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in natural vegetation of tropical, subtropical and temperate regions and are abundant in
tropical Asia12. In these areas bamboos have been used as a main material for house
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Bamboos constitute the Bambusoideae, a subfamily of the grass family, the
Graminae. About 750 bamboo species in about 45 genera are known around the world1.
Many of them are indigenous to the monsoon areas of tropical Asia, some of them occur
The genus Phyllostachys is one of the largest genera of bamboo and about 30
species are known in East Asia. In Japan, three species (P. bambusoides Siebold et
Zuccarini, P. pubescens Mazel ex Houzeau de Lahaie and P. nigra (Loddiges) Munro var.
which run horizontally under the ground and forms groves of evenly spaced culms14.
Moso bamboo (Phyllostachys pubescens Mazel), one of the largest and typical
bamboos of Japan as shown in Photo 3.1, grows to its full size within about two months,
but takes about three years for complete maturation. The largest of bamboo culms
marked by one prominent ridge, but branch-bearing nodes of mid-culms marked by two
prominent ridges, i.e., the supranodal ridge and sheath scar as the other species of this
genus. This plant is native from China. It is said that it was introduced into Kagoshima
Japan and the northern limit of cultivation of this plant in Japan is Hakodate, Hokkaido.
The fundamental architecture of the cell wall is completed at an early stage but
secondary thickening of cell walls and their lignifications proceed more gradually.
Chrystallinity changes of cellulose in the cell walls of internodes from juvenile to mature
stages shows that cellulose formed in the early stage is more or less amorphous and
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Moso bamboos not only have such ecologically unique characteristics, but also
are also economically important. Culms are used for building construction and
handicrafts, and young shoots are edible. From the viewpoint of resource management of
the biomass, asexual culms reproduction accompanied by rapid growth of the sprouts is a
very advantageous trait. We can continuously harvest culms by thinning from a stand
3.2. Anatomy
The bamboo culms are characterized by nodes occurring periodically along its
length. At the node is a solid cross wall, called the diaphragm, and the internodes are
usually hollow (Photo 3.2). The outermost part of the culm is composed of single layer
of epidermis and the inside is covered by a layer of sclerenchyma cells. The body of the
parenchyma cells (Photo 3.3 and Photo 3.4.). Sclerenchyma cells are long, dead, thick-
walled cells and serve as strengthening material. They surround the vascular bundles,
separated by parenchyma. Just inside of the outer skin of the stem a couple of layers of
heavy sclerenchyma cells are found. The length of the fibres depends on the species, and
varies in the culm. The parenchyma tissue forms a ground tissue in which the vessels and
fibres are embedded. The parenchyma cells are mostly thin-walled and connected to each
other by numerous pits. The shape of the cells is either long or cube-like. The
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Photo 3.2. The moso bamboo shoot and culms
The vascular system in bamboo is divided into two part; first was the phloem, that
part of the vascular bundle towards the epidermis, made up of sieve tubes, for conduction
of food materials principally downwards; large and thin-walled cells; second was xylem,
inside of the phloem, composed of vessels, principally for the transport of water and
minerals upwards; thicker walled cells. The pattern vascular system in the cross section
of a culm, the longitudinal variation within the whole culm, which decreasing from base
to top is described in detail by Grosser and Liese16. Bamboo culm has no cambium and
no radial cell elements, such as rays, exist in the internodes. Bamboo is therefore quite
different from wood. The culms consist of about 50% parenchyma, 40% fibers and 10%
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Photo 3.3. Cross section of moso bamboo culms
In bamboo, the parenchymatous ground tissue is lined on the outer and the inner
side by special terminal layers. The layers are heavily cutinized, such as overlaid with a
waxy covering, to prevent loss of water from the culms. Liese17 also reported that the
cutinized layer is composed of cellulose and pectin and has a tangential lamellation. A
wax layer is formed on top and the outside has a heavy deposit of silica, which is
incorporated in peripheral parts of the culms tissue. These materials make the bamboo
The anatomical bamboo structure becomes highly modified and complex in the
nodes. Here the vascular bundles and the phloem become extensively branched and
research topic.
