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Classification

of the
— ANIMAL
KINGDOM
Richard E. Blackwelder
Classification of the Animal Kingdom
CLASSIFICATION
OF THE

ANIMAL KINGDOM
Richard E. Blackwelder

SOUTHERN ILLINOIS UNIVERSITY PRESS Carbondale, Illinois


COPYRIGHT © 1963 by Southern Illinois University Press

All rights reserved

LIBRARY OF CONGRESS CATALOG CARD NUMBER 62-17618


PRINTED IN THE UNITED STATES OF AMERICA
DESIGNED BY ANDOR BRAUN
CONTENTS

INTRODUCTION 1

Simplified List of Recent Phyla 6

Simplified List of Recent Classes and Orders,


with Common Names 9

Notes on the Taxa 26


Complete List of Phyla 36
Complete List of Classes and Orders,
with Synonyms, Subgroups, and Geologic Range 39
BIBLIOGRAPHY 72
INDEX TO COMMON NAMES 75
INDEX TO LATIN NAMES 80
INTRODUCTION

THE CLASSIFICATION OF ANIMALS is Still Very much a field in which dis-


covery and revision are continuing, even after two hundred years of
study. The importance of classification in biology increases every year,
because the experimental and practical fields find increasing need for
accurate identification of animals and for understanding of compara-
tive relationships.

At least one outstanding biologist has opposed pubUcation of


this new classification on the ground that it would be accepted as final,

the classification, and would tend to make students think that all higher
classification is finished. The intention of the compiler is just the op-

posite. Just as this classification is different in detail from all previous


ones, so will future editions be still different, as we learn more about
the comparative features of animals.
It is anticipated that every new edition will spur students of the
individual groups to propose improvements. It is therefore planned to
issue corrected editions whenever appropriate. The very appearance of
these subsequent editions will emphasize the growth of understanding
of animal groups.
Only one ostensibly complete classification of animals, living and
fossil, has been published in recent years. That classification, by A. S.

Pearse of Duke University, is a good one, based on the views of many


specialists. Certain mechanical faults make it less usable than it should
be, and the need for revision gave the original impetus to preparation

of the present classification. Because Pearse did not usually indicate the
source of his arrangements, he is not here cited as an authority. Never-
theless, the two classifications are basically very similar. No other single
classification has been found that agrees so closely with the conclusions
of the present study.
It should be emphasized that, within certain limits, this classifica-

tion is not a simple compilation of the views of specific workers. In


nearly all details, choices have been made between conflicting schemes
CLASSIFICATION OF THE ANIMAL KINGDOM 2
of various authors, not on the basis of the reputation of those authors

but on my judgment of the soundness of their supporting arguments or


on my analysis of the data they present. In none of the larger groups
has the work of any single author been accepted without modification.
Several considerations have influenced the decisions embodied in
this classification.

First, a false picture is given by a simplified classification, because the


existing diversity is one of the principal features of the animal kingdom.
Therefore, no groups should be combined merely for the sake of sim-
plicity.

Second, although the previous item would seem to require coverage of


the groupings at all possible levels, to show the extreme range of division
and subdivision, this is not in fact possible. Not only are there many

conflicting groupings at certain levels, such as of phyla or orders, but


there is no practical way to show these groupings in a general classifica-

tion. It is a compromise that is believed to be effective to subdivide the


phyla only into classes, subclasses, and orders. Other possible groupings,
such as subphyla and superorders are referred to in the notes.

Third, two groups which are so distinct at any level that they cannot
be described in common terms must be separated at that level. (For
example, Pterobranchia and Enteropneusta; see the Notes on the Taxa.)

Fourth, groups which cannot be distinguished at any particular level


by the type of characters used for their neighbors must be combined at

that level. (For example, the sometime classes of Nematoda.)

Fifth, the discovery of groupings within a class, for example, does not
justify the creation of new classes for each of the subgroups. The proper
level for the new groups can only be determined by comparison with
neighboring parts of the classification.

Sixth, although uniformity in the form (endings) of names at each


level would unquestionably be helpful, it cannot now be attained with-
out adding greatly to the total of name forms and synonyms. The sys-

tems so far proposed are so diverse as to introduce further confusion of


their own. None of the systems has been widely enough accepted to be
entitled to adoption throughout the Animal Kingdom. None has been
so widely accepted on a world basis, even in one group, as to indicate
3
Introduction
universal acceptance in the near future. Indeed, even the ordinal endings
in -iformes adopted by American ichthyologists and ornithologists are
almost entirely unused in the rest of the world. The resulting names are
unnecessarily long and cumbersome. The system does not relieve any-
one from learning the shorter forms The latter are used here, with
also.

the uniform-ending forms listed as synonyms. In other groups, usage


of the source of the classification is followed as to spelling, in most
cases. There are a variety of systems in use and no obvious trend to-

ward adoption of any single system.

This classification attempts to show the various spellings as well as


the various synonyms. Each zoologist will choose which one he wishes
to use in each case.

Seventh, no single rule will suffice for choice of names where several
apply to a single taxon. Reasons for each decision are given in the text
inmany cases, but in general it has been the goal to retain the best
known names, at the most appropriate level, regardless of homonymy.
Priority is considered to be of secondary importance at these levels.

Eighth, although considerable homonymy exists at all levels, even up


to that of phylum names, there is almost no real confusion caused
thereby. Until there are direct rules to govern the decisions, there seems
to be nothing gained by replacing well known names, such as Decapoda
(either in the Cephalopoda or in the Crustacea).
This classification is in three parts, the purposes of which are quite
different. In order of preparation, these are: 1] the complete classifica-

tion, including lists of the phyla and of the classes and orders, of all ani-
mals, living and fossil; 2] the justification for unusual features in this
classification; and 3] a simplified classification of Recent animals for
student use, with common names, again including lists of the phyla
and of the classes and orders. The arrangement of these parts in the
book is just the reverse of this.

In both lists, the phyla are first arranged in four subkingdoms, and
one of these is divided into four series. Many other groupings of these
phyla are possible, and several are shown in the footnotes of the section
Complete List of Phyla. It is not here believed that these supra-phylum
groupings are of much significance at this phase of the knowledge of
animals.
In both lists of orders, these orders are arranged in the appropri-
ate classes and subclasses. No other levels, such as superorder, are rec-
CLASSIFICATION OF THE ANIMAL KINGDOM 4
ognized. They may be of use in some circumstances but seem to be of
little value in showing the arrangement of the orders on a practical
basis.

Throughout, rejected synonyms are printed in itaUcs, the accepted

class names are in capital letters, and the subclass names and the order
names are both set in capitals and lower case letters. In the footnotes,
names that also appear in the classification above are printed in small
capitals. The other names in the footnotes are somewhat in the nature

of rejected synonyms, but as most of them are really the names of non-
accepted groupings, they appear in capitals and lower case roman letters.
To the variety of spellings there is no end. No attempt is made to

list all forms, but such spellings as would appear at a separate place in
an alphabetical index are listed, along with those variations that are
used for distinct levels; e.g., Echiuroidea (phylum), Echiurida (class),
and Echiuroina (order).
In the Complete List of Orders the geologic range of each group
is shown by symbols at the right margin. The meaning of these symbols

is shown in the following table.

