Biological Sciences: Postgraduate Opportunities 2017

Biological Sciences
Postgraduate Opportunities 2017
Influencing the world since 1583

The University of Edinburgh
Biological Sciences Postgraduate Opportunities 2017 01
“Edinburgh isn’t so much a city, more 02 Introduction

04 Taught masters programmes
a way of life … I doubt I’ll ever tire of 10 Online distance learning
exploring Edinburgh, on foot or in print.” 12 Research at the School of
Biological Sciences
Ian Rankin 13 Research opportunities
Best-selling author and alumnus 16 About the School of Biological Sciences
17 Facilities and resources
18 Community
19 Employability and graduate attributes
20 Applications and fees
22 Funding
24 Campus map
25 Get in touch
02 www.ed.ac.uk/biology The University of Edinburgh
Influencing the For more than 400 years the University of

Edinburgh has been changing the world. Our
staff and students have explored space, won
TOP 50
We’re consistently ranked one of
world since 1583 Nobel Prizes and revolutionised surgery. They’ve
published era-defining books, run the country,
made life-saving breakthroughs and laid the
the top 50 universities in the world.
We’re 19th in the 2016/17 QS World
foundations to solve the mysteries of the universe. University Rankings.
Our distinguished alumni include NASA astronaut

Piers Sellers, former MI5 Director-General Dame
Stella Rimington, Olympians Sir Chris Hoy and
4 TH
We’re ranked fourth in the UK for
16 Nov 2016 Katherine Grainger and historical greats such research power, based on research
Postgraduate Open Day as philosopher David Hume, suffragist Chrystal quality and breadth.*
www.ed.ac.uk/
Macmillan, who founded the Women’s International
83%
postgraduate-open-day League for Peace and Freedom, and physicist and
mathematician James Clerk Maxwell.
International collaboration The majority of our research – 83 per

An internationally renowned centre for academic cent – is considered world leading or
excellence, we forge world-class collaborations internationally excellent.*
with partners such as the California Institute of
Technology (Caltech), Stanford University, the
University of Melbourne, Peking University, the
University of Delhi and the University of KwaZulu-
23 RD
We’re ranked 23 in the world for
rd
Natal. As a member of the League of European the employability of our graduates.**
Research Universities and the Coimbra Group,
£305m
we link up with leading institutions across Europe.
Linking research and commerce

We were one of the first UK universities to develop In 2014/15 we won £305 million
commercial links with industry, government and in competitive research grants.
the professions. Edinburgh Research and Innovation
(ERI) promotes and commercialises our research
excellence and can assist you in taking the first
step to market, through collaborative research,
20
We’re associated with 20 Nobel Prize
licensing technology or consultancy. winners, including physicists Peter Higgs,
Charles Barkla and Max Born, medical
Enhancing your career
researcher Peter Doherty and biologist
We are committed to embedding employability
in your University experience and have one of the
Sir Paul Nurse.
Russell Group’s best track records for graduate
employment. From volunteering schemes to our
sector-leading Careers Service, we provide you with 137 NATIONALITIES
Students from two-thirds of the
opportunities to develop your skills, knowledge
and experience, giving you an edge in the world’s countries study here.
competitive job market.
twitter.com/applyedinburgh
facebook.com/applyedinburgh
youtube.com/edinburghuniversity
* Research Excellence Framework (REF) 2014
instagram.com/applyedinburgh ** Latest Emerging Global Employability University Ranking
Taught masters programmes www.ed.ac.uk/pg/856 www.ed.ac.uk/pg/1
Biochemistry Biodiversity & Taxonomy

Our taught masters programmes are designed to www.ed.ac.uk/pg/763 of Plants
deepen knowledge in a subject you will typically
have studied at undergraduate level, although Animal Breeding & Genetics MSc 1 yr FT
PgDip 9 mths FT
MSc 1 yr FT
PgDip 9 mths FT
they can also offer a new direction in your
academic career. Programme description Programme description
This programme offers you an academically challenging and The understanding of plant diversity and resources has never been more
MSc and Diploma MSc 1 yr FT (2 yrs PT available for UK/EU students) career-developing study of biological systems at the molecular and important. As we face the unprecedented challenges of climate change
Taught masters programmes last 12 months, consisting of two PgDip 9 mths FT cellular level. Biochemistry is fundamental to most areas of life science; and environmental degradation, effective environmental surveillance and
it has a major impact on modern medical research and is essential in conservation depend upon detailed knowledge of plants and their habitats.
semesters of taught courses followed by an individual supervised
research project. An alternative option is the Postgraduate Programme description the pharmaceutical, nutrition, forensic, bioengineering, agricultural and
This programme is run jointly by the University and the world-renowned
The revolution in genetic mapping technology and the advent of whole environmental industries. The programme is designed to produce highly
Diploma, which lasts nine months from September to May, Royal Botanic Garden Edinburgh (RBGE), which is home to one of the
genome sequences have turned quantitative genetics into one of the skilled and motivated biochemists that are suitable for employment in
consisting of the taught courses only. world’s best living collections of plants (15,000 species across four sites,
fastest growing areas of biology. the life sciences or for further academic research. You will be taught
amounting to five per cent of known world species), a herbarium of three
to apply chemical and physical principles to biological molecules in
Based in the internationally renowned Institute of Evolutionary Biology, million preserved specimens and one of the UK’s most comprehensive
Online distance learning complex living systems in order to expand your understanding of
this MSc draws from the wealth of expertise available there, as well as the botanical libraries.
We are the biggest provider of online distance learning in the the molecular basis of the processes which take place within these
teaching, research expertise and facilities of Scotland’s Rural College, the RBGE offers collections-based biodiversity research opportunities across
Russell Group. Just like our on-campus provision, all our online organisms. Through a combination of taught courses, practical skills
University’s Centre for Genomics and Experimental Medicine, the Medical a wide spectrum of organisms and geographical regions. This diversity,
masters are delivered by dedicated academics, many of whom training and laboratory-based research, you will explore the structures,
Research Council’s Human Genetics Unit and the Roslin Institute coupled with RBGE’s world-leading research in different continents,
are leaders in their field. You can study part time towards a dynamics, interactions and metabolic pathways of biological molecules,
(birthplace of Dolly the sheep). provides an unrivalled masters programme in plant biodiversity.
from small molecules to large macromolecular complexes.
Postgraduate Certificate, Postgraduate Diploma or MSc, and the
Each year the syllabus is fine-tuned to suit current issues in evolutionary, Programme structure
timescales are flexible – you can complete the MSc in two years Programme structure
plant, human and animal genetics. This programme forms part of the This programme is full time and consists of two semesters of lectures,
or take up to six years. You can also take individual courses for Teaching and learning activities include lectures, tutorials, workshops,
quantitative genetics and genome analysis suite of programmes (see practicals, workshops and investigations, followed by a four-month
Continuing Professional Development purposes. Online distance presentations, laboratory work, practical skills training and a research
page 8), which includes the specialist routes Human Complex Trait research project. The programme includes a two-week field course in
learning is an extremely flexible option for students looking to project, literature and database searching, discussion groups and project
Genetics and Evolutionary Genetics. a tropical country (recently Belize). The programme is delivered mainly
groups and seminars.
balance further study with professional or family commitments.
Programme structure at RBGE but also at the University’s King’s Buildings campus. There are
This programme consists of two semesters of taught courses followed COMPULSORY COURSES PREVIOUSLY OFFERED INCLUDE: no option elements to the programme – all courses are compulsory.
by a research project, leading to a dissertation. Courses are taught via
See also… lectures, tutorials, seminars and computer practicals. Assessment is
Biochemistry A and B; Biophysical Chemistry; Research Project Proposal;
Practical Skills in Biochemistry A and B; MSc Project & Dissertation.
COURSES PREVIOUSLY OFFERED INCLUDE:
You may also be interested in taught masters programmes by written examinations, in-course assignments and project work. Conservation and Sustainability; Taxonomy and Plant Collections;
offered elsewhere in the University, particularly the College of OPTION COURSES PREVIOUSLY OFFERED INCLUDE : Biodiversity of Angiosperms; Evolution of Cryptogams and Fungi; Evolution
COMPULSORY COURSES PREVIOUSLY OFFERED INCLUDE:
of Angiosperms; Plant Geography; Phylogenetics and Population Genetics;
Medicine & Veterinary Medicine or the Schools of Chemistry, Applicable Mathematics; Information Processing in Biological Cells;
Population and Quantitative Genetics; Genetic Interpretation; Linkage Biodiversity of Cryptogams and Fungi; Tropical Biodiversity Field Course.
Informatics, or Engineering. Molecular Modelling and Database Mining; Preparative Methods for
and Association in Genome Analysis; Animal Genetic Improvement;
Structural Biology; Tools for Synthetic Biology; Applications of Synthetic Research
Quantitative Genetic Models; Research Proposal; Statistics and Data
www.ed.ac.uk/studying/prospectus-request Biology; Bioinformatics; Biomacromolecules; Commercial Aspects Your research project will be chosen in consultation with your
Analysis; Dissertation.
of Drug Discovery; Detailed Characterisation of Drug or Ligand supervisor, and will link directly with active research programmes at
Interactions using SPR; Drug Discovery; Programming for the Life RBGE or other research institutions. The field trip, together with training
OPTION COURSES PREVIOUSLY OFFERED INCLUDE:
Sciences; Quantitating Drug Binding; Economics and Innovation and a short practical exam, qualifies you for the RBGE Certificate in
Molecular Phylogenetics; Bioinformatics; Molecular Evolution; in the Biotechnology Industry; Functional Genomic Technologies; Practical Field Botany.
Genetics of Human Complex Traits; Functional Genomic Technologies; Protein Structure Determination; Vaccines and Molecular Therapies. Career opportunities
Evolutionary Quantitative Genetics.
Research The programme is good preparation for roles in taxonomy, while
Career opportunities Students progressing to MSc level will carry out their own research project many graduates have also continued to PhD or other research roles at
You will develop the in-depth knowledge and specialised skills required at the frontier of knowledge and can make a genuine contribution to the universities internationally. Recent graduates have entered a wide variety
to apply quantitative genetics theory to practical problems, in both the progress of original research. This also involves reviewing relevant papers, of jobs, including ecologist, plant scientist, plant surveyor, environmental
biomedical and animal science industries, and to undertake research analysing data, writing a dissertation and giving a presentation. officer and plant health inspector, for research institutions, conservation
in evolutionary genetics, population genetics and genome analysis. agencies and other employers, such as Kew Gardens, the Royal Botanic
Career opportunities Garden Edinburgh, Corrour Estate and Scottish Natural Heritage.
Entry requirements You will enhance your career prospects by acquiring knowledge of
A UK 2:1 honours degree, or its international equivalent (www.ed.ac.uk/ contemporary biochemistry from world experts in the field, by being Entry requirements
international/graduate-entry), in biological or biomedical sciences, trained in advanced analytical and presentation skills, and by having A UK 2:1 honours degree, or its international equivalent (www.ed.ac.uk/
medicine, agriculture or animal sciences, with evidence of quantitative independent research experience in a modern, world-class laboratory. international/graduate-entry), in biological or environmental science.
skills. We will also consider your application if you have a background in Relevant work experience is desirable.
Entry requirements
mathematics, statistics or physics and can show evidence of an interest English language requirements
A UK 2:1 honours degree or its international equivalent (www.ed.ac.uk/
in genetics, or if you have less than the minimum qualification but can See page 20.
international/graduate-entry) in biochemistry, biological sciences
show sufficient additional relevant experience (e.g. several years working
(with some relevant chemistry components) or chemistry (with some Fees and funding
in the animal breeding sector).
relevant biology components). For fees see page 20 and for funding information see page 22.
English language requirements
See page 20.
See page 20. Programme Directors Dr Andrew Hudson and Dr Louis Ronse de Craene
Fees and funding Tel +44 (0)131 650 8651
Fees and funding
For fees see page 20 and for funding information see page 22. Email btpmsc@ed.ac.uk
For fees see page 20 and for funding information see page 22.
Programme Director Professor Andrew Leigh-Brown

