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Structure
10.1 Introduction
Objectives
10.2 Ecology
10.3 Ecosystem and its Structure
10.3.1 The Biotic Component
10.3.2 The Abiotic Component
10.1 INTRODUCTION
The purpose of this unit is to make clear some of the basic concepts in ecology. After
defining what is ecology, we will also outline the fundamental structure of an ecosystem.
We will study how each ecosystem has an abiotic (non-living) and a biotic (living)
component and how there is a continuous interaction among these two components. In each
ecosystem there are producer organisms and consumer organisms. Producers generate food
with the help of solar energy while all the consumers are directly or hldirectly dependent on
these producers, for food. There are different kinds of food-chains and we will outline some
examples in this unit. Energy passes along the food-chain in each ecosystem - it is a
one-way traffic. We will, however, note that the various minerals continuously circulate
between the abiotic and biotic components of the ecosystem. These are the so called
biogeochemical cycles.
We will then try to follow the concept of ecological niche and also the concept of
homeostasis.
Objectives
I
After studying this unit, you should be able to
describe the components of an ecosystem,
elaborate upon the biotic and abiotic components and the interaction between
them,
explain the food-chains,food-web and energy flow in an ecosystem,
outline various biogeochernical cycles, and
explain the concept of ecological niche as well as the concept of homeostasis.
10.2 ECOLOGY
Ever since the time of Greek philosophers, there is considerable written material that is
definitely ecological. Man must have been using his ecological knowledge since time
immemorial. This must have been important for his survival in the wild environment. Even
today many tribal people all over the world are using this ecological knowledge and living
happily in the wilderness. With the tremendous increase in population, every one including
you, must understand ecology more thoroughly. This is essential for our survival and well
being.
The word ecology was recently coined by a Gerrnan biologist Emst Haeckel, in 18.69. This
word has a Greek root oikos meaning 'house'. Thus, an ecologist actually.studiesthe
organisms 'at home'.
For the sake of definition we will call ecology as the study of relationship of organisms to
their environment or surrounding. It really inclu&sThe study of the structure and function
of nature. Right here you must note that human being is also another organism and study of
his environment is equally part of ecology.
Broadly speaking, ecology can be subdivided into two major branches (1) autecology, and
(2) synecology. When you are studying one organism and its relationship with the
surrounding environment, you are doing autecologicalstudy. Study of groups of organisms
which are associated together as a unit is the realm of synecology. For example, you are
interested in a particular snake and you want to find out where it lives, what it eats, how
many eggs does it lay, etc., you are pursuing autecology. On the other hand, if you are
interested in a particular river, the kind of terrain it is flowing through, the kinds of fishes
and other animals as well as plants that are found in the river, the inter-relationshipof
plants-animals and physicochemical factors of water, etc., then the objective is
synecological. Both these aspects are important but the synecological studies give total
picture of the situation. Of course, much efforts and time are also required to complete
synecological work. The autecological approach is relatively straightforward but by no
means easy or simple.
Study of ecology is becoming more and more important as we are modifying the
environmpt surrounding us. Here, the word environment is used broadly to denote
everything that surrounds us -that is it includes other living organisms as well as non-living
things. This and subsequent units will give you sufficient idea about the basics of ecology
and its usefulness in day to day planning for economic development.
7
S,P!Q I:
a) Define ecoltrgy.
tp) Detine atnBc~~Bogy aad sy~heh;oBogy.
C) I ~ e k j ~itlg
4 ~icr~~??; I L F la.;tei17ks.opRaic
;!Isb0t~oi3i1i~ I~ c~rgCmisn~s.
d) PIQsal ;irz biotic ai~dsbdoUc coanpoe:eats of sans ecosystcn~:I
(Grass)
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is carnivore). Thus, simply stated the food chain of this type can be :
Plant Herbivore -------+ Carnivore
(Insect) (Lizard or Frog or Bird)
The detritus food-chain begins with organic matter from &ad organisms, etc. This organic
matter is degraded by microorganisms (that is detritivores). Thus, dead leaves, stems and
such parts of plants or dead animals or their organic wastes will be degraded. Thus, it can be
shown as :
Dead log of wood ----c Wood rotting fungi
The important thing about the detritus food-chain is that the activity of microorganisms I
release important inorganic nutrients that can be once again useful for other organisms. Had I
it not been for detritivores, a lot of energy would have remained locked up in the dead
bodies of organisms (Figure 10.1 (a)).
In most ecosystems, the food chains are not so simple and straightforward. An insect like
grasshopper may feed on a dozen differentplants. Similarly, a frog may eat several other
insects along with grasshopper. Thus, the food-chains are not simple isolated sequences as
discussed above. Rather there is considerable interlocking of food-chains. We then speak of
a food-web rather than a foodshain while studying an ecosystem This is because even a
single animal, during its development and growth, may be a pkt of a different food-chain
and hence a food-web.A simple food-chain in a small lake is shown ih Figure 10.1 (b).
