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Journal of Personality 00:00, Month 2017

Cognitive-Adaptive Trait Theory: A C 2017 Wiley Periodicals, Inc.


V
DOI: 10.1111/jopy.12319

Shift in Perspective on Personality

Gerald Matthews
University of Central Florida

Abstract
Traditional, biologically based trait theories have deservedly gained broad acceptance, but some long-standing core issues of
personality research remain unresolved. Recent research questions whether (a) there can be a single universal structural
model of personality superfactors, (b) current theory adequately specifies the processes that mediate behavioral and emo-
tional expressions of traits, and (c) brain-based accounts of traits adequately explain their role in real-world functioning and
adaptation. This article reviews the perspective on these issues provided by cognitive-adaptive trait theory. This theory rejects
the view that personality dimensions directly reflect brain systems. Instead, traits correspond to variation in strategies for
managing key adaptive challenges. Thus, each trait is expressed in environments that pose those challenges, and each trait cor-
responds to skills and self-knowledge that facilitate adaptation to those environments. The cognitive-adaptive theory affords
novel perspectives on trait psychometrics, theoretical accounts of mediating processes, and real-world adaptation.

Keywords: Personality traits, cognitive processes, skill, adaptation, psychometrics

Traits are dened simply as stable dispositions, that is, descrip- demonstrate that major traits have a biological basis (DeYoung
tions of individual differences in styles of thought, feeling, and & Gray, 2009; Kennis, Rademaker, & Geuze, 2013). Evidence
behavior. Modern accounts of traits (Deary, 2009) go beyond from behavior genetics on the heritability of traits is consistent
description to assign a causal status to traits. Typically, trait the- with psychobiological accounts, although linkages between
ories include both a structural description of major trait dimen- traits and specic polymorphisms have proved elusive
sions, such as the Five-Factor Model (FFM; McCrae & Costa, (Turkheimer, Pettersson, & Horn, 2014). Third, behavioral
2008), and an account of neural or social-cognitive processes expressions of traits are increasingly understood within main-
that mediate between traits and individual differences in behav- stream psychological theory. The major subdisciplines of the
ior. Different versions of trait theory differ psychometrically, eld, including biological, cognitive, social, and health psychol-
regarding which traits qualify as major superfactors, and theoret- ogy, provide an array of explanatory models for behaviors and
ically, in terms of which neural or psychological systems are the other relevant outcomes. Studies of traits are not some arcane
key causal agents. However, the leading theories, reviewed by numerology but an indispensable facet of psychological science.
Matthews, Deary, and Whiteman (2009), share common Fourth, traits predict various consequential life outcomes, sup-
assumptions, including some degree of behavioral consistency porting applications including personnel selection, clinical guid-
of the individual, stability over time, biological basis for traits, ance, and educational interventions (Ozer & Benet-Martnez,
and cross-cultural generality (McCrae & Costa, 2008), sugges- 2006; R. Roberts, Martin, & Olaru, 2015). These advances have
ting that an eventual standard theory of traits may emerge. These persuaded many researchers that traits are more than mere social
assumptions are also shared by accounts of personality that rec- constructions (Matthews et al., 2009).
ognize constructs beyond dimensional traits such as characteris- Signicant challenges remain for the trait perspective. While
tic life adaptations and integrative life narratives (McAdams & many researchers see the FFM as at least a pragmatically useful
Pals, 2006). Naturally, trait assumptions remain open to critical frame of reference, psychometric uncertainties remain over
questioning (e.g., Cervone, 2008); this article addresses the extensions to the FFM, development of more granular trait mod-
broad consensus of trait researchers. els, and the relationships between stable trait and transient state
Matthews et al. (2009) summarized four key accomplish- dimensions. Theoretical accounts of traits often take a rather
ments that support the acceptance of trait theory. First, there is
sufcient convergence of psychometric measurement models,
Correspondence concerning this article should be addressed to Gerald
including the FFM, for optimism about future prospects for a Matthews, Institute for Simulation and Training, University of Central
consensus personality structure. Second, psychophysiological Florida, 3100 Technology Parkway, Orlando, FL 32826. Email: gmatthews@
studies, which increasingly employ brain-imaging methods, ist.ucf.edu.
2 Matthews