REC Recent JUR Jurassic


QUA Quaternary mes Mesozoic
PLE Pleistocene per Permian
PLI Pliocene pen Pennsylvanian
OLI Oligocene mis Mississippian
Mio Miocene car Carboniferous
EOC Eocene (+ Paleocene) dev Devonian
ter Tertiary sil Silurian
CEN Cenozoic ord Ordovician
CRE Cretaceous cam Cambrian
TRI Triassic pal Paleozoic

There are a few points of discrepancy between the Simplified List


and the Complete List. These are intentional, to make the simplified
list more useful to students. The Complete List shows the definitive

classification that is here being proposed.


The names included under the footnote heading "Includes" may
be suborders, synonyms, rejected groups, or names of questionable ap-
plication.They are names which have at some time been used for
all

orders or more inclusive groups and are included merely to indicate


their approximate position in the scheme.
5 Introduction
Several recent schemes of classification in particular groups are
known to the compiler but are not followed herein. Some were received
too late for study (e.g., part W of the Treatise of Invertebrate Paleon-
tology). Some were not yet available in the form needed for our use
and so were not considered (e.g., Echinodermata by H. B. Fell and
Mollusca by Taylor and Sohl). There is no judgment of these schemes
implied in this action; they will be considered for a subsequent revised
edition.

It will probably be thought by some that this is an extreme classifi-

cation in separating many small groups as distinct phyla. The compiler


believes that it is a conservative classification even in this regard. He
believes that an important basic tenet of classification, too often over-
looked, is that all groups must be distinct and definable and that there-
fore forms are not to be forced into existing groups at any level if

they do not agree with what are deemed to be the important features of
that group. The important features in this case are those which caused
the group to be set aside and maintained as distinct.
It is sometimes possible to enlarge slightly the scope of a group
definition to admit forms previously unknown, but this does not justify
including widely divergent forms that cannot be defined together effec-
tively.
Simplified List

Subk Series Phylum

Kingdom AN MALI A^
I

EOZOA
^ ^
Protozoa [one-celled animals]

PARAZOA
Porifera sponges
AGNOTOZOA
Mesozoa
HISTOZOA (Metazoa)
Enterocoela
Monoblastozoa
Coelenterata (Cnidaria) hydroids, jellyfish, medusae, corals,
sea-anemones
Ctenophora comb-jellies, sea-walnuts
Acoelomata
Platyhelminthes flatworms
Rhynchocoela (Nemertinea) ribbon-worms, proboscis-worms
Pseudocoelomata
Acanthocephala spiny-headed-worms
Rotifera (Rotatoria) rotifers, wheel-animalcules
Gastrotricha
Kinorhyncha (Echinodera)
Priapuloidea
Nematoda thread-worms, round-worms
Gordiacea (Nematomorpha) horsehair- worms,
gordian-worms
Calyssozoa (Endoprocta)

of Recent Phyla

Subk Series Phylum

Coelomata
Bryozoa (Ectoprocta) moss-animals
Phoronida
Brachiopoda lamp-shells
MoUusca mollusks
Sipunculoidea
Echiuroidea
Myzostomida
Annelida [segmented worms]
Tardigrada bear-animalcules, water-bears
Pentastomida
Onychophora
Arthropoda crustaceans, arachnids, insects, etc.
Chaetognatha arrow-worms
Pogonophora beard-worms
Echinodermata
Pterobranchia
Enteropneusta
Planctosphaeroidea
Tunicata sea-squirts
Cephalochordata lancelets
Vertebrata vertebrates

^ For explanations, other synonyms, extinct groups, and other taxa above the
phylum level, see the section Complete List of Phyla.
^The Protozoa are sometimes placed in a separate kingdom of organisms
the Protista.
^ Non-Latin names can be made for each phylum
by merely using the English
form of the name, such as protozoans for Protozoa or arthropods for Arthropoda.
These are listed only where they are in common use.
Simplified List of Recent Classes and Orders

with Common Names

Class Subcl Order

PROTOZOA
FLAGELLATA (Mastigophora) flagellates

Phytomastigina [plant-like flagellates]

Chrysomonadina silicoflageUates, etc.

Coccolithophorida coccolithophores, coccoliths


Cryptomonadina
Phytomonadina ( Volvocales)
Euglenoidina
Chloromonadina
Dinoflagellata dinoflagellates

Zoomastigina [animal-like flagellates]


Rhizomastigina
Protomonadina
Polymastigina
Hypermastigina
SARCODINA
Rhizopoda rhizopods
Proteomyxa
Mycetozoa slime-molds
Amoebozoa
Testacea
Foraminifera foraminiferans, forams
Actinopoda
Heliozoa sun-animalcules
Radiolaria radiolarians
SPOROZOA
Telosporidia
Gregarinida
Coccidia
Haemosporidia
Cnidosporidia
Myxosporidia
Actinomyxidia
SIMPLIFIED LIST OF RECENT CLASSES AND ORDERS 10
Class Subcl Order

Microsporidia microsporidians
Helicosporidia
Sarcosporidia
Sarcosporidia
Globidia
Haplosporidia
Haplosporidia
CILIATA ciliates

Protociliata
Opalinida opalinids
Euciliata
Holotricha
Spirotricha tintiimids, etc.

Chonotricha
Peritricha
SUCTORIA
Suctoria

PORIFERA
sponges
CALCAREA (Calcispongea) [calcareous sponges, chalky sponges]
Solenida
Lebetida
Pharetronida
Thalamida
HYALOSPONGEA (Hexactinellida) glass-sponges
Lyssakina
Dictyonina
Lychniskophora
Heteractinida
DEMOSPONGEA
Myxospongida
Keratosida horny-sponges
Haplosclerida
Poecilosclerida
Hadromerida
Halichondrida
Epipolasida
Choristida
Carnosida
Lithistlda stone-sponges
1

1 Porifera — Coelenterata
Class Subcl Order

M ESOZOA
RHOMBOZOA
Dicyemida
Heterocyemida
ORTHONECTIDA
Orthonectida

MONOBLASTOZOA
MONOBLASTOIDEA
Monoblastidea

COELENTERATA
(Cnidaria)

coelenterates, medusae

HYDROZOA
Trachylinida
Hydroida
Milleporida (Hydrocorallinae) millepores
Stylasterina
Siphonophora siphonophores
SCYPHOZOA jellyfishes

Stauromedusae
Cubomedusae
Coronatae
Semaeostomeae
Rhizostomeae
ANTHOZOA sea-anemones, corals
Alcyonaria
Stolonifera
Telestacea
Alcyonacea soft-corals
Coenothecalia blue-corals
Gorgonacea sea-fans, horny-corals, gorgonians, sea-feathers
Pennatulacea sea-pens, sea-pansies
Zoantharia
Zoanthiniaria
CoraUimorpharia
Actiniaria sea-anemones
Scleractinia (Madreporaria) hexacorals, stony-corals
SIMPLIFIED LIST OF RECENT CLASSES AND ORDERS 12
Class Subcl Order

Ceriantipatharia
Antipatharia black-corals, thomy-corals
Ceriantharia

CTENOPHORA
comb-jellies, sea-walnuts

TENTACULATA
Cydippida
Lobata
Cestida
Platyctenea
NUDA
Beroida

PLATYH ELMINTHES
flatworms

TURBELLARIA planarians
Acoela
Rhabdocoela
Alloeocoela
Tricladida triclads

Polycladida polyclads
TREMATODA flukes
Monogenea
Aspidogastrea
Digenea
CESTODA tapeworms
Proteocephala
Tetraphyllidea
Disculicepitidea
Lecanicephala
Trypanorhyncha
Cyclophyllidea
Aporidea
Nippotaeniidea
Caryophyllidea
Spathebothridea
Pseudophyllidea
CESTODARIA
Amphilinidea
Gyrocotylidea
Biporophyllidea
3