Ecosystem : Bwic Concepts
GRAZING CARNIVORES
TYPE
OF
FOOD CHAIN
DETRITUS
TYPE
OF k
FOOD CHAIN MINERALIZATION BACTERIA, FUNGI, PROZOANS
SMALL FISH
..
D ET R I T U S
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level. Thus, the trophic levels can be stated as :
Producer Primary consumer -+ Secondary consumer-Tertiary consumer
But please note that this is not a trophic classification. It tells us about the function of a
species in question. It is also important to remember that one animal may occupy One or
more than one trophic level. This complexity is also evident from the food-web.
10.4.3 Energy, Sources of Energy and Energy Flow
Physicists define energy as the ability or capacity to do work. Work can mean movement,
hands or legs, growth, movement of a car, etc. Virtually all actions of living things reqJre
energy. Muscles do not contract without energy, so you cannot move any part of body
without spending some amount of energy. Not only that, many other reactions going on in
your body (for example movement of cellular constituents, synthesis of cellular proteins a
carbohydrates, ctc.) also require energy. At all levels of biological organization, there ex;
processes that channel energy into the various activities and that this channeling is
admirably well controlled. For this reason it is said that all living systetns, from single cell
to a complex community, are energy convertcrs more than anything else.
Energy can be divided into two basic forms : I) potential energy and 2) kinetic energy.
Potential energy is simply stored energy which is available to do work. The living cells
generally store potential energy in the form of high energy phosphate bonds. When these
bonds are broken the liberated energy can be used to do work. The most common 'energy
rich' compound used by #e cell is adenosine triphosphate or ATP. The other similar
compounds are uridine triphosphate (UTP)or guanosine triphosphate (GTP), etc. The
common reaction is :
where 'ATJ?' is a diphosphate, 'P,' is inorganic phosphatc and 'e' is energy. This reaction
reversible. Although it is beyond the scope of this course, you must know that all animals
generate their own ATP but the energy required to do so comes lrom food they eat
(vegetable or flesh). All we do is break down the sugar or proteins or Pdts and obtain the
potential energy trapped in those molecules.
Kinetic energy on the other h&d is associated with movement. Water falling down a watej
fall (this is how we run ow turbines to generate electricity at hydroelectric station), blowin
wind, the waves pounding on a sea shore or a moving car. All are examples o l kinetic
energy.
There are other forms of energy, like atomic energy, geothermal energy, solar energy or
radiant energy which is in the formof electromagnetic waves, and .also heat.
As we remarked earlier all consumers depend ultimately on solar energy. The production c
a biomass (and hence availability of plant and animal resources) in a given ecosystem is
determined primarily by solar radiation. Mean intensity of solar radialion is the most
important factor. As far as the flow of energy in an ecosystem is concerned, it directly
depends on the amount of solar energy captured per unit area in unit time by the plants.
In the presence of sunlight the chlorophyll (the green pigment) containing plants are capab
of fixing carbon dioxide to form sugar. Thus
light energy
6 CO, + 6 H,O *C,H120, + 60,
(carbon dioxide) (water) (sugar) (oxygen)
n e r e are also pigments other than chlorophyll which can also do the same thing. This
process is photosynthesis.
The amount of solar energy falling on earth is variable according to season, latitude, cloud
cover and such other factors. Much of this energy is wasted and ends up in heating the eartl
and its atmosphere or is reflected back from earth. Only a tiny fraction of incident solar
energy is fixed by the plants to prepare their food (sugar). The amount of plant biomass
developed per unit time per unit area is sometimes known as gross productivity. Plants of
course use some food for their metabolism (respiration) and the remaining biomass is then
the net productivity.
The energy present in the plants is then used by the subsequent members of the food-chain.
This entire movement and overall behaviour of energy obeys the laws of therxnodynamics.
Let us briefly see those two laws and their implications in ecology.
The first law of thermodynamics explains that energy can neither be created nor destroyed
it is merely transformed, that is it changes its fonn. The plants transform solar energy
(radiant energy) into potential energy by using the sunlight to produce sugar from carbon
dioxide. When an animal eats the plant or animal material as food, the same energy is
gradually liberated in the process of respiration to produce the energy rich phosphate
molecules. This we have discussed earlier. It is in this sense that the ecologists say that all
living organisms are merely energy converters.
The second law of thermodynamics points out to us tllat no process of transformation of
energy is possible without partial degradation of energy. By degradation we mean
conversion of concentrated and orderly form of energy into a 'dilute', unusable form. This
means there is a loss of useful energy. Please remember cnergy is not lost or destroyed,
simply it is in a form that is not serving our purpose. The most important ecological
implication of this law is that no biological process can take place with 100% efficiency.