broad-brush approach of limited use for predicting individual There may also be culturally specic factors that are of compara-
differences in behavior in specic contexts. Theory is also lim- ble importance to the Big Five in the culture concerned, such as
ited in its treatment of how traits make a difference to the indi- those associated with Chinese values (Fetvadjiev & Van de
vidual in real-life settings, and the processes mediating between Vijver, 2015).
traits and adaptation to life challenges (Cervone, 2008).
In this article, I will consider solutions to some of the chal- Contextualized Traits. The universality of the FFM is part of
lenges facing contemporary trait theory. I will do so from the its appeal. However, traits that are tied to a specic context may
perspective of research on personality and human performance be more predictive of performance. For example, in evaluative
in laboratory and real-life settings, using tasks that provide contexts, traits such as test anxiety and math anxiety may be pre-
objective measures of behavior such as accuracy and response ferred to broad traits (Zeidner, 1998). Similarly, stress response
time. The cognitive-adaptive theory of traits (CATT; Matthews, to vehicle driving is better predicted by traits tied to the driving
1999, 2004, 2008a, 2016; Matthews et al., 2009) was developed context than by broad traits (Matthews, 2002).
to explain associations between traits and performance. In this
article, I focus on the wider implications of the theory for under- Differentiating Traits and States. Transient states are
standing traits and their behavioral impacts. Specically, I will important for trait psychology, as they are frequently supposed
identify some burning issues that remain for the psychomet- to mediate trait expression. For example, arousal states played a
rics, theoretical bases, and real-world signicance of traits and central role in H. J. Eysencks theory (1967). However, there is
then outline the unique perspectives and possible solutions pro- less consensus on their psychometric treatment. One approach is
vided by CATT. to match states directly to traits, as in traitstate anxiety research
(Spielberger & Reheiser, 2009). Positive and negative affect are
seen as counterparts to E and N (Watson, 2000), leaving unde-
UNRESOLVED ISSUES IN PERSONALITY ned state mediators of other traits. In the performance context,
THEORY there is no direct correspondence between trait and state dimen-
Psychometric Uncertainties sions (Matthews et al., 2002).
McCrae and Costa (2008; McCrae et al., 2000) claim that psy-
chometric modeling of data from various personality measures Theoretical Issues: Mediators of Trait Effects
consistently reveals the ve dimensions of Extraversion (E),
Developing a theory of traits requires specifying the underlying
Neuroticism (N), Conscientiousness (C), Agreeableness (A),
and Openness (O). Behavior-genetic studies show substantial neural or psychological processes indexed by the trait, processes
heritability for each FFM dimension (typically 0.5, including that may mediate associations between the trait and behavior.
nonadditive factors). The FFM shows high rank-order stability Biological trait models offer an especially clean paradigm of this
(r .75) in adulthood, with the factor structure remaining stable kind, to the extent that traits can be linked to neural processes or
over time. The FFM factors are said to be replicated in most cul- systems (Corr, 2009; H. J. Eysenck, 1967). Unfortunately,
tures and recognizable in all. Most trait researchers see the FFM despite evidence for correlations between traits and various indi-
as at least a useful working approximation of psychometric ces of brain functioning (DeYoung & Gray, 2009), there seems
structure, but whether the FFM is a precise and universal mea- to be no one-to-one correspondence or isomorphism between
surement model remains open to debate. The extent to which specic traits and specic brain systems (Matthews, 2004; Zuck-
latent factor modeling identies the FFM remains controversial erman, 2005). The heritability of traits afrms their biological
(Boyle, 2008), and scales do not meet strict criteria for measure- basis, but it is frustratingly difcult to nd brain markers that
ment invariance across cultures (Church, 2016; Fetvadjiev & correlate highly and reliably with trait measures (for more
Van de Vijver, 2015). detailed critiques of biological trait theory, see Matthews,
The major traits correlate with various aspects of task per- 2008b, 2016; Matthews & Gilliland, 1999).
formance (for reviews, see Matthews, 2008a, 2009). Increas- Personality traits may also express psychological rather than
ingly, studies cover all of the FFM, as well as additional physiological constructs. From a clinical perspective, Beck (e.g.,
traits such as anxiety (M. W. Eysenck & Derakshan, 2011). Clark & Beck, 1999) identied various systematic distortions in
Laboratory tasks include those requiring speeded response, self-beliefs and reasoning that accompany depression and anxi-
attention, and memory; real-world tasks support job and aca- ety. If every person has a self-schema that encodes his or her per-
demic performance (Matthews et al., 2009). However, several ceived self-worth in long-term memory, along with idiographic
issues remain. self-beliefs, then we can dene self-esteem as a nomothetic con-
struct that inuences traits such as N. From a more process-
Additional Superfactors. Several authors make persuasive oriented standpoint, traits are based in cybernetic goal-seeking
cases for adding to the Big Five. For example, the HEXACO processes that regulate the self (Carver & Scheier, 2001) or in
model adds an additional Honesty factor supported by substan- information-processing constructs such as executive function
tial psychometric evidence (Ashton, Lee, & De Vries, 2014). (M. W. Eysenck & Derakshan, 2011).
Cognitive-Adaptive Theory 3

Many trait psychologists distinguish levels of theorizing apply to understanding ourselves and other people (Mischel,
associated with biological and social-cognitive underpinnings 1968). The systematic distortion hypothesis (Schweder, 2007)
for personality. The distinction is sometimes expressed as one states that traits are the product of a preexisting model of con-
between temperament and personality, where temperament ceptual associations between personality data. The model biases
refers to basic, primarily biological, individual differences evi- memory and inferencing processes when people complete per-
dent in early childhood, and personality is acquired patterns of sonality questionnaires. The application of multimethod dimen-
thought, behavior, and socialization built on the temperamental sional models that incorporate other-ratings (Chang, Connelly,
platform (McCrae et al., 2000; Rothbart, Sheese & Conradt, & Geeza, 2012) argues against the hypothesis, but, given the
2009; Strelau, 2001). This approach is starkly realized in prevalence of cognitive biases and heuristics, such critiques
McCrae and colleagues (2000) ve-factor theory (FFT). The should be taken seriously. Bias in self-perception is likely to fur-
FFM is a structural description of traits, not a theory. FFT aims ther weaken the utility of self-reports as markers for latent brain
to explain the FFM traits as representing temperamental basic characteristics.
tendencies that shape learning of characteristic adaptations,
including skills, habits, beliefs, roles, relationships, and elements Specifying Characteristic Adaptations. In FFT (McCrae &
of the self, though developmental processes. FFT is a paradigm Costa, 2008), characteristic adaptations is an all-embracing
that represents much current thinking about personality and a notion of numerous psychological constructs. The broad deni-
possible avenue toward a standard trait model. tion captures the pervasiveness of trait inuences, but it has the
For the performance psychologist, this catholic approach to disadvantage of mixing together constructs that may play differ-
theory building allows for some trait effects to be mediated ent roles in mediating trait inuences on behavior. There are sev-
directly by neural processes, and others to reect more context- eral conceptual distinctions that are important to accommodate
bound psychological processes such as strategy choice. In fact, theoretically, as stated below.
the cognitive theory of performance recognizes an equivalent
distinction between structural effects that reect variation in Stable Personality Attributes vs. States. Personality
basic processing efciency and strategic effects that reect effects in part reect temporally stable, specic operating
voluntary control over processing (Hockey, 1986). Thus, con- parameters of the mind (Matthews & Harley, 1993). Such
temporary personality theory captures something of the joint parameters moderate stimulus-response (S-R) relationships.
biological and social-cognitive character of traits that lends itself However, as explored in anxiety research (Spielberger & Rehe-
to explanatory models for individual differences in behavior. iser, 2009), some effects of personality are mediated by broader-
Nevertheless, signicant theoretical challenges remain. based changes in transient psychological states.

