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9781605351155
July 2013 | Hardback
656pp | £46.99
Request an inspection copy today at
www.palgrave.com/Futuyma3e
Evolution, Third Edition
– a comprehensive exploration of contemporary evolutionary biology
Thoroughly updated, Evolution, Third Edition, addresses major themes — including
the history of evolution, evolutionary processes, adaptation, and evolution as an
explanatory framework — at levels of biological organization ranging from genomes
to ecological communities. Throughout, the text emphasizes the interplay between
theory and empirical tests of hypotheses, to help students become more familiar
with the process of science. With a helpful list of important concepts and terms in
each chapter, dynamic figures and lively photographs the new edition of Evolution
will inspire and inform the next generation.
About the Author
Douglas J. Futuyma is Distinguished Professor in the Department of Ecology and Evolution at the
State University of New York at Stony Brook. He is the author of two previous editions of Evolution,
as well as three editions of its predecessor, Evolutionary Biology, and has authored or edited several
other books. Dr. Futuyma has received the 1997 Sewall Wright Award of the American Society of
Naturalists and the 2012 Joseph Leidy Award of the Academy of Natural Sciences of Drexel University
(Philadelphia). He has served as Editor of Evolution and is currently Editor of the Annual Review of
Ecology, Evolution, and Systematics. His major research interests include evolution of host specificity
in herbivorous insects and of interactions among insects and plants, speciation, and evolution of
community structure.
What’s New?

• Greatly revised treatment of methods of phylogenetic analysis

Chapter 4 includes updated material of feathered
• Up-to-date coverage of human evolution, based on the fossil record and genomic dinosaurs and the origin of birds, including Figure
variation 4.9 which references papers from 2010 and 2012

• Reorganized treatment of the history of changes in biological diversity, integrating

information from palaeontology and phylogenetics
• Coverage of important new developments in sexual selection and the evolutionary
advantage of sex
• Updated and expanded coverage of reinforcement in speciation, and of speciation with
gene flow – View Chapter 18: Speciation today.
• Extensive updating of very recent advances in evolutionary genomics
• Approximately 25% new figures
• 460 new citations (of approximately 1,700)

For more information, explore the

Chapter 9 includes new and updated material
Chapter-by-Chapter Guide to New Content… on nongenetic inherited variation, linkage
disequilibrium and patterns of human genetic
variation, in relation to notions of “race”
A Chapter-by-Chapter Guide to New Content