1 Ctenophora — Kinorhyncha
Class Subcl Order

RH YNCHOCOELA
ribbon-worms, proboscis-worms

NEMERTINEA
Palaeonemertea
Heteronemertea
Hoplonemertea
Bdellonemertea

ACANTHOCEPHALA
spiny-headed-worms

ACANTHOCEPHALA
Archiacanthocephala
Palaeacanthocephala
Eoacanthocephala

ROTI FERA
(Rotatoria)

rotifers, wheel-animalcules

SEISONIDEA
Seisonacea
BDELLOIDEA
Bdellacea
MONOGONONTA
Ploima
Flosculariacea
Collothecacea

GASTROTRICHA
gastrotrichs

MACRODASYOIDEA
Macrodasyidea
CHAETONOTOIDEA
Chaetonotidea

KINORHYNCHA
ECHINODERA
Echinodera
SIMPLIFIED LIST OF RECENT CLASSES AND ORDERS 14
Class Subcl Order

PRIAPULOI DEA
PRIAPULOIDEA
Priapulida

NEMATODA
nematodes, nemas, thread-worms, round-worms

NEMATOIDEA
Enoploidea
Dorylaimoidea
Mermithoidea
Chromadoroidea
Araeolaimoidea
Monhysteroidea
Desmoscolecoidea
Rhabditoidea
Rhabdiasoidea
Oxyuroidea
Ascaroidea
Strongyloidea
Spiruroidea
Dracunculoidea
Filarioidea
Trichuroidea
Dioctophymoidea

GOR DIACEA
gordian-worms, horsehair-worms

NEMATOMORPHA
Gordioidea
Nectonematoidea

CALYSSOZOA
endoprocts

ENDOPROCTA (Entoprocta)

Pedicellinida
15 Priapuloidea — Mollusca
Class Subcl Order

BR YOZOA
sea-mats, corallines, moss-animals, bryozoans, sea-mosses

PHYLACTOLAEMATA
Lophopoda
GYMNOLAEMATA
Cyclostomata
Ctenostomata
Cheilostomata

PHORON DA I

phoronids

PHORONIDA
Phoronida

BR ACH lOPO DA
lamp-shells, brachiopods

INARTICULATA
Atremata
Neotremata
ARTICULATA
Protremata
Telotremata

MOLLUSCA
mollusks

MONOPLACOPHORA
Tryblidioidea
AMPHINEURA chitons
Neoloricata
APLACOPHORA (Solenogastres)
Neomeniida
Chaetodermatida
GASTROPODA snails, slugs, gastropods
Prosobranchia
Archaeogastropoda limpets, ear-shells, turbans
Caenogastropoda
Opisthobranchia
Pleurocoela sea-hares
SIMPLIFIED LIST OF RECENT CLASSES AND ORDERS
Class Subcl Order

Pteropoda butterfly-shells, pteropods


Sacoglossa
Acoela nudibranchs
Pulmonata land-snails, slugs

Basommatophora boat-shells, ramshorns


Stylommatophora slugs

BIVALVIA (Pelecypoda, Lamellibrcmchiata) bivalves, oysters, clams,

mussels, pelecypods
Protobranchia
Filibranchia
Eulamellibranchia
Septibranchia
SCAPHOPODA tooth-shells, tusk-shells

Scaphopoda
CEPHALOPODA cephalopods
Tetrabranchiata
Nautiloidea pearly-nautilus

Dibranchiata
Decapoda squids, cuttle-fish

Octopoda octopuses, argonauts


Vampyromorpha

SIPUNCULOIDEA
[sipunculid worms]

SIPUNCULOIDEA
Sipunculida

ECHI U ROI DEA


ECHIURIDA
Echiuroina
Xenopneusta
Heteromyota
SACCOSOMATIDA
Saccosomatida

M YZOSTOM DA I

MYZOSTOMIDA
Proboscidea
Pharyngidea

ANNELIDA
[segmented worms] annelids
7

1 Sipunculoidea — Arthropoda
Class Subcl Order

CHAETOPODA
Polychaeta polychaetes
Errantia sandworms
Sedentaria tubeworms
Oligochaeta earthworms, angle-worms, night-crawlers, oligochaetes
Plesiothecata
Prosothecata
Prosopora
Opisthopora
HIRUDINEA leeches, bloodsuckers
Rhynchobdellida
Gnathobdellida
Pharyngobdellida
Acanthobdellida
ARCHIANNELIDA
Archiannelida

TAR DIGRADA
bear-animalcules, water-bears

HETEROTARDIGRADA
Arthrotardigrada
Echiniscoidea
EUTARDIGRADA
Eutardigrada

PENTASTOM DA I

LINGUATULIDA
Cephalobaenida
Porocephalida

ONYCHOPHORA
PERIPATIDEA
Euonychophora

ARTHROPODA
arthropods

MEROSTOMATA
Xiphosura
Xiphosurida horseshoe-crabs
PYCNOGONIDA sea-spiders
Eupantopoda
SIMPLIFIED LIST OF RECENT CLASSES AND ORDERS l8
Class Subcl Order

ARACHNIDA arachnids
Latigastra
Scorpionida scorpions
Pseudoscorpionida book-scorpions, false-scorpions,
pseudoscorpions
Phalangida (Opiliones) harvest-men. Daddy-long-legs
Acarida mites, ticks, chiggers
Caulogastra
Palpigradida microscorpions
Thelyphonida whip-scorpions
Schizomida
Phrynichida tailless-whip-scorpions
Araneida {Araneae) spiders, tarantulas, black-widows
Solpugida (Solifugae) solpugids, sun-spiders
Ricinuleida (Podogonata)
CRUSTACEA crustaceans
Branchiopoda
Anostraca brine-shrimps
Notostraca phyllopods, fairy-shrimps
Conchostraca
Cladocera water-fleas
Cephalocarida
Cephalocarida
Ostracoda
Myodocopida
Podocopida
Mystacocarida
Mystacocarida
Copepoda (Eucopepoda)
Calanoida
Harpacticoida
Cyclopoida
Notodelphyoida
Monstrilloida
Caligoida
Lernaeopodoida
Branchiura
Branchiura
Cirripedia barnacles
Thoracica
Acrothoracica
19 Arthropoda
Class Subcl Order

Ascothoracica
Apoda
Rhizocephala
Malacostraca
Nebaliacea
Anaspidacea
Mysidacea opossum-shrimps
Thermosbaenacea
Spelaeogriphacea
Lophogastridea
Cumacea
Tanaidacea
Isopoda pillbugs, sowbugs
Amphipoda scuds
Euphausiacea krill

Decapoda crabs, lobsters, crayfish, shrimps, prawns


Stomatopoda mantis-shrimps
PAUROPODA
Heterognatha
SYMPHYLA
Cephalostigmata
DIPLOPODA millipedes
Pselaphognatha
Ancyrotricha
Lophotricha
Chilognatha
Limacomorpha
Oniscomorpha
Ascospermophora
Colobognatha
Nematophora
Proterospermophora
Opisthospermophora
CHILOPODA centipedes
Pleurostigmophora
Geophilomorpha
Scolopendromorpha
Lithobiomorpha
Craterostigma
Notostigmophora
Scutigeromorpha
SIMPLIFIED LIST OF RECENT CLASSES AND ORDERS 20
Class Subcl Order