Mismatch of Constructs and Assessments. If dimensions Content vs. Process. Personality is related to both stable
of personality or temperament are biologically based, they elements of self-knowledge, such as self-esteem, and individual
should be most validly assessed objectively. Indeed, neo- differences in processing routines, such as self-verication or
Pavlovian studies employed objective, psychophysiological social comparison (Wells & Matthews, 2015).
tests for constructs such as strength of the nervous system. Con-
versely, characteristic adaptations in FFT (McCrae & Costa, Core Personality Attributes vs. Contextualized Learning.
2008) include the self-concept and so are better suited to Some concomitants of personality, such as stress vulnerability
questionnaire assessment. As FFT identies the Big Five as and generalized self-esteem, may be intrinsic features of the trait
dimensions of temperament (basic tendencies), it is strange apt to inuence behavior across a variety of different situations.
that self-report remains the primary mode of assessment. Typical For example, trait anxiety may be associated with amplication
psychophysiological measures are subject to limitations such as of the threat value attached to stimuli (M. W. Eysenck & Derak-
temporal instability that may limit their usefulness as indices of shan, 2011). Other characteristic adaptations cited by McCrae
traits (Strelau, 1991), but it should nonetheless be possible to et al. (2000), including skills and habits, are learned and thus
assess stable individual differences in arousability or sensitivity context dependent.
to reinforcement. On the one hand, self-reports are only mod-
estly associated with basic neural mechanisms (Matthews & Gil- Explicit vs. Implicit Processes. The emergence of
liland, 1999), suggesting a lack of convergent evidence. implicit personality measures as a counterpart to explicit assess-
Convergence between self-report and objective, implicit person- ments (Gawronski & De Houwer, 2014) requires further articu-
ality measures is also limited (Gawronski & De Houwer, 2014). lation of how conscious and unconscious elements of
On the other hand, self-reports reect self-concept, suggesting a personality are interrelated.
lack of divergence from personality in the form of characteristic
adaptations. Accommodation of Cognitive Processes. In the perfor-
mance context, the constructs and methodologies of cognitive
The Role of Self-Perceptions. A long-standing criticism is psychology have proved critical for theory (M. W. Eysenck &
that traits are essentially artifacts of the cognitive processes we Derakshan, 2011; Humphreys & Revelle, 1984). Use of broad
4 Matthews

constructs such as general arousal simply does not support effec- according to the inverted-U principle. Reinforcement sensitivity
tive prediction of individual differences in performance theory (RST; Corr, 2009; Gray, 1987) states that activation of
(Matthews & Gilliland, 1999). Instead, major traits such as E are brain reward and punishment systems inuences individual dif-
associated with a cognitive patterning (Hockey, 1986; Mat- ferences in approach and avoidance behaviors. However,
thews, 2008a), that is, a set of independent, often modest, biases acquired skills rather than basic neural or cognitive processes
in multiple processing components associated with selective typically determine whether a person gains or loses from real-
attention, memory, problem solving, response execution, and so life encounters (Ericsson & Towne, 2010; Matthews, 1999). For
forth. Advancements in cognitive neuroscience increasingly example, enhanced selective attention to threat may be a core
connect these processing-based individual differences to brain attribute of trait anxiety (Cisler & Koster, 2010). However, in
mechanisms. The theoretical problem is that while we can real-life environments in which attention to threat is important,
develop cognitive mini-theories that apply to specic perfor- such as military surveillance in a multitasking environment
mance paradigms such as vigilance or selective attention to (Panganiban, Matthews, Knott, & Funke, 2011), performance in
threat, the multiplication of mechanisms threatens the coherence part reects learned skills for visual search and attentional con-
of a cognitive account of personality. How do we explain a trait trol, skills whose acquisition and execution are inuenced by
that has multiple and distinct small expressions, but no single anxiety. Personality, as well as cognitive abilities, will substan-
cognitive or biological basis? tially inuence individual differences in skill (Ackerman, 2003,
2014), but it is the skill that mediates the adaptive costs and ben-
ets of the personality trait.
Individual Differences in Adaptation: Traits in
Real Life
Traditional personality trait theory has a linear, feedforward
COGNITIVE-ADAPTIVE THEORY OF
structure (Matthews, 2004, 2016). Genes and childhood experi- PERSONALITY
ences dene the main attributes of personality at an early age, CATT starts from the premise that trait variation reects differ-
which are then expressed in emotional and behavioral form for ent strategies for adapting to environmental opportunities and
the remainder of life. In turn, individual differences in typical pressures (Matthews, 2008a). Somewhat in the spirit of J. J.
response inuence real-world outcomes such as mental health, Gibson, we should ask not only what is inside the head, but
educational and work success, and deviance from social norms. what the head is inside of. Traits may be most relevant to manag-
This picture captures something of the stability of personality, ing human social environments that are cognitively demanding,
evidenced in the reliable but modest correlations between child- such as attending a party where one does not have much in com-
hood assessments and adult outcomes (Caspi & Shiner, 2008), mon with the other people there. There is the scope for gains,
but it does not fully illuminate personalityenvironment across such as enjoyment of social interaction and opportunities to
the life span. Two issues that are especially challenging to tradi- make potentially useful contact, but realizing those gains
tional theory are the malleability of traits and the importance of requires specialized social skills. Extraversion then reects an
contextualized skills for real-life outcomes. adaptation to the environment that increases the likelihood of
proting from it. Note that adaptation here refers to selecting,
Stable and Malleable Elements of Personality. Recent implementing, and regulating strategies for managing external
research suggests that over time spans of years, traits may be more demands and challenges (Lazarus, 1991), not the evolutionary
malleable than previously believed. Changes in social roles drive sense of the term. Natural selection plays a role in shaping the
commensurate changes in FFM traits (Hudson, Roberts, & Lodi- brain systems that, indirectly, inuence the characteristic adapta-
Smith, 2012). The identication of the FFM with basic tempera- tions identied with personality traits, but evolutionary factors
ment requires either skepticism about the evidence for malleability are beyond the present scope.
(McCrae et al., 2000) or a reexamination of which personality con- This article focuses primarily on E and N. The latter trait
structs are more and less stable. In the traditional view, adult per- reects differing strategies for handling social threat. Other
sonality inuences the environment, but there is only limited FFM traits may be conceptualized adaptively (Matthews,
feedback from the environment to personality. For example, extra- 2008a). In brief, C contrasts sustained, systematic effort against
verts acquire larger networks of friends, but having many friends opportunism as strategies for success. A contrasts cooperative
does not elevate E. By contrast, dynamic interactionist views and competitive social strategies, and O is a choice between
assume a continuous interaction in which personality and environ- self-directed reasoning, curiosity, and exploration versus tradi-
ment mutually shape one another (Endler & Parker, 1992). tional received knowledge and pragmatism in handling intellec-
tual challenges. In CATT, trait characteristics are not
Context and Skill. Traditional personality theories assume intrinsically adaptive or maladaptive. E is typically seen as adap-
that basic attributes associated with traits have direct adaptive tive, but there is a downside of E, such as poor listening skills
impact. H. J. Eysencks (1967) theory supposes that individual and lack of appreciation of proactive behavior by subordinates
differences in cortical arousal directly impact performance (Judge, Piccolo, & Kosalka, 2009). Grant, Gino, and Hoffman
Cognitive-Adaptive Theory 5