Programme Director Dr Janice Bramham
Tel +44 (0)131 650 5523
Tel +44 (0)131 650 4786
Email qgen@ed.ac.uk
Email mscbiochemistry@ed.ac.uk
www.ed.ac.uk/pg/2 www.ed.ac.uk/pg/676 www.ed.ac.uk/pg/3 www.ed.ac.uk/pg/764
Bioinformatics Biotechnology Drug Discovery & Evolutionary Genetics

Translational Biology
MSc 1 yr FT MSc 1 yr FT MSc 1 yr FT MSc 1 yr FT (2 yrs PT available for UK/EU students)
PgDip 9 mths FT PgDip 9 mths FT PgDip 9 mths FT PgDip 9 mths FT
Programme description Programme description Programme description Programme description

Bioinformatics is about the application of computer-based approaches Changing demographics and growing demand for food, fuel and The rapid transformation in the nature of drug discovery means that The revolution in genetic mapping technology and the advent of whole
to understanding biological processes. Our programme will introduce agricultural and environmental sustainability are among the key knowledge of related disciplines, and the technologies used, is essential genome sequences have turned quantitative genetics into one of the
you to the current methods used to interpret the vast amounts of data challenges the world faces today. for those considering a career in commercial or academic research. This fastest growing areas of biology.
generated by modern high-throughput technologies. MSc will help you explore the latest methods of developing drugs and
In this MSc you will learn research and development skills to enable the Based in the internationally renowned Institute of Evolutionary Biology,
therapeutic compounds for humans and animals and disease control
The aim of this MSc is to equip you with a strong background in biology, creation of new products and services. You will investigate the economic this MSc draws from the wealth of expertise available there, as well as the
agents for plants. You will learn about marketing, licensing and
plus the computing skills and knowledge necessary to navigate the vast basis for current biotechnology structures and areas of future demand, teaching, research expertise and facilities of Scotland’s Rural College, the
regulations, which are all part of the development process.
wealth of modern biological data. On completing this programme you including the global pharmaceutical industry and carbon sequestration. University’s Centre for Molecular Medicine, the Medical Research Council’s
will be able to take up PhD studies or bioinformatics posts in academia You will learn how technology can be applied to solve pressing real-world Our multidisciplinary approach links structural biology, bioinformatics, Human Genetics Unit and the Roslin Institute (birthplace of Dolly the sheep).
or in industry. The programme covers programming skills, statistical biological problems and gain the skills and expertise needed for future chemistry and pharmacology. You will investigate the fundamental
Each year the syllabus is fine-tuned to suit current issues in evolutionary,
analysis and database science as well as bioinformatics. Option courses developments in biotechnology. scientific problems and techniques of drug discovery and design, alongside
plant, human and animal genetics. This programme forms part of the
allow you to specialise in several aspects of bioinformatics. the challenges of developing principles for new therapeutic strategies. You
Programme structure quantitative genetics and genome analysis suite of programmes (see
will have hands-on experience of crystallographic computer programming
Programme structure This programme consists of two semesters of taught courses followed page 8), which includes the specialist routes Animal Breeding & Genetics
and computation for bioinformatics. You will consider the moral and ethical
The MSc comprises two semesters of taught courses followed by a research by a research project or industrial placement, leading to a dissertation. and Human Complex Trait Genetics.
aspects of the agrochemical and pharmaceutical industries through case
project and dissertation. The project is a key element in deciding how
COMPULSORY COURSES PREVIOUSLY OFFERED INCLUDE: studies, seminars and discussions. Programme structure
your career in bioinformatics should develop further. Teaching is through
This programme consists of two semesters of taught courses followed
lectures, tutorials, seminars, computer practicals and lab demonstrations. Economics and Innovation in the Biotechnology Industry; Intelligent Programme structure by a research project, leading to a dissertation. Courses are taught via
Agriculture; Principles of Industrial Biotechnology; Research Project This programme consists of two semesters of taught courses followed
COMPULSORY COURSES PREVIOUSLY OFFERED INCLUDE: lectures, tutorials, seminars and computer practicals. Assessment is
Proposal; Research Project or Industrial Placement. by a research project, leading to a dissertation.
by written examinations, in-course assignments and project work.
Bioinformatics Programming & System Management; Bioinformatics
COMPULSORY COURSES PREVIOUSLY OFFERED INCLUDE:
Research Proposal; MSc Dissertation (Bioinformatics); Statistics & OPTION COURSES PREVIOUSLY OFFERED INCLUDE: COMPULSORY COURSES PREVIOUSLY OFFERED INCLUDE:
Data Analysis. Applicable Mathematics; Molecular Modelling and Database Mining;
Biobusiness; Bioinformatics; Bioinformatics Programming & System Population and Quantitative Genetics; Genetic Interpretation; Linkage
Quantitating Drug Binding; Protein Structure Determination; Commercial
Management; Drug Discovery; Commercial Aspects of Drug Discovery; and Association in Genome Analysis; Research Proposal; Statistics and
OPTION COURSES PREVIOUSLY OFFERED INCLUDE: Aspects of Drug Discovery; Project Proposal and Literature Review;
Environmental Gene-Mining and Metagenomics; Enzymology and Data Analysis; Dissertation.
Preparative Methods for Structural Biology; Drug Discovery.
Bioinformatics 1; Human-Computer Interaction; Information Processing in Biological Production; Gene Expression and Microbial Regulation; Industry
Biological Cells; Molecular Modelling and Database Mining; Quantitating & Entrepreneurship in Biotechnology; Molecular Modelling and Database OPTION COURSES PREVIOUSLY OFFERED INCLUDE:
Drug Binding; Bioinformatics Algorithms; Bioinformatics 2; Functional Mining; Programming for the Life Sciences; Social Dimensions of
Molecular Phylogenetics; Bioinformatics; Molecular Evolution; Genetics
Genomic Technologies; Introduction to Website and Database Design Systems and Synthetic Biology; Stem Cells and Regenerative Medicine; Biobusiness; Bioinformatics 1; Chemical Medicine; Functional Genomic
of Human Complex Traits; Quantitative Genetic Models; Functional
for Drug Discovery; Molecular Phylogenetics; Next Generation Biochemistry; Tools for Synthetic Biology; Applications of Synthetic Technologies; Biophysical Chemistry for MSc Biochemistry; Introduction
Genomic Technologies; Animal Genetic Improvement; Evolutionary
Genomics; Software Architecture, Process and Management; Drug Biology; Practical Skills in Biochemistry; Vaccines and Molecular Therapies. to Scientific Programming; Biochemistry; Practical Skills in Biochemistry;
Quantitative Genetics.
Discovery; Introduction to Java Programming; Software Development; Introduction to Website and Database Design for Drug Discovery; Detailed
Practical Systems Biology.
Research and laboratory work Characterisation of Drug or Ligand Interactions Using Surface Plasmon Career opportunities
There will be a considerable practical element to the programme. You
Resonance (SPR); Bioinformatics Programming & System Management; You will develop the in-depth knowledge and specialised skills required
Research will work in a biotechnology laboratory and learn how experimental
Bioinformatics Algorithms; Information Processing in Biological Cells; to apply quantitative genetics theory to practical problems, in both the
The research project is carried out independently, but under the technology is designed and operated.
Bioinformatics 2; Protein Molecular Modelling Practical Skills; Tools biomedical and animal science industries, and to undertake research
guidance of a supervisor, during the summer, with results presented
Career opportunities for Synthetic Biology. in evolutionary genetics, population genetics and genome analysis.
in a dissertation. A wide range of projects is available through both
The programme will open up a wide variety of career opportunities,
the School of Biological Sciences and the School of Informatics. Career opportunities Entry requirements
ranging from sales and marketing, to research and development, to
This MSc is designed to help you pursue a career in the pharmaceutical A UK 2:1 honours degree, or its international equivalent (www.ed.ac.uk/
Career opportunities manufacturing and quality control and assurance. Recent graduates
international/graduate-entry), in biological or biomedical sciences,
industry or relevant government agencies, and it will provide a good
The programme is good preparation for further academic research or have found work as biologists, development scientists and researchers
background for managerial or technical roles in research, design and with evidence of quantitative skills. We will also consider your application
for technical or managerial roles in various commercial sectors, including for employers including Renishaw Diagnostics, Aridhia Informatics, the
development. It is also a solid basis from which to continue your studies if you have a background in mathematics, statistics or physics and can
medical electronics, life sciences and defence. Recent graduates are now Ministry of Food, Agriculture and Livestock, and universities internationally.
to PhD level. Recent graduates have found roles in healthcare, research show evidence of an interest in genetics.
working as bioinformaticians, software developers, data analysts and
Entry requirements and new drug screening, for a range of pharmaceuticals companies. English language requirements
system testers.
A UK 2:1 honours degree, or its international equivalent (www.ed.ac.uk/
Entry requirements See page 20.
Entry requirements international/graduate-entry), in biotechnology, biochemistry,
A UK 2:1 honours degree, or its international equivalent (www.ed.ac.uk/ Fees and funding
A UK 2:1 honours degree, or its international equivalent (www.ed.ac.uk/ molecular biology, genetics, plant sciences or related sciences.
international/graduate-entry), in biochemistry, molecular biology, For fees see page 20 and for funding information see page 22.
international/graduate-entry), in biological sciences. You must have
English language requirements biophysics or pharmacology. We will also consider your application if you
a strong background in molecular biology, biochemistry or related
See page 20. have a background in chemistry, physics or computer science. We may
sciences and some experience of computer science or mathematics. Programme Director Dr Jacob Moorad
Fees and funding consider your application if you have a background in another science
We will also consider your application if you have a background in Tel +44 (0)131 650 5442
For fees see page 20 and for funding information see page 22. or medicine; please contact us to check before you apply.
chemistry, physics, mathematics or engineering. Email qgen@ed.ac.uk
English language requirements English language requirements
Programme Director Dr Andrew Free See page 20.
See page 20.
Tel +44 (0)131 650 5338 Fees and funding
Fees and funding Email biotechmsc@ed.ac.uk For fees see page 20 and for funding information see page 22.
Programme Director Dr Paul Taylor