INSECTA (Hexapoda) insects

Apterygota
Protura proturans
Thysanura silver-fish, bristle-tails, rock-jumpers
Entotrophi campodeids, japygids
Collembola springtails, snow-fleas

Exopterygota (Heterometabola)
Ephemerida mayflies
Odonata dragonflies, damselflies, mosquito-hawks,
devil's-darning-needles, snake-doctors
Plecoptera stone-flies, salmonflies

Grylloblattoidea grylloblattids

Orthoptera (Saltatoria) grasshoppers, crickets, locusts,


katydids, mole-crickets
Phasmidia walking-sticks, stick-insects, leaf-insects
Blattaria roaches, cockroaches, croton-bugs
Mantodea praying-mantis, soothsayers, mantids
Dermaptera earwigs
Embioptera embiids, webspinners
Isoptera termites, white-ants

Psocoptera (Corrodentia) psocids, book-lice, bark-lice,


dust-lice

Zoraptera zorapterans
Mallophaga bird-lice, biting-lice

Thysanoptera thrips

Homoptera cicadas, leaf-hoppers, tree-hoppers, aphids,


scale-insects, spittle-bugs, mealy-bugs, frog-spit, psylhds,

lantern-flies, white-flies

Heteroptera bugs, bed-bugs


Anoplura sucking-lice, lice

Endopterygota {Holometabola)
Neuroptera snake-flies, serpent-flies, lace-wings, ant-lions,

dobson-flies, fish-flies, orl-flies


Mecoptera scorpion-flies

Trichoptera caddis-flies, trout-flies, case-flies

Lepidoptera butterflies, moths, skippers, blues, woolly-bears,


caterpiflars, millers

Diptera flies, gnats, mosquitoes, midges, bots, maggots,

punkies
Siphonaptera fleas, chigoes
Coleoptera beetles, weevils, fireflies, elaters, glow-worms,
water-pennies, meal-worms, wire-worms, white-grubs
21 Insecta — Echinodermata
Class Subcl Order

Strepsiptera stylopids
Hymenoptera bees, wasps, ants, sawflies, hornets, wood-wasps,
ichneumon-flies, gall-wasps, velvet-ants, horntails,
tarantula-hawks

CH AETOGN ATH A
arrow-worms

SAGITTOIDEA
Sagittoidea

POGONOPHORA
beard-worms

POGONOPHORA
Thecanephria
Athecanephria

ECHINODERMATA
echinoderms

CRINOIDEA feather-stars, crinoids, sea-lilies


Articulata
Isocrinida
Cyrtocrinida
Comatulida
ASTEROIDEA starfishes, sea-stars

Phanerozonea
Spinulosa
Forcipulata
OPHIUROIDEA brittle-stars, sand-stars, basket-stars, serpent-stars

Myophiurida
Ophiocystiida
Aganasterida
Phrynophiurida
Laemophiurida
Gnathophiurida
Chilophiurida
ECHINOIDEA sea-urchins, heart-urchins, sand-dollars
Regularia
Cidaroida
Centrechinoida
Exocycloida
SIMPLIFIED LIST OF RECENT CLASSES AND ORDERS 22
Class Subcl Order

Irregularia
Holectypoida
Cassiduloida
Clypeastroida cake-urchins, sand-dollars
Spantangoida heart-urchins
HOLOTHURIOIDEA sea-cucumbers
Aspidochirota
Elasipoda
Dendrochirota
Molpadonia
Apoda
PTEROBRANCHIA
PTEROBRANCHIA
Rhabdopleurida
Cephalodiscidea

ENTEROPNEU STA
acom-worms, tongue-worms
ENTEROPNEUSTA
Balanoglossida

PLANCTOSPH AEROI DEA


PLANCTOSPHAEROIDEA
Planctosphaeroidea

TUN ICATA
tunicates

LARVACEA
Larvacea
ASCIDIACEA sea-squirts, ascidians
Stolidobranchiata
Aspiraculata
Phlebobranchiata
Aplousobranchiata
Octacnemida
THALIACEA
Pyrosomata
Pyrosomatida
Myosomata
Cyclomyaria
Hemimyaria
Desmomyaria
23 Pterobranchia — Vertebrata
Class Subcl Order

CEPH ALOCHOR DATA


LEPTOCARDIA
Amphioxi lancelets

VERTEBRATA
vertebrates

AGNATHA [jawless fishes]


Cephalaspidomorpha
Cyclostomata lampreys, hag-fishes, sHme-eels
CHONDRICHTHYES [cartilaginous fishes]
Elasmobranchii
Selachii sharks, dogfishes, angel-fishes
Batoidea skates, rays

Holocephali rabbit-fishes
Chimaerae chimaeras, ratfishes
OSTEICHTHYES [bony fishes]

Actinopterygii [ray-finned fishes]


Chondrostei
Holostei gars
Teleostei
Choanichthyes [lobe-finned fishes]
Crossopterygii coelacanths, etc.
Dipnoi lungfishes
AMPHIBIA (Batrachia) amphibians, batrachians
Salientia
Anura frogs, toads

Lepospondyli
Urodela salamanders, newts
Apoda (Gymnophiona) caecilians

REPTILIA reptiles

Anapsida
Chelonia tortoises, turtles

Diapsida
Rhynchocephalia
Squamata lizards, snakes
Crocodilia crocodiles, gavials, alligators, caymans
AVES birds
Neornithes
Sphenisci {Spheniscijormes) penguins
Struthiones (Stmthioniformes) ostriches
SIMPLIFIED LIST OF RECENT CLASSES AND ORDERS 24
Class Subcl Order

Rheae (Rheiformes) rheas


Casuarii (Casuariiformes) cassowaries, emus
Apteryges (Apterygiformes) kiwis
Crypturi {Crypturijormes, Tinami, Tinamijormes) tinamous
Gaviae (Gaviiformes) loons
Podicipedes (Podicipediformes, Colymbae, Colymbijormes)
grebes, divers
Procellariae (Procellariiformes, Tubinares) albatrosses,

shearwaters, petrels, fulmars,


Steganopodes (Pelecani, Pelecaniformes) cormorants,
pelicans, gannets, tropicbirds, boobies, snake-birds,
frigate-birds
Ciconiae (Ciconiiformes) herons, bitterns, storks,
hammerheads, spoonbills, ibises, flamingoes

Anseres (A nseriformes) ducks, geese, swans, screamers


Falcones (Falconiformes) hawks, eagles, vultures, falcons,
caracaras, ospreys, harriers, secretary-birds
Galli (Galliformes) megapodes, pheasants, quails, grouse,

turkeys, fowls, peacocks, hoatzins


Grues (Gruiformes) cranes, limpkins, raUs, sunbitterns,
bustard-quails, plainwanderers, trumpeters, coots, gaUinules,
kagus, sungrebes, bustards
Charadriae (Charadriiformes) jacanas, snipe, oyster-catchers,
plovers, tumstones, surf-birds, woodcock, sandpipers,
avocets, stilts, phalaropes, gulls, terns, skimmers, awks,
murres
Columbae (Columbiformes) doves, pigeons, dodos,
sandgrouse, solitaires
Psittaci (Psittaciformes) parrots, parakeets, lories, macaws
Cuculi (Cuculiformes) cuckoos, plantain-eaters, touracos,
anis, roadrunners
Striges (Strigiformes) owls
Caprimulgi (Caprimulgiformes) goatsuckers, potoos,
oil-birds, frogmouths
Macrochires (Macrochiriformes, Apoda, Micropodi) swifts,

humming-birds
Colli (Coliiformes) mouse-birds, colics
Trogones (Trogoniformes) trogans
Coraciae (Coraciiformes) kingfishers, rollers, hoopoes,

hornbills, todies, motmots, bee-eaters


Pici (Piciformes) woodpeckers, toucans, honey-guides,
jacamars, puffbirds, barbets, piculets
25 Vertebrata
Class Subcl Order