12
px mp
mx
bs
vascular bundle
parenchyma
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3.3. Chemical properties
Bamboo consists of about 50-70% holocellulose, 30% pentosan and 20-25%
lignin, with some variation according to species and age of the culm. More than 90% of
the bamboo hemicellulose consists of xylan, which has a unique structure within the
hardwoods, except that alkaline extract, ash and silica content are higher than in
hardwoods. Not enough is known about the location of the silica, which amount to 0.5 to
4% of the culm and affects its pulping quality. Most silica appears to be deposited in the
epidermis, whereas the internodes and nodes contain none or little. As seen in Table 3.1,
some researcher have done test on chemical composition on several bamboo as well.
a
Chao et al.20, bSingh and Guha21 and cTamolang22; na=not available.
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3.4. Physical and mechanical properties
3.4.1. Objective
The main objective of this study was to obtain basic data on the physical and
content, node presence and loading orientation with respect to the outer culms surface.
Prefecture in Japan was chosen as raw material. From 20 culms (100 cm long), 30 cm
long pieces, one of them with the node at midsection and another without node, were cut
from each section for testing the bamboo’s mechanical properties. Every 30 cm long
piece was then split into quarter, two for air-drying at 200C and 65% relative humidity
(RH) and the other remaining in green condition (Photo 3.5). From each of the above
quarter, two bending samples (2 cm wide and 25 cm long with the thickness of the
bamboo wall), two tension samples (1 cm wide, 20 cm long and 0.1 cm center neck with
the bamboo wall thickness), and two compression samples (1 cm wide and 3 cm long
with the thickness of bamboo wall) were obtained (Photo 3.6). Minimum planning was
done on the inner and outer perimeters/layers to make the samples rectangular.
The specimens were tested in compression, tension and bending following the JIS
Z-211323 standard procedures, while shear strength was adopted from another researcher.
The bending test was done with a simple support and concentrated center load and with a
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Photo 3.5. Bamboo raw material after split into quarter.
Photo 3.6. Specimens for bending, compression and tension testing. From left:
specimens with node and without node, respectively.
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After the bending test of the green samples, 2 cm wide by 5 cm long specimen
was cut from each sample for measure of the percent shrinkages in tangential, radial and
longitudinal directions from green to air-dry condition (65% RH at 200C). The moisture
P P
Inner face Outer face
h
Outer face Inner face
b
(Down) (Up)
2 cm
15 cm
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3.4.3. Physical properties
The properties of the bamboo culms are determined by its anatomical structure
since the distribution of cell types within the culm is influenced by the shape, size,
arrangement and number of vascular bundles. These factors contribute to the bamboo’s
Anatomically, density of bamboo regarded as a function of the ratio of cell wall volume
to cell void volume; as such, it is affected by cell wall thickness and structure, cell width,
relative proportions of different types of cells and the kind and amount of extractives
As summarize in Table 3.2, the average MC was 35.79% in the green condition
and 12.48% in the air-dry condition, whereas the average of basic density or specific
gravity was 0.74. The density of bamboo is greatest at the surface of the culm and
decreases from there inward. Density also depends on vertical position in the culm,
increasing from bottom to top. The variations in density are closely correlated with the
relative proportions of vascular bundles and ground tissues; especially as the proportion
of thick-walled sclerenchyma cells increases, the density increases also24. For two
botanical species, P. pubescens (moso chiku) and P. reticulata (madake), Ota25 has
Shrinkages
Bamboo, like wood, is anisotropic and has its principal directions the longitudinal,
radial and tangential as referred to the cylindrical shape of the culms. The shrinkages of
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specimens were measured from green to air-dry conditions. Unlike wood, bamboo
shrinks right from the beginning of drying. The average shrinkages in longitudinal,
tangential and radial directions were 0.13, 3.72, and 6.15%, respectively (Table 3.2). The
greatest shrinkages occurred in the radial direction, which was almost 1.6 times as great
as shrinkage in the tangential direction. The shrinkage in the longitudinal direction was
negligible. These result was agreed with Ota25, for P. pubescens and P. reticulata the
radial, tangential and longitudinal shrinkage values from green to oven-dry were 6.65,
4.45 and 0.33%, respectively. Shrinkage is least in longitudinal direction, but in contrast
Moisture content, %
Green 33.3 51.43 35.8 6.73
Air-dry 11.3 13.15 12.5 1.50
Shrinkages, %
Longitudinal 0.09 0.21 0.13 0.02
Tangential 3.12 5.22 3.72 0.59
Radial 4.85 8.41 6.15 0.86
Number of samples, 20; min, minimum value; max, maximum value; SD, standard deviation from
the sample mean; shrinkages calculated from green to air-dry condition.
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