Programme Director Dr Simon Tomlinson
Tel +44 (0)131 650 7058
Tel +44 (0)131 651 9554
Email drugdiscovery@ed.ac.uk
Email bioinfmsc@ed.ac.uk
Human Complex Quantitative Genetics Synthetic Biology Systems & Synthetic Biology
Trait Genetics & Genome Analysis & Biotechnology
MSc 1 yr FT (2 yrs PT available for UK/EU students) MSc 1 yr FT (2 yrs PT available for UK/EU students) MSc 1 yr FT MSc 1 yr FT
PgDip 9 mths FT PgDip 9 mths FT PgDip 9 mths FT PgDip 9 mths FT
Programme description Programme description Programme description Programme description

The revolution in genetic mapping technology and the advent of whole The revolution in genetic mapping technology and the advent of whole This academically challenging and career-developing programme focuses We offer an opportunity to train in one of the newest areas of biology:
genome sequences have turned quantitative genetics into one of the genome sequences have turned quantitative genetics into one of the fastest on research and development using biological and chemical principles the application of engineering principles to the understanding and
fastest growing areas of biology. growing areas of biology. This MSc is one of a suite of programmes offering and systems to create new products, services and industries. design of biological networks. This new approach promises solutions to
specialist routes in genetics. The other programmes are Animal Breeding some of today’s most pressing challenges in environmental protection,
Based in the internationally renowned Institute of Evolutionary Biology, You will employ elements of the developing field of synthetic biology to
& Genetics, Evolutionary Genetics, and Human Complex Trait Genetics. human health and energy production.
this MSc draws from the wealth of expertise available there, as well as the bring about significant changes and major innovations that address the
teaching, research expertise and facilities of Scotland’s Rural College, the Based in the internationally renowned Institute of Evolutionary Biology, challenges of rapidly changing human demographics, resource shortages, This MSc will provide you with a thorough knowledge of the primary
University’s Centre for Molecular Medicine, the Medical Research Council’s this MSc draws from the wealth of expertise available there, as well as the energy economy transition and the concomitant growth in demand for design principles and biotechnology tools being developed in systems
Human Genetics Unit and the Roslin Institute (birthplace of Dolly the sheep). teaching, research expertise and facilities of Scotland’s Rural College, the more and healthier food, sustainable fuel cycles, and a cleaner environment. and synthetic biology, ranging from understanding genome-wide data to
University’s Centre for Molecular Medicine, the Medical Research Council’s designing and synthesising BioBricks. You will learn quantitative methods
Each year the syllabus is fine-tuned to suit current issues in evolutionary, Programme structure
Human Genetics Unit and the Roslin Institute (birthplace of Dolly the sheep). of modelling and data analysis to inform and design new hypotheses
plant, human and animal genetics. This programme forms part of the You will learn through a variety of activities, including lectures,
based on experimental data. The University’s SynthSys centre is a hub
quantitative genetics and genome analysis suite of programmes (see Each year the syllabus is fine-tuned to suit current issues in evolutionary, workshops, presentations, laboratory work, field work, tutorials,
for world-leading research in both systems and synthetic biology.
right), which includes the specialist routes Animal Breeding & Genetics plant, human and animal genetics. Applicants who wish to select their seminars, discussion groups, project groups and problem-based learning
and Evolutionary Genetics. area of specialisation during the programme should apply for this activities. You will attend problem-based tutorial sessions and one-to-one Programme structure
umbrella programme. meetings with your personal tutor or programme director. The programme consists of two semesters of taught courses followed by
Programme structure
a research project and dissertation, which can be either modelling-based
This programme consists of two semesters of taught courses followed Programme structure You will carry out research at the frontier of knowledge and can make
or laboratory-based.
by a research project, leading to a dissertation. Courses are taught via This programme consists of two semesters of taught courses followed a genuine contribution to the progress of original research. This involves
lectures, tutorials, seminars and computer practicals. Assessment is by a research project, leading to a dissertation. carrying out project work in a research laboratory, reviewing relevant COMPULSORY COURSES PREVIOUSLY OFFERED INCLUDE:
by written examinations, in-course assignments and project work. papers, analysing data, writing reports and giving presentations.
COMPULSORY COURSES PREVIOUSLY OFFERED INCLUDE: Information Processing in Biological Cells; Social Dimensions of Systems
COMPULSORY COURSES PREVIOUSLY OFFERED INCLUDE: COMPULSORY COURSES PREVIOUSLY OFFERED INCLUDE: and Synthetic Biology; Dissertation project; Practical Systems Biology;
Population and Quantitative Genetics; Genetic Interpretation; Linkage
Applications of Synthetic Biology; Tools for Synthetic Biology.
Population and Quantitative Genetics; Genetic Interpretation; Linkage and Association in Genome Analysis; Research Proposal; Statistics and Applications of Synthetic Biology; Tools for Synthetic Biology; Social
and Association in Genome Analysis; Genetics of Human Complex Traits; Data Analysis; Dissertation. Dimensions of Systems & Synthetic Biology; Environmental Gene Mining &
Quantitative Genetic Models; Research Project Proposal; Statistics and Metagenomics; Research Project Proposal; MSc Project and Dissertation.
Data Analysis; Dissertation. OPTION COURSES PREVIOUSLY OFFERED INCLUDE: Neural Computation; Probabilistic Modelling and Reasoning; Functional
OPTION COURSES PREVIOUSLY OFFERED INCLUDE: Genomic Technologies; Bioinformatics Programming & System
Selected according to degree specialisation: Quantitative Genetic
OPTION COURSES PREVIOUSLY OFFERED INCLUDE: Management; Stem Cells & Regenerative Medicine; Statistics and
Models; Molecular Evolution; Genetics of Human Complex Traits; Animal Introduction to Scientific Programming; Commercial Aspects of Drug
Data Analysis; Biobusiness; Gene Expression & Microbial Regulation;
Molecular Phylogenetics; Bioinformatics; Molecular Evolution; Functional Genetic Improvement; Functional Genomic Technologies; Molecular Discovery; Stem Cells & Regenerative Medicine; Biological Physics;
Bioinformatics Algorithms; Biological Physics; Computational Cognitive
Genomic Technologies; Animal Genetic Improvement; Evolutionary Phylogenetics; Bioinformatics; Evolutionary Quantitative Genetics. Biochemistry; Enzymology & Biological Production; Next Generation
Neuroscience; Molecular Phylogenetics; Next Generation Genomics;
Quantitative Genetics. Genomics; Machine Learning & Pattern Recognition; Drug Discovery;
Career opportunities Biophysical Chemistry; Bioinformatics Programming & System
Drug Discovery; Biochemistry; Environmental Gene Mining &
Career opportunities You will develop the in-depth knowledge and specialised skills required
Management; Economics & Innovation in the Biotechnology Industry;
Metagenomics; Economics & Innovation in the Biotechnology Industry;
You will develop the in-depth knowledge and specialised skills required to apply quantitative genetics theory to practical problems, in both the Industry & Entrepreneurship in Biotechnology; Introduction to Scientific
BioBusiness; Molecular Modelling & Database Mining; Industry &
to apply quantitative genetics theory to practical problems, in both the biomedical and animal science industries, and to undertake research in Programming; Practical Skills in Biochemistry; Mathematical Biology.
Entrepreneurship; Practical Skills in Biochemistry; Practical Systems
biomedical and animal science industries, and to undertake research in evolutionary genetics, population genetics and genome analysis. Recent
Biology; Functional Genomic Technologies; Information Processing in Career opportunities
evolutionary genetics, population genetics and genome analysis. graduates have taken on roles as geneticists, DNA analysts, data analysts
Biological Cells; Data Mining & Exploration; Gene Expression & Microbial The programme is designed to give you a good basis for managerial or
and researchers.
Entry requirements Regulation; Bioinformatics; Principles of Industrial Biotechnology. technical roles in the pharmaceutical and biotech industries. It will also
A UK 2:1 honours degree, or its international equivalent (www.ed.ac.uk/ Entry requirements prepare you for entry into a PhD programme.
international/graduate-entry), in biological or biomedical sciences or A UK 2:1 honours degree, or its international equivalent (www.ed.ac.uk/
Career opportunities
medicine, with evidence of quantitative skills. We will also consider your international/graduate-entry), in biological or biomedical sciences,
You will enhance your career prospects by acquiring current, marketable Entry requirements
knowledge and developing advanced analytical and presentational A UK 2:1 honours degree, or its international equivalent (www.ed.ac.uk/
application if you have a background in mathematics, statistics or physics medicine, agriculture or animal sciences, with evidence of quantitative
skills, within the social and intellectual sphere of a leading international international/graduate-entry), with a strong background in one or more
and can show evidence of an interest in genetics. skills. We will also consider your application if you have a background in
university. The School of Biological Sciences offers a research-rich of the following areas: bioinformatics, computer sciences and modelling,
mathematics, statistics or physics and can show evidence of an interest
English language requirements environment in which you can develop as a scientist and entrepreneur. molecular biology and related sciences, engineering or biotechnology.
in genetics.
See page 20. We may also consider your application if you have a background in
English language requirements Entry requirements
Fees and funding physics or mathematics.
A UK 2:1 honours degree or its international equivalent (www.ed.ac.uk/
See page 20.
For fees see page 20 and for funding information see page 22. international/graduate-entry). You should have a strong background English language requirements
Fees and funding in one or more of the following areas: biotechnology, biochemistry, See page 20.
For fees see page 20 and for funding information see page 22. molecular biology and the related sciences. We may also consider your
Programme Director Professor Andrew Leigh Brown Fees and funding
Tel +44 (0)131 650 5523 application if you have a background in engineering (particularly
Email qgen@ed.ac.uk Programme Director Professor Andrew Leigh Brown bioengineering), physics or mathematics.
Tel +44 (0)131 650 5523 English language requirements Programme Director Professor Peter Swain
Email qgen@ed.ac.uk See page 20.
Tel +44 (0)131 650 5451
Fees and funding Email mscssb@ed.ac.uk
Programme Director Dr Louise Horsfall