Passeres (Passeriformes) songbirds, warblers, thrushes,


shrikes, creepers, nuthatches, titmice, vireos, finches,
tanagers, blackbirds, starlings, orioles, crows, jays, magpies
swallows, butcher-birds, wrens, thrashers, mockingbirds,
kinglets, flycatchers, wrentits, dippers, honey-creepers,

grosbeaks, buntings, broadbills, woodhewers, antbirds,


ovenbirds, lyrebirds, bulbuls, larks, babblers, wagtails,
waxwings, weaverbirds, drongos, wattlebirds, bowerbirds,
birds-of-paradise, etc.
MAMMALIA mammals
Prototheria
Monotremata monotremes, platypus, echidna
Metatheria
MarsupiaUa marsupials, opossums, Tasmanian-wolf,
bandicoots, phalangers, koalas, kangaroos
Eutheria (Placentalia) placentals
Insectivora insectivores, tenrecs, hedgehogs, shrews
Dermoptera colugos, flying-lemurs
Chiroptera bats, vampires
Primates lemurs, tree-shrews, aye-aye, lorises, bush-babies,
tarsiers, monkeys, marmosets, macaques, baboons, guenons,
langurs, apes, gibbons, lars, chimpanzees, orangutans, gorillas,
men
Edentata sloths, anteaters, armadillos

Pholidota pangolms
Lagomorpha hares, rabbits, pikas
Rodentia squirrels, chipmunks, marmots, pocket-mice, pacas,
kangaroo-rats, beavers, rats, mice, muskrats, lemmings, voles,
dormice, porcupines, capybaras, guinea-pigs, chinchillas
Cetacea porpoises, dolphins, whales
Carnivora dogs, wolves, foxes, bears, raccoons, coatis,
kinkajous, pandas, weasels, minks, otters, badgers,
wolverines, skunks, civets, hyenas, cats, seals, sea-lions,
walruses
Tubulidentata aardvarks
Proboscidea mastodons, mammoths, elephants
Hyracoidea hyraxes
Sirenia sea-cows
Perissodactyla horses, zebras, tapirs, rhinoceroses
Artiodactyla pigs, peccaries, hippopotamuses, camels, llamas,
alpacas, guanacos, deer, giraffes, pronghorns, cattle,
antelopes, sheep, gazelles, musk-oxen, goats, etc.
Notes on the Taxa

Subkingdoms and their subdivisions


Animalia. Division of the Animal King- Eozoa and Agnotozoa. In the choice of
dom into four subkingdoms is seen in names for subkingdoms, it has been felt
many recent classifications, although some that only slight advantage results from
writers prefer to list the Parazoa, Mesozoa, having a single phylum known by a differ-
and Eumetazoa as branches of Metazoa in ent name than is used for its subkingdom.
contrast to the Protozoa. The use of the However, with only four subkingdoms in-
additional level "branch" is difficult to jus- volved and with two of these consisting of
tify where so few groups are involved, un- two or more phyla, it appears to be rea-
less there is definite information on the sonable to be uniform in this respect and
phylogeny of these groups. Such knowl- use separate names for the subkingdom
edge of the relationships of the groups is and phylum that include the protozoans
lacking, or, at best, highly speculative. The and for the subkingdom and phylum that
groups are generally recognized at the include the mesozoans.
phylum and subkingdom levels, and these
seem to be adequate for classification ex- Parazoa. The Porifera have long been rec-
cept within the Metazoa proper (see be-
ognized as constituting a group distinct
low).
from the rest of the many-celled animals.
Inasmuch as it is often stated that ani-
The extinct Cyathospongia, under one of
mals are either one-celled or many-celled,
the three available names, were placed
it would seem to be necessary to accept the
with the sponges by Okulitch and others,
older subdivision of Animalia into two sub-
and as a separate phylum in the Parazoa
kingdoms, Protozoa and Metazoa. The fact
by Pearse and others.
is, however, that many undoubted proto-

zoans exist only in aggregations of many


cells, often with as much division of labor Histozoa. This name is accepted here be-
between cell types as in some undoubted cause of the great ambiguity of the more
metazoans. Removal of the Protozoa to a familiar name Metazoa. As explained
separate kingdom Protista solves part of above, the incorporation of many cells into
this problem, but the remaining animals one body is not distinctive of any major
still represent the three very different basic group of animals, even if single-celled
structures: 1] a vase-like cylinder open adult structure is found exclusively in one
at one end and with several types of cells group. It was the desire to retain Metazoa
in the walls but with the internal cavity which has led many writers to list the
not serving for digestion, 2] a solid body Parazoa, Mesozoa, and Eumetazoa as
consisting of one layer of cells around a branches of a subkingdom Metazoa. Inas-
central cell or group of cells, and 3] a much as Metazoa cannot be effectively de-
multicellular body with internal cavities of fined, to the exclusion of all Protozoa, it

which one is usually a digestive tract and seems to be more realistic to recognize
with walls of one, two, or three layers of three or four subkingdoms of animals on
cells. the basis of the general body construction.
These three types of construction are Attempts to divide it on the basis of cell
so different as to require recognition as pri- number are arbitrary and misleading.
mary divisions of the kingdom. With the The use of any of these subkingdom
Protozoa (when these are treated as ani- groups is of questionable value; it is the
mals), they form the four subkingdoms phyla that are important and that are most
employed here: Eozoa, Parazoa, Agnoto- often definable. The distinction between
zoa, and Histozoa. even Protozoa and Metazoa is so com-
27 Notes on Subkingdoms and Phyla
pletely nebulous that no longer serves
it point it appears that the most useful group-
any clear purpose. Use of the subkingdom ing is the one based on the type of body
and series names herein is simply a recog- cavity. Accordingly four groups are here
nition of their use in many recent classifi- recognized, those phyla with an enterocoel
cations. or gastrovascular cavity only, those with
Division of the Histozoa (Metazoa) no cavities except a digestive tract, those
been at-
into grades, series, or divisions has with a pseudocoel, and those with a coe-
tempted many times. Most of the groups so lom. These groupings are all well known
adopted in the past are employed in the from Hyman (1940) and other works.
face of obvious defects in the form of ex- No satisfactory name exists for the
ceptions. Not all Radiata show any form first of these four groups. Radiata is inap-

of radial arrangement, and not all radially plicable to many forms. Protaxonia is
arranged animals are put in the Radiata. based on a concept of embryonic axes
Bilateria included animals with quite di- which would include extraneous groups.
verse body plans, some with virtually no Enterozoa and Enterocoela were originally
paired structures, no obvious "sides," no applied to much larger concepts. On the
anterior and posterior, and only a remote whole, Enterocoela is the most appropriate
similarity to the obviously bilateral ani- in meaning, and it is adopted here.
mals. Some groups placed in the Schizo-
coela form their coelom in the enterocoe- Radiata and Bilateria. The Histozoa or
lous manner, and at least one group placed Metazoa have sometimes been divided into
in the Enterocoela forms its coelom by the the Radiata (Coelenterata and Cteno-
schizocoelous method. Articulata has in- phora) and the Bilateria (all others) on
cluded animals that are not segmented. the basis of their general body arrange-
And so on. ment. The distinction is here held to be a
It is here concluded that the histozoan fictitious one, because Ctenophora are
(or eumetazoan) phyla cannot be grouped much less radial than some Echinoder-
readily into clear-cut series. The number mata, and such an animal as a bryozoan is
of these phyla is not so great as to force so completely different in body arrange-
subdivision of the subkingdom, but custom ment from an annelid worm that it is

seems to be sufficient justification for indi- meaningless to say that they are both bi-
cating some grouping of them. At this lateral.

