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Introduction
An ecological succession is the replacement of populations in habitat as it moves toward
a stable state. The progress of a succession is determined by changes in plant number
and species over time.
The environment may be altered in substantial ways through the activities of organisms,
including humans, or when the climate changes. Although these alterations are
sometimes abrupt (ex. natural disasters), in most cases species replace others, resulting in
long-term gradual changes in ecosystems.
Ecosystems tend to change with time until a stable system is formed. The type of
ecosystem that is formed depends on the soil or climatic limitations of a given
geographical area.
Pioneer organisms are the first organisms to establish themselves in a succession. These
organisms could include lichens on bare rock or annual grasses on an abandoned farm
hay field. Pioneer organisms modify their environment, thus establishing conditions
under which more advanced organisms can live. An example of this could include
lichens breaking down rock to form small pockets of soil where small plants such as
grasses may become established.
During a succession, each community over time modifies its environment, often making
in time making it more difficult for it to succeed and more favorable for the community
that follows it.
Successions may be categorized as being primary or secondary successions.
A primary succession occurs in an environment where plants previously did not exist. A
primary succession would occur when plants become established on a lava flow from a
volcano. A secondary succession refers to the return of an area to its natural vegetation
following a disruption or removal of the original dominant community. A forest fire
which burns a forest that is a thousand years old would be an example of an event that
could trigger a secondary succession.
Common secondary successions in the Northeastern United States might involve the
abandonment of a plowed field. Within a year or two, annual grasses will take over the
field. These in turn will slowly be shaded out by larger shrubs and blackberry briers.
Eventually these plants will be shaded out by larger sun-loving trees such as cherries,
poplars (alders) and birches. In time these trees will be replaced by larger trees forming
a climax community. In the colder regions of the Northeastern US, the climax
community is often dominated by trees such as beech and maple, while in warmer
climates oak, hickory and even sycamore are more predominate.
The climax community is a self-perpetuating community in which populations remain
stable and exist in balance with each other and their environment. The species diversity
and numbers may remain stable for hundreds or thousands of years in a climax
community. This climax community will persist until a catastrophic change of a major
biotic or abiotic nature alters or destroys it. Wind storms, floods, and disease are just a
few examples of such catastrophic events.
After the climax community has been destroyed, the damaged ecosystem is likely to
recover in stages that eventually result in a stable system similar to the original one.
Image source
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Using the information in the reading passage, define the following terms.
1. ecological succession – new organisms being added to a habitat after the death of
older ones
2. primary succession – succession that leads to a new environment for the first time
3. secondary succession – old environment is revived after a period of time
4. pioneer organism – first living things to pop up from succession
5. climax community -- small plants replaced by larger plants
View of an early ecological succession taken from NY-86 looking toward
Whiteface Mountain in the Adirondack Mountains of New York State.
(Personal photo)
In this activity you will apply what you have learned about succession to the ecological
changes that occur in an abandoned hay field similar to the one pictured above. After
5 years, several types of trees have begun to grow where there was once only hay. You
will observe, by counting, the number and types of some major plants that grow in this
field 5, 15, 40 and 100 years after abandonment.
Part One:
For each data table and diagram on pages 4 through 7, count how many of each species
are found in the field, using the key as a guide. Add up all of the plants to find the total
number of plants present in the field. You will now calculate the percentage of each
species at each observed time period. Use the following formula, rounding your answer
to the nearest tenth.
Number of specific plant ÷ TOTAL number of plants = _______ X 100 = ______%
Please note that the air and soil temperatures in the sample area have also been indicated.
(For the purposes of this activity, we will assume the area is under the influence of the
same type of air mass producing extremely similar climatological conditions in each
different year.)
(Major Plants Observed 5 years after ecosystem disruption)
Beech Tree ◇ 0 0%
● ● ●
■ ● ○
● ○ ●
● ■
■ ■ ●
● ■ ■ ●
● ▲ ● ● ●
(Major Plants Observed 15 years after ecosystem disruption)
Blackberry ● 3 10.7%
TOTAL 28 100%
○ ● ◇ ●
● ▲
■ ▲
○ ▲ ■ ■
■ ○ ○
▲ ■ ● ▲
○ ■ ■ ■
□ ▲ □
▲ ▲
(Major Plants Observed 40 years after ecosystem disruption)
Blackberry ● 0 0%
TOTAL 31 100%
▲ ▲
○ ○ ◇
□ ▲ □ ○
□ ◇ □ ○ ◇
◇ ▲ ○
▲ ◇
▲ □ ○ □ ▲
▲ ◇ ▲ □ ◇
□ ▲
(Major Plants observed 100 years after ecosystem disruption)
Blackberry ● 0 0%
TOTAL 46 100%
□ ◇ ◇
◇ □ □ ◇
◇ ○ ◇ ◇ □
□ ◇ □ ○ ○ ◇ ○
□ □
◇ ◇ ○ ◇ ○ ◇ □
□ ○ □ □
□ ○ ◇ □ □ ○
◇ ◇ □ □ ◇ ◇
□ ○ □
Procedure B: Using the data in the tables you just completed, construct two separate line
graph involving the plant species and insert them below. The types of plant species that
need to be on each graph are listed below. Plot the % of plant species on the y-axis and
time in years on the x-axis. Be sure to include a graph title and a key/legend.
Graph A: Yellow Birch Tree, Sugar Maple Tree, and Beech Tree