Tel +44 (0)131 650 5363
Email mscsyntheticbiotech@ed.ac.uk
Online distance learning

www.ed.ac.uk/pg/834 www.ed.ac.uk/pg/766
Drug Discovery & Next Generation

Protein Biotechnology Drug Discovery
MSc 2–6 yrs PT MSc 2–6 yrs PT
PgDip/PgProfDev 2–4 yrs PT PgDip/PgProfDev 2–4 yrs PT
PgCert 1–2 yrs PT PgCert 1–2 yrs PT
CPD options CPD options
Programme description Programme description

The modern pharmaceutical industry encompasses the development This online programme will provide you with the stimulus, guidance and
of ‘biologics’ (for example antibodies or protein hormones), as much knowledge to develop a career around new approaches to drug discovery.
as it does traditional, small-molecule drug discovery.
You will study the challenges in developing novel drugs; the science
You will study the design and potential uses of different families of proteins underlying emerging fields of drug discovery; the application of new ideas
and will examine the experiences of successful entrepreneurs in the field to the field; how drug discovery relates to real-world health problems; the
who have been involved in the commercialisation of biopharmaceuticals. commercial aspects of drug discovery; and potential future developments.
Your research project will focus on the early phases of an industrial The programme offers a research-rich environment in which you can
biologics design programme. develop as a scientist and entrepreneur.
Programme structure Programme structure
You will learn through a variety of teaching methods, including online You will be taught through a variety of teaching methods, including online
tuition, peer-to-peer discussion and individual study. You will take six tuition, peer-to-peer discussion and individual study. For the MSc, you will
taught courses, which will be a mixture of compulsory and option courses, take six taught courses – a mixture of compulsory and option courses. In
followed by a research project leading to a dissertation in your final year. your final year, you will pursue a research project leading to a dissertation.
Individual courses can be taken for Continuing Professional Development You can take individual courses for Continuing Professional Development
purposes or you can study for a Postgraduate Certificate, Postgraduate purposes or study to qualify for a Postgraduate Certificate, Postgraduate
Diploma or MSc. The standard MSc duration is three years but we also Diploma or the MSc. The standard MSc duration is three years but we also
offer a fast track two-year option depending on your eligibility at the end offer a fast-track, two-year option depending on your eligibility at the end
of Year 1. Alternatively, you can spread your programme over a maximum of Year 1. Alternatively, you can spread your programme over a maximum
of six years, through intermittent study, allowing you to accommodate of six years through intermittent study, allowing you to plan around work
work and other commitments. You can expect to spend seven to 13 and other commitments. You can expect to spend between seven and
hours a week on your studies, depending on your chosen schedule. 13 hours a week on your studies, depending on your chosen schedule.
COURSES PREVIOUSLY OFFERED INCLUDE: COURSES PREVIOUSLY OFFERED INCLUDE:

Commercial Aspects of Drug Discovery; High Throughput Drug Commercial Aspects of Drug Discovery; Chemistry for Drug Discovery;
Discovery; In Silico Drug Discovery; Introduction to Modelling Biological Druggable Systems; High Throughput Drug Discovery; In Silico Drug
Systems; Measuring Drug Binding; Molecular Modelling; Professional Skills Discovery; Introduction to Modelling Biological Systems; Measuring
in Drug Discovery; Structure Determination of Drug Target; Systems Drug Binding; Modelling Metabolic Pathways; Molecular Modelling;
Approach to Modelling Cell Signal Transduction; Research Grant Proposal. Professional Skills in Drug Discovery; Structure Determination of Drug
Targets; Systems Approach to Modelling Cell Signal Transduction.
Career opportunities
You will enhance your career prospects with marketable analytical Career opportunities
and presentation skills. You will enhance your career prospects with marketable analytical
and presentation skills.
Entry requirements
A UK 2:1 honours degree, or its international equivalent (www.ed.ac.uk/ Entry requirements
international/graduate-entry), in biotechnology, biochemistry, chemistry, A UK 2:1 honours degree, or its international equivalent (www.ed.ac.uk/
medicine, molecular biology, pharmacology or related sciences. Other international/graduate-entry), in biotechnology, biochemistry, chemistry,
scientific backgrounds are considered on a case-by-case basis. We may medicine, molecular biology, pharmacology or related sciences.
also consider your application if you have another scientific background;
please contact us to check before you apply.
See page 20.
English language requirements Fees and funding
See page 20.
Fees and funding
Programme Director Dr Paul McLaughlin
Tel +44 (0)131 650 7060
Programme Director Dr Paul McLaughlin Email mscngdd@ed.ac.uk
Tel +44 (0)131 650 7060
Email mscddpb@ed.ac.uk
Research at the School of Biological Sciences Research opportunities