(2011) found pizza stores with extraverted leaders only achieved attained). Self-regulation encompasses here many constructs
higher prots when employees were passive. Similarly, the investigated in personality research as potential mediators of
affective downside of N is balanced by greater awareness and trait expression, such as self-esteem, self-efcacy, affective
foresight for risk (Panganiban et al., 2011) and the motivational states, appraisal, coping, and emotional control. Self-regulation
advantages of defensive pessimism (Tamir, 2005). Indeed, adap- is dynamically related to skills. Actual skill typically correlates
tive value may vary with cultural and historical circumstances; with positive elements of self-regulation, such as higher self-
for example, high C is more adaptive in an orderly society in efcacy, but self-regulation is also biased by personality. For
which long-term effort pays off. example, trait anxiety and N appear to relate to both objective
CATT diverges from typical denitions of traits. Deary social skills decits in threatening contexts and to underestima-
(2009) points out that the trajectory of modern theory has been tion of personal competency (Matthews, 2004). Conversely,
toward replacing supercial, descriptive accounts of traits as col- quality of self-regulation can enhance or limit objective skill
lections of covarying behaviors with accounts of traits as latent execution (Kanfer & Ackerman, 1996). Effective self-regulation
causal entities, typically identied with properties of brain sys- may improve task focus and maintain motivation, whereas dys-
tems such as reticulo-cortical arousability (H. J. Eysenck, 1967). functional styles of self-regulation may lead to distraction due to
By contrast, in CATT, traits are constructs that emerge from col- personal worries and withdrawal of effort from the task. Thus,
lections of neural and cognitive processes that serve a common there is a three-way correspondence between traits, environ-
adaptive purpose. The emergent nature of traits poses challenges ments matched to the trait, and skills that are central to the trait
for both assessment and for causal modeling. Self-report scales and support adaptation.
assess explicit processes, and data on correlates of the FFM are Figure 1 illustrates the cognitive-adaptive perspective on E.
biased accordingly. Multimethod assessment including objec- The primary trait adaptation is to demanding social encounters.
tive tests provides a possible solution. From the causal perspec- In addition to skills for social interaction, such as uency of
tive, CATT attributes observed associations between traits and expression, persuasiveness, and nonverbal expression (e.g.,
behaviors to multiple processes, with different processes playing Kluemper, McLarty, Bishop, & Sen, 2015), extraverts also
a mediating role in different contexts. For some research ques- require capabilities for tolerance of social stress, such as apprais-
tions, we can treat the trait as a unitary construct, whereas others ing situational demands as challenges rather than threats, and
may require discriminating the separate component processes. preferring task-focused coping (Carver & Scheier, 2001). Multi-
For example, C may be associated with neural and information- ple information-processing and neural correlates of traits, on the
processing functions for effortful control, positive affect in per- left of Figure 1, support acquisition of these skills. Research on
formance settings, skills for time management and task comple- the cognitive patterning of E (Matthews, 2008a; Matthews &
tion, and self-efcacy for task performance (Matthews & Zeidner, 2012) identies multiple basic processing correlates of
Zeidner, 2012; R. Roberts et al., 2015). With a broad focus on E, including superior divided attention under high processing
job performance, we can treat these different attributes as a pack- load, inhibition of distractor stimuli, and verbal uency. These
age, identied with C as a causal inuence on success. A more processing attributes, predominantly set during childhood
granular approach is to discriminate the different attributes as through genetic and environmental inuences, facilitate learning
causal inuences on behavior, rather than the trait itself. social skills such as keeping track of multiple speakers in a con-
The theory recognizes two classes of inuence on adaptive versation, ignoring irrelevant speech, and intervening in ongoing
behaviors that mediate trait effects: cognitive skills and self- conversations. Processing attributes contribute indirectly rather
knowledge. Skills are dened in cognitive models such as that of than directly to adaptation: The skills and self-regulative pro-
Anderson (2007). For example, a car sales agent will have a cesses that they support are more important for individual differ-
standard script to encourage positive attitudes in potential cus- ences in adaptation. Biologically based trait correlates such as
tomers that can be modied to deal with a range of types of car arousability similarly inuence skill acquisition indirectly; for
buyers with only limited conscious thought. Personality traits example, overarousal in social settings limits social skill devel-
are related to cognitive skills such as time management (C; Mac- opment. Thus, CATT is a variant of investment theories (Acker-
Cann, Fogarty, & Roberts, 2012) and information seeking (O; man, 2014; Hudson et al., 2012). Consistent with traditional
cf. Goff & Ackerman, 1992), as well as cognitive-social skills thinking on traits, basic cognitive and neural inuences are
such as interacting with strangers (E), detecting possible malice determined early in life, and they remain quite stable in
or criticism (N), and seeing someone elses perspective (A). adulthood.
Use of skills is accompanied by self-regulation based on sta- The more malleable aspects of personality are represented by
ble self-knowledge (explicit and implicit). Especially in chal- the adaptive triangle shown to the right of Figure 1. Cognitive
lenging environments, the person needs to monitor the skills, self-knowledge, and contextualized behaviors interact
effectiveness of skill execution and process feedback, set appro- continuously. Adaptation operates over different timescales:
priate goals, and determine whether additional effort is needed The model shown in Figure 1 concerns typical outcomes of typi-
to compensate for lack of skill. Self-regulation is also intimately cal behaviors, such as the extravert repeatedly interacting with
related to emotional response, which may be positive (e.g., pride others at parties. High levels of expertise support greater benets
in accomplishment) or negative (e.g., sadness if goals are not when the skill is exercised, leading to increased opportunities to
6 Matthews

Figure 1 A cognitive-adaptive model of extraversion.