Phyla and rejected phylum groups


Cyathospongia. Recent works on this ex- at the other. The cells are thus simultane-
tinct phylum have adopted either Pleo- ously both "ectoderm" and "endoderm";
spongia or Archaeocyatha as the phylum they are ciliated on both surfaces. Repro-
name. Both of these names are of later duction is asexual (by transverse fission),
date (1937) than Cyathospongia (1935), and there are indications of a sexual proc-
and Archaeocyatha has been used more ess in the form of fusion of two individ-
consistently for one of the included classes. uals. It is possible that a ciliated unicellu-
There seems to be no firmly established lar larva results from the sexual process.
usage that prevents us adopting the oldest Hyman suggested (1940) the phylum
name. status for this peculiar animal, but she did
not propose such a phylum directly and
Mesozoa. There appears to be no reason left Salinella unassigned to any group.
for not adopting the name now inwide use
for this phylum, especially as it is the old- Graptozoa. The graptolites have had a
est name (1877). The name has been also more varied history than most other major
used at the subkingdom level, where the groups. They were for years assigned to the
synonym Agnotozoa seems to be more ap- Hydrozoa in the Coelenterata, but have
propriate. more recently been transferred to the
Hemichordata. In 1959 Hyman examined
Monoblastozoa. A new phylum named the arguments supporting the hemichor-
here for the unique metazoan Salinella, date assignment. After effectively dispos-
which has too long been left excluded ing of all of these, she left the group
from the classifications of animals. It con- without clear assignment, although she pre-
tains only one genus and one species. The sumably retained them in the Hydrozoa, as
animal consists of a single layer of similar she had doubtfully done in 1940.
cells surrounding an internal tubular tract Some features of the skeleton of grap-
which has a "mouth" at one end, an "anus" tolites are not duplicated in the Hydrozoa,
CLASSIFICATION OF THE ANIMAL KINGDOM 28
and the nature of the material of this exo- Coelenterata. In some recent works (es-
skeleton is not known. It is not possible to pecially Hyman, 1940) this phylum has
say definitely the graptolites are
that been called Cnidaria because Coelenterata
coelenterates, because the nature of the has at other times included such groups as
body cavity is not known, but there is no Ctenophora and Porifera. This reason for
evidence that not a coelenteron. The
it is abandoning the universally known name
graptolites maythus reasonably be in- Coelenterata would, if applied to other
cluded in the Coelenterata, even placed in modern phylum concepts, result in chang-
the Hydrozoa because of general similari- ing most of the familiar names, including
ties, but information is simply lacking that Porifera, Annelida, Arthropoda, Hemi-
would enable one to say that they did have chordata, and Chordata. Such a change
the features of the coelenterates. cannot, in the opinion of the writer, be jus-
Inasmuch as there is some evidence of tified by any benefits resulting therefrom.

bilaterality, the skeletal tubes are different If it is thought to be necessary, the prior
from those of Hydrozoa in manner of for- name Nematozoa would also have to be
mation, and the almost universal form of considered. (Furthermore, inclusion of
the colonies is unmatched in the Hydrozoa, the Graptozoa or Conularida (as in Hy-
it is also reasonable to emphasize the dif- man) would likewise necessitate a change
ferences by separating the group from the in the phylum name, by this same argu-
Hydrozoa as a class. Because of the lack of ment.)
knowledge of all soft-part features and the
possibility that some of these also are with- Aschelminthes. The proposal of this name
out counterpart in the Coelenterata, it is by Hyman (1940) for all of the Pseudo-
here preferred to emphasize this uncer- coelomata except the Entoprocta has been
tainty by separating the group as a phylum adopted by some later works, but her alter-
distinct from Coelenterata. nate conclusion that the subphyla each be
The class Hydrozoa is already one of treated as a separate phylum has also been
the most diverse in the Animal Kingdom. followed by some. It is surely premature to
It seems undesirable to increase further its claim that the Aschelminthes has been
heterogeneity by including an additional conclusively accepted. Hyman (1951) re-
series of different features. The distinctive moves one of the original seven groups
features of the Coelenterata cannot be ad- (Acanthocephala) as a separate phylum.
duced to help us with the graptolites, so The definition given for the emended
these fossils cannot be included in that phylum Aschelminthes in 1951 contains
phylum upon any firm basis. no clear-cut distinctions. Unless such ex-
ist, the supposed phylum must be con-
cluded to be an indefinable assemblage.
Conularida. The same arguments as ap- The fact that the included subgroups are
plied above to the Graptozoa are cogent mostly small and less well known is of no
for the separation of the Conularida as a value in determining whether they are
separate phylum also. Apparently no coe- phyla, subphyla, or classes. It is here be-
lenterates have a chitinophosphatic skele- lieved that they are adequately distinct by
ton, which fact alone makes the inclusion clear-cut features of fundamental nature
of these animals in that phylum unsatisfac- (body plan, ciliation, "segmentation" of
tory. Of course, here -Iso there is no direct cuticle, presence of jaws, presence of flame
evidence that the animals were actually bulbs or solenocytes, musculature, nature
coelenterate in nature. of nervous system, etc.) to be considered
This group has recently been assigned separate phyla.
to the Scyphozoa, although also placed
sometimes as a phylum near the Annelida Nemathelminthes and Trochelminthes.
or as a member of some other phylum. Al- These two names were formerly applied to
though quite easily restored to look very the thread-like and the ciliated animals
much like elongate scyphozoans, the fossil more recently combined into the Aschel-
remains of these animals show consistent minthes. These groupings are also difficult
differences in the steep-sided pyramidal to define. It istherefore preferred to treat
form with four distinct sides, the closing their components as separate phyla. Nema-
of the aperture by lobes of the side faces, thelminthes usually included the Nema-
and the chitinophosphatic nature of the toda, Gordiacea, Acanthocephala, and
periderm. The arguments in the Treatise sometimes the Chaetognatha. The Troch-
(F) for combining these with the Scypho- elminthes included the Rotifera and
zoa seem very weak. Gastrotricha.
29 Notes on Subkingdoms and Phyla
Nematoda. Hyman isfollowed here in re- from scars or galls, they cannot be as-
jecting recent proposals to change this signed to orders.
name to Nemata.
Protarthropoda. The inclusion of Tardi-
grada, Onychophora, and Pentastomida in
Gordiacea. The best form of this name at
the Arthropoda as a subphylum (Pro-
thephylum level is a moot question. Little
tarthropoda, Pararthropoda, or Oncopoda)
isgained but confusion by using Nemato-
has been done, in every case traced out,
morpha, whereas Gordioidea has generally
without direct consideration of whether
been used at the class or order level. Gor-
they have the basic features of arthropods,
diacea seems to remain as the most distinc-
or whether the resulting agglomeration
tive phylum name.
can be defined. Apparently it cannot be
defined, and these three groups individually
Calyssozoa / Endoprocta. Entoprocta is
have only a few of the basic arthropod fea-
accepted by Hyman for this group, but this
tures. Until more correlation is demon-
name (or the more distinctive spelling
strated, held that they cannot reasona-
it is
Endoprocta) is much more often applied
bly be combined with the Arthropoda.
at the class level. Calyssozoa was proposed
originally (and followed by Kamptozoa) Tardigrada. This group is generally
for the group as a phylum. Inasmuch as placed in either the Aschelminthes or the
names are needed at both levels, Calys- Arthropoda. Either position is untenable
sozoa is accepted here for the phylum and if Cuenot (1949) is correct in asserting
Endoprocta for the single class. (The that the animals are coelomate and entero-
spellings Endoprocta and Entoprocta are coelous. It is distinguished from Onycho-
both ascribed to Nitsche (1870). Both phora and Pentastomida, as well as Ar-
have been used extensively, but the former thropoda, by features of considerable im-
is more distinct from Ectoprocta and is portance. It is certainly entitled to phylum
therefore adopted here.) status, even if the correct position for the
phylum is still unknown.
Myzostomida. This peculiar and little- Pentastomida. This group is generally
known group has previously been listed as placed in the Arthropoda, sometimes even
a class of Annelida, as a subclass of
in the order Acarida. Some of its charac-
Chaetopoda, or as part of the Polychaeta, ters have been ascribed to parasitic degen-
but apparently never as a phylum. eration. They apparently have no cilia, do
These animals are disc-shaped, are have a chitinous cuticle, and do have an
non-segmented although with some paired arthropod type of nervous system. They
organs, have jfive pairs of ventral append- lack an exoskeleton, jointed appendages,
ages and four pairs of suckers, have ten or Malpighian tubules or coxal glands, circu-
more pairs of marginal cirri or tentacles, latory organs, tracheae, and nephridia, and
lack blood-vascular and respiratory sys- their appendages are of the type seen in
tems as well as multiple nephridia, have the the Onychophora and the Tardigrada.
central nervous system consisting of a sin-
As it would be impossible to place
gle large ventral stellate ganglion and two
these definitely in any class of arthropods,
nerve rings around the oesophagus and and since they lack many arthropod fea-
pharynx, have a complete digestive tract
tures, it seems best to emphasize their dif-
but with the stomach branching through- ferences by treating them as a separate
out the body, and have a trochosphere
phylum.
larva. These features would make the An-
nelida impossible to diagnose, and they re- Hemichordata {Branchiotremata, Adelo-
sult in an animal whose peculiarities are chorda). Nearly all recent classifications
only obscured by inclusion in the An- recognize a phylum Hemichordata that in-
neUda. cludes the Enteropneusta, the Pterobran-
Prenant (1960) in the Traite concludes chia, and perhaps such other groups as the
that these animals are annelids but suffi- Graptozoa. A good example of this is
ciently distinct to be made a class. It is Hyman (1959). It is difficult to understand
here believed that the features cited by this grouping when every attempt at defi-