We’re one of the largest and most highly rated www.ed.ac.uk/pg/7 www.ed.ac.uk/pg/8
centres of our kind in the UK and one of the top
25 worldwide for life sciences*. Cell Biology Evolutionary Biology
With the majority of our research judged as either world leading
or internationally excellent in the Research Excellence Framework
(REF) 2014, and with a successful record of research grant
Case study: Edinburgh’s PhD 3 yrs FT (6 yrs PT available for UK/EU students) PhD 3 yrs FT (6 yrs PT available for UK/EU students)
applications, we can offer an environment that boasts cutting-edge research with impact MPhil 2 yrs FT (4 yrs PT available for UK/EU students) MPhil 2 yrs FT (4 yrs PT available for UK/EU students)
equipment and facilities to encourage research excellence and MSc by Research 1 yr FT (2 yrs PT available for UK/EU students) MSc by Research 1 yr FT (2 yrs PT available for UK/EU students)
innovation. You’ll be part of an active Graduate School and will
benefit from the support of around 130 principal investigators, New hope for Rett Research environment Research environment
syndrome sufferers
many of whom hold independent personal fellowships funded The Institute of Cell Biology (ICB), including the Wellcome Trust Centre As a research student at the Institute of Evolutionary Biology you will join
by prestigious bodies. Through our innovative skills database, for Cell Biology and its specialist PhD programme, hosts 38 laboratories the UK’s largest and most dynamic community of researchers in the field.
carrying out world-class research into fundamental mechanisms relating With about 30 research groups, which include about 50 PhD students,
you’ll have access to a broad range of expertise within our
to cellular function. We apply a wide range of approaches including you’ll benefit from a busy programme of seminars, journal clubs and
comprehensive research areas, the output of which has been Rett syndrome is a severe autistic-spectrum disorder with molecular genetics, cell and structural biology, systems biology other research-furthering activities and will be encouraged to publish
assessed as second in the UK for the volume of ‘internationally delayed onset that affects one in 10,000 girls, which includes approaches and mathematical modelling. your findings. You’ll also take part in student-led courses and talks.
excellent’ and ‘world-leading’ research. around 16,000 in the US, and an estimated 2,400 here in the
Our research groups comprise one of the largest communities of cell We have an enviable record in academic career success. Our institute
UK. This regressive disease causes loss of speech and hand
biology researchers in the UK. Many of our staff are Fellows of the Royal reflects the School-wide figure of 84 per cent for students achieving
Research routes movement, coupled with autistic behaviour, an undersized Society and acknowledged leaders in their field. Most PhD students who academic posts after they complete their PhD.
Our six discipline-based research institutes cover biology from brain (microencephaly), and growth retardation. A cure or train at the ICB continue with their academic career or join research in
molecular structure to evolutionary and population biology. therapy was thought to be most unlikely. However, leading We go beyond the study of animal behaviour to focus on the underpinnings
the industry, often becoming independent group leaders.
of the science of evolution, down to the level of DNA, to look at just how
geneticist Professor Sir Adrian Bird from the University’s
Diversity of interests organisms fundamentally develop as a result of natural selection.
PhD School of Biological Sciences may have found the answer We offer a wide scope of projects covering different areas of cell biology,
As a PhD candidate you pursue a research project under continuous to this devastating disease. Our current research falls under three broad themes: evolutionary
including the synthesis, processing, localisation and degradation of RNA;
guidance, resulting in a thesis that makes an original contribution to and quantitative genetics, evolutionary ecology and behaviour, and
epigenetic control of gene expression, chromosome function and genome
biodiversity and ecology. Researchers are working on everything from
the sector. You will gain specialist background knowledge for your Project background stability; mechanisms of cell growth and duplication; the rules that govern
viral evolution and host-parasite co-evolution to natural selection in the
intended research and develop the skills to research in that field. The syndrome was previously believed to be a developmental cellular architecture; biotechnology; synthetic biology; and microbial
wild and plant conservation genetics.
We offer two routes to a PhD: a four-year programme that includes or neurodegenerative disease because of its early appearance regulation. A variety of courses in microscopy, bioinformatics and
a substantial training element and a three-year programme for and the gradual deterioration of those affected. However, proteomics are available to all our PhD students. Facilities
students with a strong background in their chosen area. Topping our list of impressive research tools is Edinburgh Genomics,
leading-edge research led by Professor Bird has presented the Facilities one of the UK’s biggest and best university-based genomics facilities.
very real prospect of a future cure. By developing a genetic The ICB provides an exceptional working environment. The Wellcome Trust
Attracting visiting researchers from all over the UK and internationally,
MPhil mouse model for Rett syndrome that mimics the genetic Centre for Cell Biology is home to the Central Optical Instrumentation
this next-generation facility opens up new ways of creating research
The Master of Philosophy (MPhil) resembles a PhD but generally mutation that causes its symptoms, Professor Bird opened up Laboratory (COIL), while image analysis at the boundary of biology and
data. If your study involves bioinformatics, the Ashworth Bioinformatics
takes two years and does not carry the same requirement for a new avenue of research. In 2007, Dr Jacky Guy and other physics is provided through the nearby Collaborative Optical Spectroscopy
Support Service provides a vital link with the expertise and facilities on
original contribution to knowledge. You’ll pursue your individual scientists in Professor Bird’s team introduced a modified Micromanipulation & Imaging Centre (COSMIC). The ICB is proud of its
offer through our world-leading School of Informatics.
research project under supervision and submit a thesis. cutting-edge proteomics facility and automated drug-screening platforms
MeCP2 gene into Rett model mice, which allowed controlled English language requirements
that facilitate discovery of new molecules that impact on cellular functions.
expression of normal MeCP2 protein – a lack of which had See page 20.
MSc by Research been determined to be the cause of Rett syndrome. English language requirements
An MSc by Research is based on a research project tailored to See page 20. Fees and funding
your interests. It lasts one year full time or two years part time. Mutant female mice carrying this modified gene exhibited Fees and funding
The project can be a shorter alternative to an MPhil or PhD, or the characteristics of Rett syndrome until normal MeCP2 For fees see page 20 and for funding information see page 22.
a precursor to either – including the option of an MSc project expression was activated, after which they rapidly regained
expanding into MPhil or doctorate work as it evolves. normal behaviour. This striking result indicated that the
developmental or degenerative changes seen in Rett
Entry requirements patients are reversible, and overturned previous
A UK 2:1 honours degree, or its international equivalent, in a understanding of the disease.
relevant subject. You should have some research experience and
be able to demonstrate that you have a good understanding of Project results
the field you propose to study. These potentially life-changing findings by the research
team at the University have inspired worldwide awareness
High rate of career success campaigns, a documentary, and fundraising programmes
We find the overwhelming majority of our students have an aimed at supporting further research. They underpin the
academic career in mind when applying for research degrees. rationale of multiple clinical trials now under way in both
Our latest figures show that 84 per cent of our graduates enter Europe and the US to test both symptom-relieving drugs
an academic research or teaching career on graduation, and are and gene therapy to combat the underlying cause.
now establishing themselves in universities and other research
institutions worldwide. It’s a significant achievement, one that
demonstrates the quality of both our research and our standing
Leading-edge research led by
within the international academic community. We also fully support Professor Bird has presented the
our students who plan to develop careers outside academia. very real prospect of a future cure.
* T imes Higher Education World University Rankings 2014/15
See more online: www.ed.ac.uk/research/impact
Immunology & Molecular Plant Sciences Quantitative Biology, Stem Cell Research
Infection Research Biochemistry & Biotechnology
PhD 3 yrs FT (6 yrs PT available for UK/EU students) PhD 3 yrs FT (6 yrs PT available for UK/EU students) PhD 3 yrs FT (6 yrs PT available for UK/EU students) PhD 3 yrs FT (6 yrs PT available for UK/EU students)
MPhil 2 yrs FT (4 yrs PT available for UK/EU students) MPhil 2 yrs FT (4 yrs PT available for UK/EU students) MPhil 2 yrs FT (4 yrs PT available for UK/EU students) MPhil 2 yrs FT (4 yrs PT available for UK/EU students)
MSc by Research 1 yr FT (2 yrs PT available for UK/EU students) MSc by Research 1 yr FT (2 yrs PT available for UK/EU students) MSc by Research 1 yr FT (2 yrs PT available for UK/EU students) MSc by Research 1 yr FT (2 yrs PT available for UK/EU students)
Research environment Research environment Research environment Research environment

Immunology and infection research has a proud tradition here at the With world-class facilities and lively collaborations, our Institute of Closely allied with the Institute of Cell Biology, our group studies the Our role in the development of stem cell research began in the early 1990s,
School of Biological Sciences. As a researcher, you’ll be following in the Molecular Plant Sciences provides you with the ideal environment in structure and impact of biomolecules, from their atomic structure and with researchers at the former Centre for Genome Research producing the
footsteps of Nobel Prize winners – from malaria pioneer Sir Ronald Ross which to pursue postgraduate research. We’re one of the most active assembly into molecular machines, to studying how molecular signals are subject’s early publications. We grew into the first stem cell research
to the 2001 laureate Sir Paul Nurse – and other great names who have university departments for plant sciences in the UK, hosting one of the transduced in animals and plants. We offer a combination of world-class institute in the UK, and this formed the basis of what is now a world-leading
made groundbreaking discoveries over the years. Our leadership in country’s few plant-based research and teaching institutes. Each of our facilities and training that sets us apart as one of the premier centres for this centre for multidisciplinary research in mammalian stem cell biology
infectious disease research has emerged alongside the School’s strength 11 research groups makes a vital contribution to new discoveries. You’ll type of research in the UK. You’ll be supported by well-published staff who and regenerative medicine. As a PhD student, you’ll study in a unique
in population biology and quantitative genetics; our close association be able to choose from a broad range of topics, applying multidisciplinary are leaders in their field and will benefit from a comprehensive programme environment, thanks to our location within the MRC-funded Scottish
with biologists in these areas can be a valuable asset to your work. approaches – from computational biology to molecular genetics and cell of seminars, symposiums, distinguished visitors and social events. Centre for Regenerative Medicine (SCRM). This unique cross-disciplinary
biology – and to subjects ranging from plant growth, development and initiative is aimed at studying the properties and behaviour of stem cells
The Institute of Immunology & Infection Research has more than 20 Finding answers
evolution to plant pathogen interactions and biotechnology. to develop new therapies.
research group leaders, including seven professors. While genetics is We investigate molecular structure using techniques such as biomolecular
a core discipline, our work extends over molecular and cell biology, Broad scope nuclear magnetic resonance, X-ray crystallography, computational Field of focus
immunology, evolutionary biology and epidemiology. We offer a wide range of research interests, including plant biotechnology, structure prediction and cryo-electron microscopy. We also cover the full Research groups within SCRM cover a diverse range of areas, exploring
developmental biology; speciation and natural variation (making use of our panoply of methods for protein production and biological characterisation. the possibilities offered by embryonic, fetal and adult tissue stem cells.
Themes
strong connections with the Royal Botanic Garden Edinburgh); circadian Each group focuses on a highly specific aspect of research. The themes
We cover a number of overlapping themes of study. In fundamental Our labs are highly active in deciphering the molecular events that drive
clocks and photobiology; bio-imaging; biochemistry and cell biology; we address include: the properties of stem cells and their niches; control
immunology, we look at how B and T lymphocytes interact and develop growth and development. By combining experimental and mathematical
and molecular plant pathology. of stem cell self-renewal and differentiation in vitro and in vivo; the role
in response to antigen challenge, and how responses are initiated by modelling approaches we are able to gain a more holistic ‘systems level’
of stem cells in the ageing process; and the mechanisms underlying tissue
dendritic cells and dampened by regulatory cells. Helminth, allergy and Facilities understanding of molecular signalling.
degeneration and repair. The Centre’s overarching goal is to harness this
wound repair models are used to understand immune responses in both Substantial investment has been made in ensuring the equipment and
A growing area of our research is the development of technologies that knowledge to develop new therapies. We attract highly committed and
practical and evolutionary terms. Immune regulation of disease projects facilities you’ll use are as good as those anywhere in the world. You’ll
combine chemical library generation with proteomics and high-speed motivated students and reward them with an interdisciplinary research
aims to design immunological interventions to ameliorate pathology or work in purpose-built laboratories and specialist facilities for plant
imaging methods to allow identification of new ligand-protein interactions. environment that fosters achievement at the highest levels.
to enhance host immunity, in conjunction with new vaccines against growth, including a high-throughput Arabidopsis genetics facility, a suite
Through links with the College of Medicine & Veterinary Medicine, we aim
parasites and therapies for autoimmunity. of biochemistry equipment, controlled environment growth rooms and
to make a real impact on healthcare through drug discovery projects.
Facilities
glasshouses, including a modern category three containment glasshouse. Since 2011, SCRM has been housed in a spectacular, specially designed
Work on molecular biology and genetics of parasites focuses on the
Our strong links with the Centre for Synthetic and Systems Biology, Facilities building that provides high-quality research facilities, including:
identification of vaccine candidates and virulence factors in malaria
SynthSys, and the Royal Botanic Garden Edinburgh give you access Your work will benefit from our strong links with a number of on-site
parasites, the cell biology of trypanosomes and proteomics and • a state-of-the-art centralised cell culture facility for isolation and
to an even broader range of facilities and expertise. research centres, including the Wellcome Trust Centre for Cell Biology,
transcriptomics of helminth worm parasites. culture of primary and established cell lines including embryonic
SynthSys, the Centre for Synthetic and Systems Biology, the Centre for
In molecular microbiology, we look at how microRNAs regulate immune
English language requirements Science at Extreme Conditions, and the Centre for Translational and
and induced pluripotent stem cells;
See page 20.
signalling and how pathogens manipulate these processes. Host-pathogen Chemical Biology. We offer outstanding laboratories, equipment and • a clinical-grade GMP cell culture facility;
population biology integrates conventional immunology, pathogen Fees and funding training, particularly in the areas of protein production and characterisation,
• a Specific Pathogen Free animal facility;
research and systems-level quantitative biology, using mathematical For fees see page 20 and for funding information see page 22. cryo-EM and macromolecular X-ray crystallography, mass-spectrometry,
approaches in both experimental models and the epidemiology of robotised qPCR, real-time bioluminescent imaging and plant phenotyping. • a transgenic service covering derivation and provision of mouse
human and animal infections. embryonic stem cells, blastocyst injection, morula aggregation
English language requirements and production of defined genetic alterations;
Facilities See page 20.
Our institute offers comprehensive facilities, including tissue-culture • ultrasound micro-injection equipment;
Fees and funding
facilities, state-of-the-art 17-colour flow cytometry and five-dimensional • a flow cytometry service consisting of cell sorters, MoFlo, FACS Jazz
confocal microscopy, as well as equipment such as fast protein liquid and FACS Aria II, operated by facility staff and analysers,and the LSR
chromatography, phosphorimaging and real-time polymerase chain Fortessa and FACS Calibur that can be operated by users following
reaction. There is also ready access to molecular technologies such mandatory training;
as automated DNA sequencing, DNA arrays and mass spectrometry The University is
on the King’s Buildings campus. piloting PhDs by distance • histology;
learning. If you’re interested • imaging facilities including standard compound microscopy, confocal,
English language requirements in studying with us this way,
See page 20. STED super-resolution, high-content and timelapse imaging;
we’re keen to investigate
Fees and funding possibilities in some of our • quantitative real-time polymerase chain reaction capabilities; and
For fees see page 20 and for funding information see page 22. areas of research. • Fluidigm Biomark and CellPrep for single cell transcriptomics.
Your colleagues will include members of the University’s College of
Medicine & Veterinary Medicine who can provide a clinical perspective and,
as you’ll be located a stone’s throw from the Royal Infirmary of Edinburgh,
“I completed a PhD in Structural Biology at the University you’ll also have access to hospital facilities and practising clinicians.
of Edinburgh. I originally joined the Walkinshaw group English language requirements