practice the skill (e.g., extraverts receive more party invitations) familiar people, (d) possibly benecial, and (e) unable to be
and further skill improvement. Self-regulative factors such as addressed by a single coping strategy (Matthews, 2004). For
self-efcacy may also motivate the person to choose activities in example, a person in a troubled intimate relationship may have
which the skill is required (e.g., extraverts seek out parties), lead- to work to decode mixed signals coming from the other person,
ing to greater expertise, practice, and self-condence. Con- as well as use a variety of coping strategies to repair the relation-
versely, a typical adaptation for introverts is working in the ship or to leave it with hope for the future. Faced with this com-
absence of immediate reward or help from others, supported by plex problem-solving task, the typical high N strategy is to
skills for sustaining attention, reective problem solving, and remain vigilant for possible threats and to act to anticipate and
boredom tolerance (Matthews, 2008a). avoid possible threats. Hence, N is associated with sensitivity to
Often, social skills and condence will align, but individuals possible criticism and a preference for avoidant coping (Mat-
show a range of adaptive patterns. The extravert with high con- thews, 2004). Such a strategy may be adaptive in picking up
dence but low actual skill risks becoming a saloon-bar bore; lim- early signs that an intimate relationship is in trouble. Fear of
itations in social perception may limit the impact of negative social disapproval may also be motivating and associated with
feedback from others on self-concept. Adaptation may also high levels of preparation and contribution to tasks performed at
change over time. Bendersky and Shah (2013) discuss how, in work (Smillie, Yeo, Furnham, & Jackson, 2006; Tamir, 2005).
business team contexts, the condence and enthusiasm of extra- However, such strategies also carry potential costs of overesti-
verts (reecting self-regulation) is initially perceived positively mating threat and excessive stress response, worry, and rumina-
by other team members. Over time, extraverts contributions to tion (Wells & Matthews, 2015). Conversely, the characteristic
the groups tasks may fail to match initial expectations, leading low N strategy is to wait for a clear threat to become evident and
to a loss of status within the group. then to deal with it directly (e.g., through task-focused coping).
Variation in N reects alternate strategies for handling threat Maladaptive outcomes associated with high N including
(see Figure 2). N especially moderates response to social threat emotional disorder may reect dynamic adaptive processes,
(e.g., Brock & Lawrence, 2014), a type of stressor that poses including dysfunctional self-regulation. Social anxiety is associ-
unique challenges. Social threats are often (a) extended over ated with avoidance of feared situations, leading to skill degrada-
time, (b) ambiguous or disguised in nature, (c) associated with tion and irrational negative beliefs about social encounters (Wells
Cognitive-Adaptive Theory 7

Figure 2 A cognitive-adaptive model of neuroticism.

& Matthews, 2015). The high N individual is vulnerable to styles knowledge, and adaptation that constitute the adaptive triangle.
of metacognition that focus attention on negative thoughts, lead- By contrast with FFT (McCrae & Costa, 2008), traits are associ-
ing to perseverative worry that keeps social threats salient, limits ated with characteristic adaptations, not basic tendencies. Traits
constructive reappraisal of the threat, and blocks exposures to the are indirectly, not directly, related to brain systems. Even the
threat that would allow acquisition of compensatory social skills. more reliable associations between traits and metrics of neural
Bendersky and Shah (2013) describe a more benign dynamic in functioning tend to be small in magnitude (Matthews &
the business context. High N individuals do not inspire condence Amelang, 1993; Matthews & Gilliland, 1999). Conversely,
in others and tend to be assigned lower status in work groups. associations between traits and self-regulative constructs, such
However, because it is relatively easy for them to exceed expecta- as self-beliefs and styles of appraisal and coping, are often quite
tions, they gain status within the group over time. large, as illustrated by the convergence of low N with other ele-
ments of core self-regulation (Ferris, Johnson, Rosen, & Tan,
2012).
Personality From the Cognitive-Adaptive
Second, while traditional psychobiological theory favors
Perspective establishing isomorphisms between major traits and specic
Historically, impetus for psychometric studies of personality brain systems, such as reward and punishment systems (Corr,
came from the example of the periodic table of the elements 2009), CATT rejects isomorphism. Mappings between brain
(Mershon & Gorsuch, 1988). That is, there is a precise taxon- attributes and traits may be complex; for example, one attribute
omy of personality dimensions that provides a universal frame may contribute to multiple traits (Zuckerman, 2005).
of reference for all traits. In addition, the periodic table metaphor Third, the isomorphism assumption encourages the search
suggests that each trait corresponds to a specic property of the for a master-process for each trait that explains the majority of
brain, as each element is dened by atomic number. CATT observed trait expression. In H. J. Eysencks (1967) theory,
departs from this metaphor in several respects. individual differences in cortical arousability inuence condi-
First, standard personality dimensions, including the Big tioning, task performance, and regulation of level of stimulation,
Five, are associated with the interlocking patterns of skills, self- potentially explaining numerous expressions of extraversion-
8 Matthews

Table 1 Comparison of Principles of Standard Trait Theories With Cognitive-Adaptive Theory

Type of Issue Standard Trait Theory Cognitive-Adaptive Theory


Psychometric Psychometrics defines a universal set of five (or Superfactor assessments are essential research tools,
more) superfactors. but no precise universal model can be defined.
Universal superfactors are more fundamental than Universal and contextualized traits are equally
contextualized traits. important for adaptation but differ in scope.
Trait and state dimensions correspond directly. Trait and state dimensions reflect different sets of
adaptive challenges and are not directly
commensurate.
Theoretical Trait superfactors correspond directly to brain Neural influences on traits are indirect; trait struc-
systems that are the source of trait coherence. ture corresponds to key adaptive strategy variation.
Trait coherence derives from common functionality
of multiple processes.
Biased self-perceptions are peripheral to trait Biased self-perceptions are one of several aspects of
definition. self-knowledge contributing to traits.
Cognitive correlates of traits are secondary to neural Cognitive correlates of traits provide a platform for
processes or are learned characteristic adaptations. acquiring the skills characteristic of the trait.
Adaptive/ Applied Trait stability reflects brain stability; characteristic Trait stability reflects multiple processes, including
adaptations are more malleable. skill and self-knowledge generalization across life
transitions, and personsituation interactions.
Adaptive outcomes reflect feedforward from basic Adaptive outcomes reflect levels of contextualized
brain systems to learning and performance. skills, functionality of self-knowledge in context,
and level of exposure to relevant environments.