Prenant make it necessary to remove the nition consists primarily of variable or


Myzostomida from the Annelida, just as relative characters. The components are so
the Sipunculoidea and Echiuroidea had Hyman can only discuss them
distinct that
previously been removed. separately.Almost none of her statements
Myzostomids are reported from sev- apply throughout the phylum.
eral geologic eras. As these are known only In this situation we only obscure the
CLASSIFICATION OF THE ANIMAL KINGDOM 30
diversityand the differences by pretending wallsdo occur, the extreme differences in
that they can be included in one phylum. arrangement of the digestive tract, the ab-
Besides the Enteropneusta and the Ptero- sence of paired structures in tunicates,
branchia, there have been placed here also the presence of the tunic and the substance
the Planctosphaeroidea, the Phoronida, tunicin in tunicates, the reversible blood
and the Graptozoa. The latter two have al- flow in tunicates, and so on, all seem to
ready been accepted as distinct phyla in deny a similarity so close as to justify in-
a previous part of this classification (fol- clusion in one phylum. With the tunicates
lowing many other classifications). The included, the Chordata are extremely dif-
Planctosphaeroidea, consisting of certain ficult to define effectively, except on the
ciliated larvae of unknown affinities, can- two features of notochord and pharyn-
not be associated with any known adults. gotremy. With the tunicates excluded, the
It is therefore impossible to combine them list of features held in common by the re-

confidently with any phylum. It seems maining groups is considerably increased.


necessary to maintain them as a distinct The Cephalochordata can much more
group at the present time. reasonably be united with the vertebrates.
Nevertheless, the oral hood, the atrial sys-
Chordata. It is now more than seventy- tem, the brown funnel, the multiple paired
five years since the vertebrates and some solenocytic nephridia, the single-layered
supoosed relatives were first combined un- epidermis, the peculiar liver-pouch, the ab-
der this name. Nearly all subsequent clas- sence of a heart, the multiple ductless gon-
sifications have accepted this arrangement, ads, and other features seem to show that
although the included groups vary some- the group is "more widely separated from
what. At the extreme the Chordata have the lowest fish than the lowest fish from a
included the Hemichordata, Tunicata, and bird or mammal" (Parker & Haswell,
Cephalochordata, as well as the Verte- 1897). It seems appropriate to recognize
brata. these substantial differences at the phylum
The inclusion of the Hemichordata is level.
unequivocally rejected by Hyman (1959) The use of thename Chordata for the
and others. The arguments seem to be well restricted concept seems to be unnecessary
founded, involving the absence of any sub- as well as inappropriate. The name is
stantial similarity in major features. younger and less well known than Verte-
The Tunicata, included in the Chor- brata. It would seem to be an unnecessary
data as a matter of course in many works, name, based on overemphasis on a very
are excluded here because the similarities few features held in common by the groups
appear to be far outweighed by the differ- involved. At the most it might be consid-
ences between tunicates and vertebrates. ered to be a sort of "superphylum," but
Even if an homologous notochord be pres- even thus it could reasonably include only
ent, even if perforations of the tracheal the Cephalochordata and the Vertebrata.

Sources of class and order arrangements


Protozoa. The five classes of Protozoa are most recent works, including the Treatise
almost universally agreed upon in recent of Invertebrate Paleontology (E, 1955),
works. The use of the subphyla is not so but most paleontology books list also the
widespread. Protociliata may be consid- Receptaculitida as incertae sedis. Inasmuch
ered to belong in the Plasmodroma rather as it cannot be justified in any of the three
than in the Ciliophora, being there treated classes, it must stand as a separate class.
as a separate class. Suctoria are sometimes (The Nidulitida are now thought to be
united with Ciliata as a subclass. Telo- algae rather than sponges, and the Pleo-
sporidia (Amoebosporidia) are sometimes spongea are treated as a separate phylum).
separated from Sporozoa as a third sub- Many works divide each of the three
phylum. classes into subclasses. In some cases these
This is the classification of Kudo subclasses are based on features now be-
(1954) except for: 1] the elevation of lieved to be of minor importance or taxo-
Haplosporidia and Sarcosporidia to sub- nomic value, and the view of de Laubenfels
class level, following Hyman (1940); and is here adopted that subclasses do not add