See page 20.
as a bioinformatics PhD candidate but through Professor See also… Fees and funding
Walkinshaw’s supportive and motivational mentorship took up Much of our research is interdisciplinary and collaborative. For fees see page 20 and for funding information see page 22.
the exciting challenge of X-ray crystallography, which I have You may find your preferred research area in the prospectus
of the College of Medicine & Veterinary Medicine, or those
continued to investigate through a postdoctoral fellowship of the Schools of Chemistry, Informatics or Engineering.
at the University of British Columbia, Vancouver, Canada.” www.ed.ac.uk/studying/prospectus-request
Liam Worrall, PhD Structural Biology
About the School of Facilities and resources

Biological Sciences Whatever tools you need to conduct your research,
you’ll find the latest at the School of Biological Sciences.
Our School features a vibrant community of staff and students Our Edinburgh Genomics sequencing
that continues a 400-year history of scientific exploration with service leads the way, playing host to the
Collections of the University
The University of Edinburgh has one of
innovative work that seeks to shape tomorrow’s world. National Environment Research Council
the world’s great collections, which has
National Sequencing Facility and Medical
been growing ever since its foundation
Research Council Hub Facility.
in 1583. Our collections include rare
books, archives and manuscripts, art,
We are constantly seeking to shed light on Research institutes Advanced microscopy and flow cytometry
the secrets of life through our wide-ranging historical musical instruments and a
Our world-class research takes place are two of our major strengths. Our
research. More than half of our research wide range of museum objects from
in six research institutes: microscopy facilities house state-of-the-art
was assessed as world-leading in the geological specimens to anatomical
widefield and confocal fluorescence, and
Research Excellence Framework (REF) models. If laid out end to end, we
• The Institute of Cell Biology transmission and scanning electron
2014 placing us in the UK’s top three for would have almost 60 kilometres of
• The Institute of Evolutionary Biology microscopes. Our flow cytometry
overall research quality and confirming shelving and storage space devoted to
facilities offer cutting-edge analytical
our position as one of the world’s leading • The Institute of Immunology and our heritage material, from 1st-century
and sorting capabilities.
biological science research groups. Infection Research Greek papyrus fragments to new works
of sculpture. This is curated by specialist
• The Institute of Molecular Plant Sciences Extensively equipped
Through our membership of Scottish staff across 45 sites and used for our
We have protein production and biophysical
Universities Life Sciences Alliance, we • The Institute for Stem Cell Research teaching and research and by the wider
characterisation facilities, including surface
work with colleagues in other institutions • The Institute of Quantitative Biology, public community.
plasmon resonance, ITC and spectroscopic
to maintain the country’s world-class Biochemistry and Biotechnology. equipment. We host more than 1,000m2
research base. Our commitment to The Centre for Research Collections in
of controlled environment growth space
pioneering science, supported by a Many of our researchers also participate the Main Library is the hub for all our
and glasshouse space, and have computing
healthy flow of grant funding, helps create in one or more of our numerous collections, where specialist curators
resources for evolutionary biology, including
an environment where world firsts are cross-disciplinary research centres. make them available for study, research
four dedicated computer clusters for
possible. For example, the first genetically and pleasure. Postgraduate students
phyloinformatics, comparative genomics,
engineered vaccine against hepatitis B was Programmes are welcome to study original objects
population genetics and quantitative trait
developed here at the School of Biological Postgraduate teaching takes every and have made many important
locus genetics.
Sciences. By joining us you will experience advantage of our School’s expertise to research discoveries while working on
the archives. You will find an incredible
a unique opportunity to examine life give you outstanding opportunities to Excellent facilities
processes at the very highest level. study within your chosen field, from range of material in our collections that
We are based almost wholly at the
programmes taught at the world-famous is available nowhere else in the world.
University’s King’s Buildings campus, about
Thriving community Royal Botanic Garden Edinburgh to those 15 minutes by bus from the city centre.
Our School houses about 130 principal that develop biofuels and new medicines. You can take advantage of the Noreen and
investigators, both academic teaching Our students can be found discovering Kenneth Murray Library, named after the
staff and independently funded senior how to develop new products for human pioneers of the first genetically engineered
research fellows, about 400 research and animal health, or applying the latest hepatitis B vaccine, as well as the KB
assistants and technicians and more engineering principles to the exploration Learning and Teaching Cluster and the
than 200 PhD students. of biological networks. social and sports facilities at KB House.
All postgraduate research students are
Some 90 administrative and technical staff Effective outcomes given their own desk space in shared
support the School’s academic activities. The School of Biological Sciences student offices.
continues to make its mark on the future.
Entrepreneurial opportunities are supported
by Edinburgh Research & Innovation (ERI),
the University’s commercialisation office,
with a dedicated business development
team embedded in the School. Across the
University, ERI has helped create more than
The School of Biological 400 companies during the last 50 years.
Sciences has held an Athena
Swan Silver Award since 2013
for the advancement of
women in science.
www.athenaswan.org.uk “I was very excited to be able to attend one of the best
universities in the world. The programme is intensive and
the professors kept us busy. Sometimes I struggled, but I was
happy because I gained so much as a student here.”
Stamatina Fragkogianni, MSc Bioinformatics
Community Employability and