introversion. By contrast, supported by evidence on cognitive Affect, and Intellect, but in each case the nature of the chal-
patterning (Matthews, 2008a), CATT assumes that traits are dis- lenge requires considerable further denition. For example,
tributed across multiple processes to be understood at different exerting power is relatively straightforward if one arrives at a
levels of explanation (Matthews, 2000). Thus, mediating mecha- leadership role with levers of inuence already at ones dis-
nisms for trait effects vary according to the context and the task posal (Bill Gates and Mark Zuckerberg are reputed to be
performed. introverted; Bernstein, 2015). Doing so under social pressure
Fourth, what gives traits coherence is not a single master- is difcult, and more closely linked to E. Traits reect adapta-
process, but their adaptive focus. E is correlated with numerous pro- tions to environmental challenges, but environments are noto-
cesses, often with small magnitude, but they all serve a common riously difcult to dene and categorize. Indeed, even core
goal of facilitating adaptation to demanding social environments. environmental challenges vary somewhat across cultures and
This perspective on traits provides novel answers to long- across generations. Consequently, traits are necessarily some-
standing issues in personality theory that differ in some respects what fuzzy in nature, and we should not expect a precise uni-
from traditional trait theory perspectives. Table 1 summarizes versal structural model. Indeed, as H. J. Eysenck (1967)
the key differences. Psychometrically, CATT questions whether recognized, arguments over trait denition cannot be resolved
there will ever be a truly universal trait model, given that even by psychometrics alone.
the central adaptive challenges for humanity vary at least some- Universality of trait structure speaks to a common human
what across cultures, contexts, and generations. The theory also nature, variation to the moderation of adaptive challenges by
rejects the idea of well-dened brain systems constituting the culture. However, the FFM shows some variation across cul-
basis for each trait; instead, trait coherence reects the contribu- tures, especially non-Western and/or nonindustrialized cultures
(Church, 2016). For example, Gurven, Von Rueden,
tion of multiple, dissociable processes to adaptation. Finally, in
Massenkoff, Kaplan, and Lero Vie (2013) failed to reproduce
real life, the consequences of traits depend primarily on acquired
the standard FFM in the indigenous, largely illiterate, Tsimane
skills and self-knowledge. Next, these differences in perspective
foragerhorticulturalist people of Bolivia. They attributed the
from standard models are discussed in more detail.
cultural difference in personality structure to the distinctive
social environments of small-scale societies. For example, artis-
tic interests are expressed in a communal setting so that there is
Psychometric Uncertainties more intercorrelation of E and O items than in industrialized
Psychometric studies of the FFM do not identify ve key populations. Gurven et al. (2013) also suggest that Westernized
brain attributes, but ve prototypical environmental chal- cultures may have more complex factor structures than prein-
lenges that support contrasting adaptive strategies. Goldberg dustrial ones because there are more environmental niches
(1990) dened these challenges as Power, Love, Work, available.
Cognitive-Adaptive Theory 9

Additional Superfactors. From the cognitive-adaptive per- Theoretical Issues: Mediators of Trait Effects
spective, whether there are ve, six, or more superfactor traits is Mismatch of Constructs and Assessments. Using self-
not critical. A metaphor is the question of how many planets reports to investigate individual differences in neural processes
there are in the solar system. There is a nite number of major
is problematic. Thus, we could measure traits objectively,
planets, which is important for astronomers to dene, but that
through psychophysiology, and explore their antecedents and
number reects the contingent processes guiding planetary for-
consequences. Personality theorists have been curiously reluc-
mation rather than any basic law of physics or chemistry. Simi-
tant to adopt this approach. In child temperament research espe-
larly, denition of what is a major or minor trait remains
cially, there are well-established associations between traits
somewhat arbitrary. The controversy over whether Honesty in
dened through behavioral observations and psychophysiologi-
the HEXACO model is a major Big Six trait dimension
cal measures (e.g., Kagan, 2014; Rothbart et al., 2009). How-
(Ashton et al., 2014; McCrae & Costa, 2008) is then equivalent
ever, using the behavioral measures to predict physiological
to the debate over whether Pluto is a planet. It is more important
response does not directly address the extent to which tempera-
to determine whether Honesty has antecedents and conse-
ment can be directly explored through psychophysiology.
quences that differentiate this trait from the Big Five in theoreti-
One of the challenges here is that individual differences in psy-
cally interesting or practically important ways. Generally, there
chophysiological metrics do not seem to support a superfactor
remain signicant psychometric issues surrounding structural
structure akin to the Big Five, as FFT (McCrae et al., 2000) pre-
models of personality, including the FFM (Boyle, 2008), and
dicts. In a recent study of individual differences in workload
psychometrics alone may not lead directly to causal models of
trait action. response, Matthews, Reinerman-Jones, Barber, and Abich (2015)
assessed EEG, ECG, eye tracking, and hemodynamic responses
Contextualized Traits. The FFM represents the major chal- to multiple task scenarios in 151 participants. Factors could be
lenges of being human (Goldberg, 1990). However, if traits cor- dened within specic response systems (e.g., EEG) but not
respond to environmental challenges, there should also be traits across systems. In other words, individual differences in brain
for more specialized environments. Thus, the existence of con- functioning may have numerous degrees of freedom that do not
textualized traits falls naturally out of the cognitive-adaptive map readily onto extant personality structures. Research on bio-
approach. For example, in developed cultures, formalized test- logical bases for personality might focus more productively on
ing of students is important enough that different strategies for more narrowly dened endophenotypes, consistent with the
handling the pressures of being evaluated will exist, dening the molecular genetic evidence that traits are inuenced by many
test anxiety trait (Zeidner, 1998). The general social threat linked genes of small effect (Montag & Reuter, 2014). Improvements in
to N overlaps with more narrowly dened evaluation threats, brain-recording technology may provide better denition, but for
and so the traits correlate while remaining distinct constructs. In now there is much still to be learned about dimensions of brain
evaluative contexts, traits such as test anxiety and math anxiety functioning and how they are expressed in behavior. From the
are more predictive of performance than broad traits (Zeidner, cognitive-adaptive perspective, brain characteristics (and genes)
1998). The structure of contextualized traits differs from the are all upstream from the core constructs of personality, which are
FFM because environmental pressures do not align with those more tractable psychometrically and theoretically.
of life in general. For example, aggressiveness is more promi-
nent as a driver personality dimension than E because trafc The Role of Self-Perceptions. CATT accommodates a role
environments limit the range of social responses (Matthews, for biased self-perceptions in inuencing personality. Stable
2002). They provide more scope for frustration and conict self-beliefs are encoded in memory during childhood and form
between vehicles than opportunities for social inuence, which part of the self-schema in adults (Clark & Beck, 1999). There is
typically require people to converse. extensive evidence for cognitive bias and distortion in evaluat-
ing the personalities of others (e.g., Denrell, 2005) and of the
Differentiating Traits and States. A direct correspondence self, especially in the emotional disorder literature. Clark and
between trait and state dimensions is often assumed, whether on Becks (1999) classic studies show low self-esteem in depres-
theoretical grounds or through development of parallel scales sion is linked to systematic errors in thinking. Within CATT,
(Spielberger & Reheiser, 2009). CATT rejects this view because such biases are a part of self-regulation and may feed styles of
short-term adaptive challenges of goal-directed behavior do not processing that shape adaptation, such as self-verication and
directly map onto the longer-term challenges associated with self-enhancement (Robinson & Sedikides, 2009). Thus, the the-
trait variation (Matthews & Zeidner, 2004). In the performance ory sees self-perception biases as an integral part of the major
context, for example, major state factors are associated with traits, with the power to shape behaviors such as overcondent
commitment of effort (task engagement), managing overload engagement in risky behaviors (Wohleber & Matthews, 2016).
(distress), and reection on the personal relevance of the task Biases are thus more than nuisance response styles, consistent
(worry). These dimensions correlate modestly with traits such as with evidence that variance in social desirability is actually
C and N in predictable ways, but they do not directly align with associated with substantive personality traits (Connelly &
traits (Matthews et al., 2002). Chang, 2016).
10 Matthews