Already we have seen that much of the solar energy incident on plant is not fixed. Further
when an herbivore feeds on this plat11 again some energy is lost, this continues along the
food-chain. Obviously the nature cantlot dford to have too long food-chains. This is the
reason why a unit area of land can feed more vegetarian people lhan if the s'ame area is used
to grow grass on which meat-providing ani~nalsa ~ cgrown: and then tlle meat is pr0vided.a~
a food to people.
Energy also moves only oncc through any given trophic level of Ule food-chain
(Figure 10.2). As we shall see the ininera1 nlaller is used and reuscd in cyclic manner
Energy is gradually degraded along the food-chain 'and finally dispersed as heal.
ORGANIC MATTER
WASTES AND
DEAD BODIES
SAQ 2
a) Explain the cc~lccptof footi-chain with a suit;lblc cxample.
b) llefinc gr;rzing aritl rlctritus t'cn)tl-chains.
c) What is a food-web '!
d) Statc two laws ol' tberlllodyniklllics.
e) Dci'inc potoiitiiil c~lol'gy;ultl ki~lcliccnergy.
A
Ecology These biogeochernical cycles can be broadly divided into two types. One, gaseous type in
which reservoir is almosphere and second, sedimentary type in which reservoir is l.Ile
earth's crust.
We can describe nitrogen cycle as an example to tell you how there is exchange of material
from abiotic to biotic and biotic to abiotic componenls of ui ecosystem. Herc nitrogcn in Ule
air serves as reservoir pool and this nitrogen is fixed by certain planls with the help of
micro-organisms and by electrification process in nature. As an output, nitrogen is fixed
from atmosphere and then ~t is used again and again through various biological processes.
This is how nature uses its resources efficiently.
-1 NITROGEN I N ATMOSPHERE -7
ELECTRIFICATION & NITROGEN F I X I N G
PHOTOCHEMICAL BACTERIA 8 A L 6 A E
FIXATION I
1
/
, DENITRIFICATION
4 NITRATES NITRATES fl
PROTEIN SYNTHESIS
PLANTS -4 ANIMALS
BACTERIA \
EXCRETION,
DECAYING PARTS OF
WASTE PRODUCTS
PLANTS h ANIMALS B Y THE
OF ANIMALS
ACTION OF BACTERIA h F U N G I
4
NITRITE
This is also true for other cycles such as phosphorus, O,, CO,, elc.
Now, we can lean other cycles with the help of a few diagrams :
1) Phosphorus cycle (also known as sedimenlary cycle)
2) Carbon Cycle
PHOTOSYNTHESIS
IGNEOUS ROCKS
NATURAL PROCESSES
DETRPIZTS WSD
OCEAN
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10.8 SUMMARY
In this unit you have noted some basic concepts in ecology. We have defined ecology as tlle
science dealing with the study of the structure and fu~ictionof an ecosystem. The ecosystem
itself is a structural and functional unit of ecology. Many ecosystems together form the
biosphere.
The ecologist may concern himself with single organism (autecology) or with groups of
organisms in an ecosystem (synecology). I
Each ecosystem, you must note, has a living (bioti~)and a non-living (abiotic) components.
These two components are continuously exchanging materials with each other. This is what
we have discussed as biogeochemical cycles.
~ a c ecosystem
h also has a definite trophic structure and you can speak about the trophic
status of each organism. The plants which manufacture food are called as producers while
all the rest organisms are consumers. You have observed that the food energy present in I
abiotic component
C biotic component I
Different minerals follow different path, so is the case with the gases like Ecosystem : 13;lsic Concepts
nitrogen, oxygen.
b) Each organism of an ecosystem has a definite function, a definite role to play.
This functional status of an organism is called as ecoIogical niche. The
function of an organism can be analyzed only when we h o w more about the
place where the organism lives (spatial niche), the energy sources, etc, of an
organism (trophic niche) and environmental gradient (say, of temperature) in
which it lives (multidimensional niche). The all encompassing concept is that
of ecological niche which includes all the aspects discussed above.
c) Homeostasis is the tendency of an ecosystem to maintain equilibrium and
resist changes (Homeo = same, Stasis = standing). External agents are
constantly disturbing the equilibrium yet as soon as disturbance is over the
system r e m s to nonnal. This is possible due to various feedback control
I signals that operate to maintain equilibrium. Such controls are at the
individual, population and ecosystem level. Maintenance of constant blood
sugar level or maintenance of right body temperature are examples of
homeostasis. Regulation of prey population by a predator is an example at the
ecosystem level.