The School forms one of the largest academic groupings of biological
graduate attributes
scientists in the UK. Whether you are following a taught postgraduate
programme or embarking on a research degree, you will be welcomed We offer a research-rich environment in which to develop as a scientist
into a supportive and enthusiastic community of students and staff. or entrepreneur and you will gain skills that benefit your personal and
professional development whichever direction your career takes. The
University provides a range of services and opportunities to help you make
The MSc experience reflects the importance we place on minimum of 34 fully funded PhD the most of your time here and the School offers professional internship
As an MSc student, there is always the collaborative approach to research. studentships annually and provides schemes to MSc and PhD students.
something to keep you motivated and Through these connections, you’ll come enhanced training to its students.
inspired – from the University’s Festival into contact with researchers from varied
of Creative Learning to the Edinburgh academic backgrounds and gain insights More information:
International Science Festival. Each of into new approaches and techniques. www.eastscotbiodtp.ac.uk Professional internships plus online advice and learning materials. We sustain and continually develop links
our MSc programmes has a dedicated We are linked with: We offer EASTBIO PhD students the Workshops and learning resources cover key with employers from all industries and
administrator who is there to help you The Innovation Forum opportunity to pursue a professional topics tailored to different academic stages, employment sectors, from the world’s top
with any queries. • Edinburgh Bioinformatics Edinburgh hosts a local branch of the internship, supported by a Postgraduate including getting started with your studies; recruiters to small enterprises based here
• The Centre for Translational & Chemical Innovation Forum, a global network of Placements Coordinator. critical reading, writing and thinking; in Edinburgh. Our employer team provides
Research support Biology researchers and entrepreneurs active at managing your exams; and planning a programme of opportunities for you to
All postgraduate research students are the universities of Cambridge, Oxford, All EASTBIO students spend three months for and writing up your dissertation. meet employers on campus and virtually,
members of our Graduate School, which • Edinburgh Infectious Diseases London, Edinburgh, Copenhagen, Lausanne, of their PhD programme pursuing a and advertises a wide range of part-time
enjoys an active academic and social • The MRC Centre for Regenerative Barcelona, Hong Kong and Tokyo. This is a non-academic internship. This is designed IAD also provides a comprehensive and graduate jobs.
calendar. We provide every opportunity Medicine student-led initiative seeking to promote to help you develop a broad range of programme of transferable skills training,
to join in – from peer support groups and • SynthSys – Centre for Synthetic and innovation by building bridges between professional skills. resources and support for researchers More information:
journal clubs to BioDocSoc, our society Systems Biology academia, industry and government, and completing a doctorate. The workshop www.ed.ac.uk/careers/postgrad
run by and for research students and staff. linking innovative minds across disciplines. Science communication programme is designed to help you
• The Wellcome Trust Centre for Cell
There are plenty of opportunities to successfully prepare for the various Connect.ed
Inspiring environment Biology More information: develop science communication skills. milestones of your PhD, from getting Edinburgh encourages its alumni to stay
Our environment of shared knowledge and • The Wellcome Trust Centre for Infection, www.inno-forum.org The student-run BioPOD series of podcasts started with your research, to writing up in touch with current students who share
expertise has led to groundbreaking and Immunity & Evolution. covers stories from across the School – you and preparing for the viva. Workshops an academic background or are interested
globally recognised research achievements. might find yourself the subject of one, or cover topics such as writing skills, in a similar career path. Connect.ed is a
Our research institutes provide a forum for Many of our research projects also involve helping to produce them. Similarly, our reference management tools, statistics, networking system run by the Careers
development of ideas, collaboration and collaborations with Scotland’s Rural Press Gang works with the University’s preparing for conferences, delivering Service that provides an informal and
dissemination of results, along with an College, the Royal Botanic Garden Press Office and the Scottish Initiative for presentations, time and project confidential opportunity for alumni to
environment for training and mentoring Edinburgh and many other academic Our research Biotechnology Education to spread the management, and personal development. share their occupational knowledge and
research students and early career and commercial organisations. students produce word on the School’s pioneering work. IAD also offers online resources and experience with current students, who
researchers. BioSkills, a database developed regular podcasts about Many of our students are part of the planning tools to help get your research can contact them for advice and guidance
by our own researchers, makes it easy to The School of Biological Sciences is lead School news and events. EUSci team, publishing a regular science started, plus support for tutoring and on their future career.
identify fellow researchers in Edinburgh who partner in the EASTBIO Doctoral Training www.ed.ac.uk/ magazine, and there are also opportunities demonstrating and research public
can share their skills and expertise with you. Partnership, a collaboration between biology/biopod
to get involved with the annual Edinburgh engagement and communication. More information:
the universities of Aberdeen, Dundee, International Science Festival. www.ed.ac.uk/careers/connected
Links and partnerships Edinburgh and St Andrews. Funded by the Careers Service
Our participation in the work of a wide Biotechnology and Biological Sciences Institute for Academic Development Our Careers Service plays an essential part Backing bright ideas
variety of interdisciplinary research centres Research Council, EASTBIO awards a All postgraduate students can benefit in your wider student experience at the LAUNCH.ed is the University’s award-
from our Institute for Academic University, offering world-class careers winning programme for student
Development (IAD), which provides and personal development guidance and entrepreneurs. Each year, LAUNCH.ed
information, events and courses to support. We support you to recognise the works with hundreds of students to assess
develop the skills you will need throughout wealth of possibilities ahead, while at their ideas and develop their business skills
your studies and in the future. IAD events university and after graduation, helping and helps many start their businesses.
also offer the perfect opportunity to meet you explore new avenues, tap into your We have helped Edinburgh students and
and network with other postgraduates talents and build your employability with alumni launch almost 100 new businesses
from across the University. confidence and enthusiasm. in the last three years, ranging from
language tuition to robotics companies.
Further information is available online: The Service provides specialist support
www.ed.ac.uk/iad/postgraduates for postgraduate students. From exploring More information:
career options to making decisions, from CV www.LAUNCH.ed.ac.uk
For taught postgraduates, IAD provides writing to interview practice, from Employ.
a popular study-related and transferable ed internships to graduate posts and from
skills support programme. It is designed careers fairs to postgraduate alumni events,
to help you settle into postgraduate life, we help you prepare for the future.
succeed during your studies and move
confidently to the next stage of your career.
We offer on-campus and online workshops
and one-to-one study skills consultations,
Applications and fees

We have an online application process for all postgraduate programmes.
It’s a straightforward system with full instructions, including details
of any supporting documentation you need to submit.
When applying, you will set up an account, Research applications Tuition fees The University is working with the
which lets you save your application and For funded studentships our main The following table provides an overview Scottish Government to try to protect
continue at another time. application deadline is usually around of indicative fee levels for programmes the fee status of EU students enrolling
December. Occasionally funded commencing in 2017. in the 2017/18 academic year for the
Full guidance on our application system is studentships are advertised later in the duration of their course. However there
available at: www.ed.ac.uk/postgraduate/ academic year. Please check our website Figures marked * show the fee level set is a risk that EU students enrolling in the
applying (www.ed.ac.uk/biology/postgraduate) for the 2016/17 academic year. All other 2017/18 academic year may become
for full deadline details. We accept figures are indicative of expected fee subject to international tuition fees for
General requirements applications from students with their levels for your studies during the 2017/18 any years of study which follow the UK
Our usual entrance requirement for own funding all year round. academic year. Because these figures are exit from the EU. In those circumstances
postgraduate study is a UK 2:1 degree, or indicative, it is important you check online we are committed to working with the
its international equivalent (www.ed.ac.uk/ English language requirements before you apply and check the up-to-date Government to ameliorate the impact
international/graduate-entry), in a subject Students whose first language is not fee level that will apply to your specific of that change for individual students.
related to your chosen programme. You English must show evidence of one programme: www.ed.ac.uk/student-
will also need to meet the University’s of the qualifications below: funding/tuition-fees/postgraduate The Scottish Government has already
language requirements (see below). confirmed that the fee status of existing
• IELTS Academic: total 6.5 (at least Please note: students and students enrolling in the
Entry requirements for individual 6.0 in each module). • International students starting full-time 2016/17 academic year will remain
programmes can vary, so check the • TOEFL-iBT: total 92 (at least 20 in taught programmes of study lasting unchanged for the duration of their studies.
details for the specific programme each module). more than one year will be charged
you wish to apply for. a fixed annual fee. For UK/EU students
• PTE(A): total 61 (at least 56 in each Annual fee
of the Communicative Skills sections). • All other students on full-time and Taught programme 1-year FT £13,800
References part-time programmes of study lasting
For applications to taught programmes, • CAE and CPE: total 176 (at least 169 Taught programme 2-years PT £6,900
more than one year should be aware
the normal requirement is one reference, in each module). MSc by Research 1-year FT £7,400
that annual tuition fees are subject to
although an additional reference may MSc by Research 2-years PT £3,700
• Trinity ISE: ISE II (with distinctions in revision and are typically increased by
be requested in individual cases. For MPhil 2-years FT £4,121*
all four components). approximately five per cent per annum.
applications to research programmes, two PhD 3-years FT £4,121*
This annual increase should be taken
references are required. You should check PhD 6-years PT £2,061*
Please note: into account when you are applying
the entry online for exact requirements
• English language requirements can be for a programme.
for your intended programme of study. Online Distance Learning
affected by government policy so please • In addition to tuition fees, your
For general guidance on references, visit: Annual fee
ensure you visit our degree finder to programme may be subject to an
www.ed.ac.uk/postgraduate/references £16,700
check the latest requirements for your application fee and additional costs/
programme: www.ed.ac.uk/ programme costs may apply. Please check
Deadlines For international students
postgraduate/degrees the latest programme information online.
Taught masters applications Annual fee
Some programmes have application • Your English language certificate must Taught programme 1-year FT £27,400
deadlines. Please check the programme be no more than two years old at the Tuition fees for EU students MSc by Research 1-year FT £23,700
beginning of your programme. EU students enrolling in the 2017/18
entry online for details. For all other MPhil 2-years FT £19,100
academic year – and possibly the following
programmes, you are encouraged to • We also accept recent degree-level study PhD 3-years FT £19,100
academic year – will be admitted as
apply no later than one month prior to that was taught and assessed in English
Scottish/EU fee status students and are
entry to ensure there is sufficient time to in a majority English speaking country * Figure shown is the 2016/17 fee level
eligible for tuition fee support from the
process your application. However, earlier (as defined by UK Visas & Immigration). All other fees quoted are indicative of 2017/18 fee
Student Awards Agency for Scotland (SAAS).
application is recommended, particularly levels. Because these figures are indicative, it is
important you check online before you apply and
where there is a high demand for places or Abbreviations: IELTS − International English
Future changes to the fee status of check the up-to-date fee level that will apply to your
when a visa will be required. Should you Language Testing System; TOEFL-iBT − specific programme: www.ed.ac.uk/student-funding/
EU students enrolling in the 2017/18
wish to submit a late application, please Test of English as a Foreign Language tuition-fees/postgraduate
academic year will depend on the timing
contact us for guidance. If you are Internet-Based Test; PTE(A) – Pearson Test
and terms of the UK’s exit from the
applying for funding we encourage you of English (Academic); CPE – Certificate of
European Union and would also require
to submit your programme application Proficiency in English; CAE – Certificate in
changes to existing UK and Scottish
as early as possible, as in most cases you Advanced English; Trinity ISE – Integrated
legislation. Current indications are that
will need a programme offer before you Skills in English.
the UK would leave the EU at the earliest
can make your funding application. Most
in 2019 so any changes would not take
funding deadlines are no later than June. www.ed.ac.uk/english-requirements/pg
effect before the academic year 2019/20.
Funding
A large number of scholarships, loans and other funding schemes are available
for your postgraduate studies. It is only possible to show a small selection in print.
To see the full range, please visit: www.ed.ac.uk/student-funding/postgraduate.
Awards are offered by the School of Scholarships at the • Highly Skilled Workforce Research council awards Loans available for study at Other sources of funding
Biological Sciences, the University of University of Edinburgh Scholarships Research councils offer awards to masters, the University of Edinburgh The following are examples of the many
Edinburgh, the Scottish, UK and international A number of scholarships are available to MPhil and PhD students in most of the The University of Edinburgh is a scholarships and support schemes available
governments and many funding bodies. • Beit Trust UK nationals permanently domiciled in Schools within the University of Edinburgh. participating institution in the following to students from particular countries who
Beit Trust and the University of Scotland, and to EU nationals domiciled All studentship applications from the loans programmes, meaning we certify meet certain eligibility criteria.
Here we list a selection of potential sources Edinburgh Scholarships jointly fund either on mainland EU or in Scotland, research councils must be made through your student status and can help with
of financial support for postgraduate postgraduate students from Malawi, who have been accepted on an eligible the University, through your School or the application process. • Chevening Scholarships
students applying to the School of Zambia and Zimbabwe to undertake full- or part-time masters programme. College office. Awards can be made for A number of partial and full funding
Biological Sciences. a masters: www.beittrust.org.uk The scholarships, which are funded by both taught and research programmes. • The Canada Student Loans scholarships are available to one-year
the Scottish Funding Council and subject Program masters students: www.chevening.org
• China Scholarships Council/University
Tuition fee discounts of Edinburgh Scholarships (China)
to annual confirmation, cover the UK/EU Normally only those UK/EU students The University is eligible to certify • Commonwealth Scholarships
We offer a 10 per cent discount on tuition fee: who have been resident in the UK for the Canadian student loan applications:
A number of scholarships for PhD study Scholarships available to students who are
postgraduate fees for all alumni who have www.ed.ac.uk/student-funding/sfc-hsw preceding three years are eligible for a full www.ed.ac.uk/student-funding/
to candidates who are citizens and resident in any Commonwealth country,
graduated with an undergraduate degree • Julius Nyerere Masters award. For some awards, candidates who canadian-loans
residents of China: www.ed.ac.uk/ other than the UK: www.dfid.gov.uk/cscuk
from the University. We also offer a 10 per Scholarship (Tanzania) are EU nationals and are resident in the
student-funding/china-council • Erasmus+ • F ulbright Scholarships (USA)
cent discount for international graduates One scholarship is available to citizens of UK may be eligible for a fees-only award.
• Edinburgh Global Research An Erasmus+ loan supports students Scholarships open to US graduate
who spent at least one semester at the Tanzania who are normally resident in www.ed.ac.uk/student-funding/
Scholarships accepted for a masters programme in an students in any subject wishing to
University of Edinburgh as a visiting Tanzania who are accepted on a research-councils
These scholarships are designed to attract Erasmus+ country. For more information: study in the UK: www.iie.org/fulbright
undergraduate: www.ed.ac.uk/student- full-time masters programme: www.
high-quality international research http://ec.europa.eu/education/
funding/discounts ed.ac.uk/student-funding/nyerere The University also offers a number • M
arshall Scholarships (USA)
students to the University: www.ed.ac.uk/ opportunities/higher-education/
of scholarships in partnership with the mastersloans_en.htm Scholarships available to outstanding
student-funding/global-research • School of Biological Sciences following overseas government agencies: US students wishing to study at any
Key Research Scholarships • Postgraduate Loans (PGL)
• Edinburgh Syrian Postgraduate UK university for at least two years:
Taught masters programmes Within the School of Biological Sciences, • Chile England
Scholarships www.marshallscholarship.org
Masters by Research programmes a number of highly competitive research National Commission for Scientific and Eligible students from England,
A number of scholarships are available • Scotland’s Saltire Scholarships
Research programmes scholarships are available each year to undertaking a taught or research
to postgraduate students from Syria Technological Research (CONICYT):
new postgraduate research students, masters, can apply to Student Finance A number of scholarships open to students
studying a full-time one-year masters: www.conicyt.cl
including funding from BBSRC, NERC, England for a loan of up to £10,000 who are citizens permanently and
www.ed.ac.uk/student-funding/ • Colombia ordinarily resident in Canada, China,
MRC, Wellcome Trust and our School towards fees or maintenance costs:
postgraduate/syria Administrative Department of Science, India, Pakistan and the USA for one year
International Scholarships: www.ed.ac.uk/ www.gov.uk/postgraduate-loan
biology/postgraduate/pgr/fees-funding Technology and Innovation (Colciencias): of masters study: www.ed.ac.uk/student-
www.colciencias.gov.co • Postgraduate Loans (SAAS) funding/saltire
• School of Biological Sciences Scotland and EU
Taught Postgraduate Bursaries • Ecuador The Student Awards Agency Scotland
Secretaria Nacional de Educacion Funding for online distance learning
The School of Biological Sciences offers offers tuition fee loans to eligible
Superior, Ciencia y Tecnologia (SENESCYT): The University offers several scholarships
awards to overseas students joining students undertaking full- or part-time
www.educacionsuperior.gob.ec specifically for online, part-time
taught masters programmes: postgraduate study. For a full list of postgraduate programmes, including the
www.ed.ac.uk/student-funding/ • Iraq eligible programmes: www.saas.gov.uk Edinburgh Global Online Distance Learning
biological-bursaries Ministry of Higher Education and • U
S Student Loans Masters Scholarship, for which applicants
• The University of Edinburgh Scientific Research: The University is eligible to certify loan to many masters programmes can apply:
PhD Scholarships www.iraqiculturalattache.org.uk applications for US loan students. Full www.ed.ac.uk/student-funding/e-learning/
A number of scholarships, open to UK, EU • Mexico details on eligibility and how to apply online-distance
and international PhD students: National Council of Science and can be found online: www.ed.ac.uk/
www.ed.ac.uk/student-funding/ Technology of the United Mexican student-funding/us-loans
development States (CONACYT):
www.conacyt.mx
Banco de Mexico and the Banco de
Mexico’s FIDERH trust (FIDERH):
“The Scottish Government’s initiative to attract international www.fiderh.org.mx
students from Canada, China, India and the US through