Specifying Characteristic Adaptations. The multiplicity of attention, via performance metrics. Both C and E were associ-
psychological constructs that may mediate trait effects is chal- ated with superior executive functioning, but seemingly for dif-
lenging, and there is no simple resolution to the issue. However, ferent reasons. The effect of C was mediated by the subjective
the cognitive-adaptive perspective may contribute to conceptual state of task engagement, which is associated with challenge
clarity, especially in distinguishing objectively measurable skills appraisal and task-focused coping in performance settings. Fur-
that are associated with the trait from self-regulative constructs. ther investigation might require an analysis at the self-
A limitation of much contemporary personality research (includ- knowledge level to understand how C inuences understanding
ing some of this authors studies) is the ambiguity of subjective of the personal relevance of the task and willingness to invest
measures of mediating constructs. For example, individual dif- effort in it. By contrast, E was unrelated to task engagement, and
ferences in coping sometimes mediate trait effects on well-being the effect of E on executive function was not mediated by indi-
(Matthews et al., 2009). Task-focused coping is associated with vidual differences in subjective state. In this case, we might
more positive outcomes (Matthews et al., 2009), but does that probe individual differences in the parameters of neural or cog-
mean that people use task focus more often, that they are better nitive architecture supporting executive control of attention. In
at implementing task-focused strategies, that they gain con- an applied context, such as individual differences in resistance
dence from believing they have a plan of action, or that they to distraction in social settings, we might also be concerned with
encounter more situations for which task focus is appropriate? contextualized skills for directing attention.
The theory highlights the need for greater clarity of thought
about explicit and implicit aspects of personality. Both skills and
self-regulation have conscious and unconscious elements. Individual Differences in Adaptation: Traits in
A cognitive-adaptive perspective separates different explana- Real Life
tions. In a recent study, Zeidner and Matthews (2016) showed
Stable and Malleable Elements of Personality. Traditional
that perceived social support mediates the association between
trait theory (e.g., McCrae & Costa, 2008) discriminates stable
emotional intelligence and well-being. However, there are sev-
and malleable elements fairly sharply. Basic tendencies,
eral alternate, though potentially complementary, explanations
for this nding. Perceiving oneself as socially supported (self- grounded in neural processes, show similar stability to the brain.
regulation) may elevate well-being in the absence of any objec- Following brain maturation, further changes will be relatively
tive benet. The individuals skills in securing and gaining from small in magnitude. By contrast, characteristic adaptations are
social support (skill) may improve well-being via better objec- learned and may change with external circumstances.
tive outcomes. Perceived social support may signal ones partic- CATT is in agreement with traditional theory on the relative
ipation in a mutually supportive network (dynamic adaptation). stability of neural processes but attributes less of adult trait sta-
These different support processes may well coexist, but they can bility to direct effects of brain systems on personality, given that
be distinguished and tested empirically. traits are an indirect rather than a direct expression of individual
differences in brain functioning. Instead, the theory attributes
Accommodation of Cognitive Processes. CATT was stability to several separate factors, which might be dissociated.
developed initially (Matthews, 1999) to accommodate the diver- Stability of basic neural and information-processing characteris-
sity of cognitive correlates of traits revealed in information- tics will nudge adaptation in similar directions in different con-
processing studies, along with the modest magnitude and con- texts. For example, biologically based stress vulnerability will
text sensitivity of traitperformance associations. The theory encourage the high N adaptive strategy across multiple types of
proposes that contextualized skills, such as social skills, may be threat.
more central to traits than basic information-processing compo- Skills and self-knowledge also contribute to personality
nents, such as those for speeded response, attention, and mem- change and stability, in complex ways. The key issue for skills is
ory. Like brain characteristics, their role in trait denition may how much they generalize to new contexts as the persons life
be indirect rather than direct. A chastening feature of the theory circumstances change, including taking on new social roles
is that it emphasizes the complexity of associations between per- (Hudson et al., 2012). Thus, stability of external circumstances
sonality and cognitive processes, including the necessity of is an inuence on stability of traits. The level of generalization,
referring to different explanatory levels, ranging from basic neu- and hence the extent to which trait stability is promoted, depends
roscience to high-level strategizing shaped by self-knowledge. on the various factors that control skill transfer (Anderson,
These different types of level can be formally distinguished 2007), including variability in environmental exposures or
within cognitive science theory (Matthews, 2000). Researchers choices over time. Internalized self-knowledge may also be a
naturally favor parsimony and elegance in theory, but we have source of either stability or change depending on whether self-
to live with a messier reality. regulatory processes maintain a stable self-concept in changing
For example, Matthews and Zeidner (2012) investigated trait circumstances or promote personal reinvention. Constructs such
and state correlates of Fan, McCandliss, Sommer, Raz, and as narrative identity capture longer-term processes of this kind,
Posners (2002) Attention Network Test, which assesses indi- including idiographic elements (Cervone, 2008; McAdams &
vidual differences in functioning of three brain networks for Pals, 2006).
Cognitive-Adaptive Theory 11

