)&)%7"5&2).,"##);#00#2)/59626'&2)'7"3O
P7.#).,&.).,#"#)&"#).K7)&".#"6#0 .7K&"3).,#)"6$,.)&%3)&)06%$2#)
;#6% &.).,#)2#@.O)
F70.)7@).,#)'7"3)'7%060.0)7@)&)2770#)5#0#%',(5#)K6.,)
6%.#"'#22/2&")$"7/%3)0/90.&%'#)=Q,&".7%R0):#22(BO
45%67.6')@2/63)
+"&%0@#")S P/."6.67%
T%3)

Das könnte Ihnen auch gefallen