1. Evolutionary Biology 6. The Geography of Evolution 10. Genetic Drift: Evolution at Random
• Added example of hypothetico-deductive method • A description of new methods of historical • Expanded introduction to random processes in evolution
(prediction and verification) in evolutionary biology biogeographic analysis • Updated material on reduced variation caused by
• New material on the spread of Homo sapiens beyond population bottlenecks
Africa (including admixture of Neanderthal and • New, updated section on molecular clocks
2. The Tree of Life: Classification and Phylogeny Denisovan genes) • Rewritten and updated material on the origin of modern
• Revised, updated discussion of statistical methods of Homo sapiens, human effective population size, and
phylogenetic analysis genetic correlates of explosive population growth
• New box describing additional phylogenetic methods 7. The Evolution of Biodiversity
• Updated discussion of incomplete lineage sorting • Improved integration of paleontological and
• New section on applications and extensions of phylogenetic approaches to estimating clade 11. Natural Selection and Adaptation
phylogenetics diversity through time • Added discussion of selection sorting in both time
• Updated material on patterns of temporal change and space
in clade diversity • New material on effective environment depending
3. Patterns of Evolution • Updated discussion of diversity-dependent on species
• New material on molecular convergence diversification • Expanded treatment of testing hypotheses of
• Updated discussion of comparative genome sizes • New section on why some kinds of organisms adaptation by experiments
and numbers of genes in genome are more diverse • Material on adaptive evolution moved to this
• Updated and expanded section on adaptive radiation chapter and expanded
• New section on adaptation to global warming
4. Evolution in the Fossil Record
• Updated material on feathered dinosaurs and the 8. The Origin of Genetic Variation
origin of birds • Updated discussion of effects of regulatory mutations 12. The Genetic Theory of Natural Selection
• Updated information about the Hominin fossil record, • New material on molecular causes of pleiotropy • Updated examples of gene flow vs. selection, the
new genomic data, and human origins • Updated material on fitness effects of mutation in effect of gene flow on local adaptation, and
noncoding regions of the genome, and on the frequency-dependent selection
frequency of neutral and nonneutral mutations • Updated treatment of a classic example: industrial
5. A History of Life on Earth • New description of epistatic mutations melanism in the peppered moth
Updated references and information on: the origin of • New material on detecting selection from geographic
life/earliest life; laboratory evolution of RNA mixtures; variation
the evolution of multicellularity; comparative genetics 9. Variation: The Foundation of Evolution • Updated treatment of selection in the human genome
of choanoflagellates and animals; age of animal • New section on nongenetic inherited variation • New section dealing with adaptation based on new vs.
phyla; the evolution of angiosperms; phylogeny of • Recast material on linkage disequilibrium standing variation
vertebrate groups; early mammalian diversity; causes • Updated references on patterns of human genetic • McDonald-Kreitman test for selection moved from
of Pleistocene megafaunal extinction variation, in relation to notions of “race” Chapter 10
13. Phenotypic Evolution 17. Species 21. Evolution and Development
• Substantial rewriting and shortening of some sections • Expanded treatment of phylogenetic species concepts Contributed by John R. True
• Updated material on QTL for human traits • Updated discussion of ecological isolation, hybrid • Updated coverage of the evolutionary developmental
• New treatment of directional selection in inviability, and the genetics of behavioural isolation biology of flower development
contemporary human populations • Expanded treatment of genetic incompatibility based • New material on the evolutionary developmental
• New treatment of evolution of phenotypic plasticity on duplicate genes biology of flower symmetry
• Updates on genetic constraints, modularity, and • Revised section on genetic divergence and exchange • Updated section on genome-level and pathway
evolution of pleiotropy perspectives of developmental evolution
• New section on whether adaptation can rescue species • Updated discussions of protein sequence changes in
from extinction 18. Speciation regulatory and morphological evolution
• Expanded section on evidence for allopatric speciation • New perspectives on macroevolution and the role of
• Updated discussion of ecological speciation developmental arrest in morphological evolution
14. The Evolution of Life Histories • Expanded treatments of sexual selection and • New material on the integration of ecological
• Rewritten section on age schedules of reproduction speciation and of peripatric speciation developmental biology
• Section on “Life Histories and Mating Strategies” • Revised section on sympatric speciation; new material
moved to this chapter from Chapter 15 on genomic studies of speciation with gene flow
22. Macroevolution: Evolution above the
Species Level
15. Sex and Reproductive Success 19. Coevolution: Evolving Interactions • Updated treatment of stasis, gradualism and saltation,
• Updated material on the costs of sex, clonal among Species and the evolution of complex characteristics
interference, sex and parasites • Revised section on aposematism and mimicry • New material on developmental biology and
• Treatment of the newest models of the advantage of • Expanded and reorganized section on parasite–host complexity
sex and recombination (separating advantageous interactions and infectious disease • Revised section on evolvability
alleles from disadvantageous alleles elsewhere in • Revised material on evolution of competitive • New section on predictability and contingency in
the genome) interactions evolution
• Reorganized material on the evolution of mate choice • New section on evolution and community structure
• New sections on reversed and mutual sexual selection,
and on the effects of sexual selection on adaptation 23. Evolutionary Science and Creationism
• Updated treatment of sexual conflict 20. Evolution of Genes and Genomes • Updated discussion of evolution in relation to health
Contributed by Scott V. Edwards and medicine
• New section on diverse players and processes in • New material on evolutionary approaches to pest
16. Conflict and Cooperation genomes management
• Reorganized discussion of social interactions and • New information on the fraction of the genome
cooperation that is functional
• Expanded material on the direct benefits of cooperation • Discussion of developmental processes and rates of
• New section on the evolution of spite protein evolution For more details visit
• Updated and reorganized sections on social insects • Discussion of what newly sequenced genomes reveal
and on kin selection vs. group selection about major events in the history of life www.palgrave.com/Futuyma3e
• Expanded treatment of evolutionary approaches to • Revised material on molecular convergence as to download the complete chapter
human behaviour evidence for natural selection
by chapter guide.
Media and Supplements

For the Lecturer

Instructor’s Resource Library
The Instructor’s Resource Library includes a variety of resources to help you develop your course
and deliver your lectures, including:
• Textbook Figures and Tables: All the figures, photographs and tables from the textbook are
provided as JPEGs (both high- and low-resolution).
• PowerPoint Presentations: For each chapter, all of the chapter’s figures and tables are provided
in a ready-to-use PowerPoint presentation, making it easy to quickly insert figures into your own
Online Quizzing
lecture presentations. A set of online quizzes is available via the
Companion Website. These can be assigned or
• Answers to the textbook end-of-chapter Problems and Discussion Topics
released for student self-study and lecturers can
• Quiz Questions from the Companion Website also add their own questions to the quizzing system,
to create custom quizzes. The results can be viewed
• Data Analysis and Simulation Exercises from the Companion Website, with answers
online or downloaded. (Lecturer registration is
required for student access to the quizzes.)

For the Student

Companion Website
The Companion Website features review and study tools to help students master the material
presented in the textbook, including:
• Chapter Outlines and Summaries: Concise overviews of the important topics covered in
each chapter.
• Data Analysis Exercises: Expanded for the Third Edition, these inquiry-based exercises involve
students in working with data and analysing methods and conclusions from published papers.
• Simulation Exercises: Interactive modules that allow students to explore many of the
dynamic processes of evolution, and answer questions based on the results they observe.
• Online Quizzes: Assignable quizzes that cover all the major concepts introduced in each
chapter.
• Flashcards & Key Terms: Easy-to-use activities that help students learn all the key
terminology introduced in each chapter.
Example Simulation available on the Companion Website
• The complete Glossary
 July 2013 | Hardback
656pp | £46.99
9781605351155

Request your inspection copy online at

www.palgrave.com/Futuyma3e or contact
Lecturer Services on lecturerservices@palgrave.com
or +44 (0)1256 302794

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