Fundacion Mexicana para la Educacion,
la Tecnologia y la Ciencia (FUNED):
the Saltire Scholarship Scheme, as well as the University www.funedmx.org
of Edinburgh’s help and support for international students,
has helped provide me with an opportunity that I would never
have conceived of prior to starting my studies at Edinburgh.”
Robert Starr, MSc High Performance Computing, Scotland’s Saltire Scholarship
Campus map Get in touch

The School of Biological Sciences is on the King’s Buildings Detailed maps Contact us
can be found at: For more information about taught
campus, where our teaching and administration takes place
www.ed.ac.uk/maps MSc programmes, contact:
across several buildings. The King’s Buildings campus is around
two miles from Edinburgh city centre and is well served by buses, Vicky Mactaggart
including a free University shuttle service during semester time. Biology Teaching Organisation
2105, James Clerk Maxwell Building
King’s Buildings
Mayfield Road
Edinburgh, UK
EH9 3JZ
Tel +44 (0)131 651 7052
Email pgtbiol@ed.ac.uk
www.ed.ac.uk/biology/taught-masters
For enquiries relating to specific

programmes, please refer to the contact
details provided for each programme listing.
For more information about our

postgraduate research programmes,
please contact:
Dr Caroline Proctor
Graduate School of Biological Sciences
2.06, Mary Brück Building
King’s Buildings
Mayfield Road
Edinburgh, UK
EH9 3DW
Tel +44 (0)131 650 5327
Email gradbiol@ed.ac.uk
www.ed.ac.uk/biology/postgraduate/pgr
To discuss your PhD proposal, you should

identify potential supervisors at:
www.ed.ac.uk/biology/people
Visit us
The University’s Postgraduate Open Day
is your opportunity to come and meet
current staff and students. Our next
campus-based Open Day takes place
on Wednesday 16 November 2016. For
more information, visit: www.ed.ac.uk/
postgraduate-open-day
The University also runs online information

sessions for prospective postgraduate
students throughout the year. For more
A702 SOUTH
information, visit: www.ed.ac.uk/
postgraduate/online-events
We are here!
The School of Biological Sciences
School of Biological
welcomes visitors at any time. We can
Sciences
give you a tour of the campus and arrange
a meeting with your potential Programme
Director. If you are unable to visit, we can
arrange a video call.
University building
Illustration by:
Katy Wiedemann, MA Illustration student
The front cover shows seven specimens of land dwelling
gastropod shells. These were collected by Charles Darwin while
on St Helena, 8-14 July 1836, during HMS Beagle’s return voyage
from the Galápagos Islands.
The items in this illustration are part of the University’s unique
Centre for Research Collections, a rich resource for all our
students, staff and the wider community.
#drawntoedinburgh
This publication is available online

at www.ed.ac.uk/postgraduate and
can be made available in alternative
formats on request. Please contact
communications.office@ed.ac.uk
or call +44 (0)131 650 2252.
Printed on Essential Silk FSC Mix, a Forestry Stewardship Published by:

Council certified paper stock. It was manufactured to Communications and Marketing,
ISO 14001 certified environmental management The University of Edinburgh
standards, using an elemental chlorine free process. Designed by:
We have made every effort to ensure the accuracy Tayburn
of the information in this prospectus before going Photography by:
to print. However please check online for the Paul Bock
most up-to-date information: www.ed.ac.uk Paul Dodds
On 23 June 2016 the UK electorate voted in a Edinburgh Inspiring Capital
national referendum to leave the European Union. Yao Hui
At the time of going to print, there was no immediate, Tricia Malley & Ross Gillespie
material change known that would impact applicants Real Edinburgh
for 2017 entry. However we recommend that Shutterstock
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The University’s standard terms and conditions
will form an essential part of any contract between
the University of Edinburgh and any student offered a
place here. Our full terms and conditions are available
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terms-conditions
© The University of Edinburgh 2016. No part of
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Biological Sciences: Postgraduate Opportunities 2017

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Biological Sciences

Postgraduate Opportunities 2017

Influencing the world since 1583

“Edinburgh isn’t so much a city, more 02 Introduction

Influencing the For more than 400 years the University of

Our distinguished alumni include NASA astronaut

International collaboration The majority of our research – 83 per

Linking research and commerce

Taught masters programmes www.ed.ac.uk/pg/856 www.ed.ac.uk/pg/1

Biochemistry Biodiversity & Taxonomy

Programme Director Professor Andrew Leigh-Brown

Bioinformatics Biotechnology Drug Discovery & Evolutionary Genetics

Programme description Programme description Programme description Programme description

Programme Director Dr Paul Taylor

Programme description Programme description Programme description Programme description

Programme Director Dr Louise Horsfall

Online distance learning

Drug Discovery & Next Generation

Programme description Programme description

COURSES PREVIOUSLY OFFERED INCLUDE: COURSES PREVIOUSLY OFFERED INCLUDE:

Research at the School of Biological Sciences Research opportunities

Research environment Research environment Research environment Research environment

of Edinburgh. I originally joined the Walkinshaw group English language requirements

About the School of Facilities and resources

Community Employability and

Applications and fees

students from Canada, China, India and the US through 

Campus map Get in touch

For enquiries relating to specific

For more information about our

To discuss your PhD proposal, you should

The University also runs online information

This publication is available online

Printed on Essential Silk FSC Mix, a Forestry Stewardship Published by:

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“Edinburgh isn’t so much a city, more 02 Introduction

students from Canada, China, India and the US through