Stability also reects the various forms of dynamic person are changeable (Sackett & Walmsley, 2014). Individuals can be
situation interaction (Caspi & Roberts, 2001). For example, high trained to use day planners, checklists, and other aids supporting
N may be maintained by awareness of social threat (reactive personal organization even if they are low in C. Such interven-
interaction), by styles of interpersonal interaction that elicit neg- tions would likely enhance a contextualized measure of work-
ative emotions in others (evocative interaction), and by associat- place C more than the general trait. Training also moderates the
ing with or valuing like-minded others (proactive interaction). predictive validity of traits. For example, one factor in the sur-
Thus, CATT largely aligns with more dynamic perspectives on prisingly weak correlation between E and sales performance is
trait development (e.g., Hudson et al., 2012). Where it may add that managers train introverted sales staff in the contextual skills
novelty is in separating skills from self-regulation. For example, required (Grant, 2013). According to the present analysis, inter-
reactive interaction in highly neurotic individuals might be a ventions that change skills important across multiple contexts
product of a learned style of stimulus encoding that is prone to will inuence traits. Training in stress management techniques
categorize events as threats, inuenced by selective attention that can be applied in a variety of work and nonwork contexts
bias. It might also reect a self-regulative style that dictates the might reduce N, consistent with evidence that cognitive-
importance of active search for possible threats and personal behavioral interventions for anxiety can lower N (Barlow,
vulnerability. Both types of processes appear to be implicated in Sauer-Zavala, Carl, Bullis, & Ellard, 2014). Further research is
emotional disorders (Wells & Matthews, 2015). The role of needed to substantiate this hypothesis (e.g., B. W. Roberts et al.,
skills highlights the importance of habitual styles of information 2017).
processing, of which the person may have limited awareness,
whereas the role of self-knowledge implicates self-regulative
processes, including self-verication (Tamir, 2005). CONCLUSIONS
Modern trait theory provides a set of answers to traditional ques-
Context and Skills. By contrast with traditional trait perspec-
tions that are in many ways satisfactory. We can measure mean-
tives, individual differences in declarative and procedural skills
ingful and stable individual differences, establish mappings
(Anderson, 2007) acquired within a specic context are central
between traits and biological and psychological attributes, and
to traits in CATT (Matthews, 1999). Thus, individual differ-
use trait assessments to support decision making in various
ences in contextualized skill will often be more important as
applied settings. CATT (Matthews, 1999, 2008a) seeks to
mediators of trait expression than basic neural or cognitive pro-
cesses. The common advantage of both multiple narrowly address some of the remaining issues. In doing so, it contrasts
dened scales and contextualized measures over general trait with theories that dene explicit biological bases for traits (Corr,
measures in predictive validity (Boyle, 2008; Matthews et al., 2009; Eysenck & Eysenck, 1985) and FFT (McCrae & Costa,
2009) reects the context-bound nature of many skills, although 2008). In some instances, theoretical differences are nuanced,
some key social skills, such as emotional expressivity and clarity for example, in dening which aspects of personality are mallea-
of communication, will generalize across numerous contexts. ble. Other differences are more fundamental. In particular,
Furthermore, the more specialized the environment, the weaker CATT rejects the view that traits can be identied with any sin-
the validity of standard traits will be. Vehicle driving, for exam- gle or small number of causal processes that mediate associa-
ple, is a microworld that is distinct from other everyday life set- tions between traits and behavior, including individual
tings. Standard traits such as those in the FFM have real but differences in brain systems for arousal or motivation. Traits are
modest validity for predicting adaptive criteria such as risky distributed across numerous processes at different levels of
behavior and stress response, whereas scales specialized for the abstraction from the neural substrate, and they gain coherence
driving context perform better (Matthews, 2002). from their adaptive function, not from isomorphism with any
A skill-centered perspective on traits also suggests greater underlying structure or process.
scope for interventions in applied settings than standard trait the- Finally, CATT suggests the limits of natural science meta-
ory (e.g., FFT) does. R. Roberts et al. (2015) link the FFM to phors for personality. It is appealing to think of traits as xed,
skills that can be trained to increase educational potential and essential characteristics of the person akin to height and blood
workforce readiness. For example, Durlak, Weissberg, Dym- type. From the cognitive-adaptive perspective, the essentialist
nicki, Taylor, and Schellinger (2011) demonstrated in a meta- position is often useful as a working approximation, given that
analysis that school-based social and emotional learning (SEL) traits are fairly stable and fairly universal. However, this position
programs are effective in various respects, including improving ceases to be useful when we address several specic research
tolerance of academic stressors (associated with N), setting questions, including the relationship between traits and brain
appropriate academic goals (C), and resolving conicts with characteristics, the role of cognitive processes in expressions of
peers and teachers (A). Importantly, such interventions typically personality, and the scope for personality modication in
do not seek to change personality in some global sense but to applied settings. Instead, future research should focus more on
develop skills that are relevant to the students circumstances. the role of individual differences in generalizable skills, includ-
In the workplace, underlying dispositions are fairly stable, ing those for building interpersonal relationships, interpreting
but the patterns of behavior reecting personality characteristics ones place in the world, acquiring new information, and
12 Matthews

navigating the world of work. Key questions for the future are & A. Thapar (Eds.), Rutters child and adolescent psychiatry (5th
the role of basic neural and cognitive processes in skill acquisi- ed., pp. 182199). London: Blackwell.
tion, the role of self-knowledge in supporting acquisition and Cervone, D. (2008). Explanatory models of personality: Social-
learning of skills, and the dependence of various forms of per- cognitive theories and the knowledge and appraisal model of per-
sonsituation interaction on individual differences in skill. sonality architecture. In G. J. Boyle, G. Matthews, & D. H.
Saklofske (Eds.), The Sage handbook of personality theory and
testing: Volume 1. Personality theories and models (pp. 80101).
Declaration of Conflicting Interests Thousand Oaks, CA: Sage.
Chang, L., Connelly, B. S., & Geeza, A. A. (2012). Separating
The author declared no potential conicts of interest with respect method factors and higher order traits of the Big Five: A meta-
to the research, authorship, and/or publication of this article. analytic multitraitmultimethod approach. Journal of Personality
and Social Psychology, 102, 408426.
Church, A. T. (2016). Personality traits across cultures. Current
Funding Opinion in Psychology, 8, 2230.
The author received no nancial support for the research, Cisler, J. M., & Koster, E. H. (2010). Mechanisms of attentional
authorship, and/or publication of this article. biases towards threat in anxiety disorders: An integrative review.
Clinical Psychology Review, 30, 203216.
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