2] a few cases of different choice from effectively to the classification of this


available synonyms. group. Aside from this, the arrangement
here adopted is substantially that of both
Porifera. Three classes are recognized in Hyman and the Treatise (E).
31 Notes on Subkingdoms and Phyla
Cyathospongia. The status and subdivi- The orders of Stromatoporoidea are
sions of this extinct phylum are taken accepted from Shrock & Twenhofel (1953)
from Okulitch in the Treatise (E, 1955). after Kuhn (1939). Other recent works di-
The oldest of the three names for the phy- vide the group into families, using no or-
lum is accepted here. (Archaeocyatha ders. The orders of Hydrozoa are those of
remains as a class). Hyman (1940) plus the extinct Spongio-
morphida. The orders of Scyphozoa are
Mesozoa. In adopting this phylum most of those of Hyman (1940) plus the extinct
the views of Hyman (1940) are accepted, Lithorhizostomeae, being thus those listed
except that the two orders are deemed to in the Treatise (F) after the removal of the
be amply distinct in basic development and Conularida. The orders of Anthozoa are
histology to be treated as classes. This is
those of the Treatise, and they are those of
the arrangement of Lankester (1901). It is Hyman (1940) except for the Cerianti-
also believed that Lankester was justified patharia and the extinct orders.
in separating the Heterocyemida from the
other Dicyemida, and they are accordingly Ctenophora. The classification of Hyman
given ordinal rank; for this class the name (1940) is accepted both as to classes and
Rhombozoa is available. orders. Nearly all recent works agree on
this arrangement.
Monoblastozoa. (See remarks under
Phyla, above.) Platyhelminthes. Three classes are gener-
ally recognized here, but the most recent
Graptozoa. The orders are taken from the
monographic work on the tapeworms
Treatise (V, 1955).
(Wardle & McLeod, 1952) seems to justify

Conularida. This arrangement is taken the recognition of the Cestodaria as a class


from the Treatise (F, 1956), but the treat- distinctfrom the Cestoda. Hyman (1951)
ment as a phylum is new here (see remarks included these in the Cestoda as a subclass,
under Phyla, above). but she found them sufficiently distinct to
require separate treatment in all respects
Coelenterata. The classification of the from the rest of the tapeworms (Euces-
classes of this phylum adopted here is the toda).
usual one except for two features: 1] two At one time the Temnocephaloidea
extinct classes are added, and 2] one were treated as a class intermediate between
group often listed as an order or subclass Turbellaria and Trematoda. Although it
is given class rank. The first two classes has been claimed that this arrangement is
are dealt with as in the Treatise (F). The now abandoned by all workers, it does re-
third one requires discussion here. appear in Dawes' (1946) monographic
Stromatoporoidea. This group has re- study of the Trematoda. In deference to
cently been included in the Hydrozoa. The Hyman's studies on the Turbellaria, the
principal arguments in favor of this seem group is herein placed in the Turbellaria
to be that there are other Hydrozoa show- as a suborder of Rhabdocoela.
ing some of the same peculiar colonial fea- The orders of Trematoda are taken
tures. Itappears that this is an argument from Dawes (1946), of Cestoda and Ces-
for re-examining these other groups (such todaria from Wardle & McLeod (1952),
as Spongiomorphida), because their pre- and of Turbellaria from Hyman (1951).
served hard-parts show few features of
Recent Hydrozoa. It seems best to empha- Rhynchocoela. Many recent works have
size the considerable structural differences divided this phylum two classes, the
into
between stromatoporoids and typical hy- Enopla and the Anopla. While accepting
drozoans by not merging them in one class. this subdivision, Hyman (1951) considers
The only reasons that can be given "the great similarity of structure through-
for retaining the stromatoporoids in the out the phylum" as reason for not making
Coelenterata while removing the grapto- these two groups classes. She therefore lists
lites from that phylum are that the grap- them as subclasses, there being no class
tolites form a somewhat more distinct mentioned. There is no rule preventing the
group and that the recent extreme diver- subdivision of a phylum directly into sub-
gence in views on their position in the Ani- classes, but it is unfamiliar and disconcert-
mal Kingdom lend credence to their more ing.
isolated position. Retention of the stroma- The features cited by Hyman as dis-
toporoids does not at present alter the defi- tinguishing the two subclasses seem to be
nition of the Hydrozoa. no more fundamental than those used for
.

CLASSIFICATION OF THE ANIMAL KINGDOM 32


distinguishing the orders. therefore
It is Priapidoidea. The three known species be-
here considered preferable to consider the long to two genera. There appears to be
nemertines as consisting of a single class no basis for separating these at the ordi-
of four orders. (The orders are those cited nal level (Hyman, 1951), and therefore
byHyman, 1951.) there is a single class with one order.
Inasmuch as there are two well-known
names available for this one-class phylum, Nematoda. Although the treatment of
it seems reasonable to retain one for the
this group as a phylum differs from Hy-
phylum and the other for the class. There man's (1951) treatment of it as a class of
is little reason to choose either way, but Aschelminthes, her view is accepted that
Hyman's argument that Schultze (1850- there are no subdivisions worthy of rank
51) was the "zoologist who first clearly above the ordinal level. The single class
understood the group" may be used as can be distinguished from the phylum by
adopting Schultze's name,
justification for the older but less familiar spelling Nema-
Rhynchocoela, for the phylum. toidea.

Gordiacea. There appears to be little of


Acanthocephala. This is another one-
basic nature in the differences between the
class phylum, for which no class names are
available. The orders are listed as in Hy- Gordioidea and the Nectonematoidea. Hy-
man (1951).
man (1951) is therefore followed in plac-
ing these as orders, although in the status
Rotifera. The decision made above, not of the group as a whole a different view is
to employ the Aschelminthes for six groups adopted (see Aschelminthes, above).
of pseudocoelomate animals, results in
elevation of these six groups to phyla. This
Calyssozoa / Endoprocta. Asingle class
and order make up this phylum, and the
raises the question of whether the orders of
only questions which arise are about the
the former class Rotifera should be raised
names to be used. The order has been
to classes. This has been done by some
called Pedicellinida by Boettger (1952),
classifiers, but there is considerable hesi-
the name Entoprocta or Endoprocta has
tation to doing so here. The Seisonidea
generally been used for the class, and the
appear from Hyman's remarks to be suffi-
first name proposed for the phylum is
ciently distinct to be considered a separate
Calyssozoa of Clark (1921). It seems least
class, but it is not so clear that Bdel-
confusing to accept these rather than dup-
loidea and Monogononta can be distin-
licateone name at several levels. (The
guished by equally fundamental characters.
spellingEndoprocta is here preferred over
In this dilemma, the three groups are ten-
Entoprocta because of its greater difference
tatively treated as classes, with the three
groups within the Monogononta treated as
from Ectoprocta.
orders. Bryozoa. Hyman's (1959) division of this
phylum into two classes with six orders,
Gastrotricha. The differences between the
following many earlier workers, is ac-
two groups of gastrotrichs, as described by
cepted here, as in most current paleonto-
Hyman, including the protonephridia,
logical works.
pharyngeal pores, and the body cavity sub-
The single order of the Phylactolae-
division, appear to justify the elevation of
mata seems to be without a name. The
the two groups to the level of classes. Each
name first used for the group was Lopho-
then consists of a single order.
poda, abandoned by later workers in favor
of Phylactolaemata. It is here revived for
Kinorhyncha. Although this group is here
the ordinal level.
considered to be a distinct phylum rather
The argument for replacing Bryozoa
than a class of Aschelminthes, there seems
with Ectoprocta because of removal of the
to be no reason for not following Hyman
Endoprocta has been answered above un-
(1951) in considering its subdivisions as
der Coelenterata. Removal of one group is
of less than ordinal rank. The characters
not considered justification for changing
distinguishing the three subdivisions are
the name of a phylum (or other taxon)
principally matters of degree, including
no fundamental clear-cut distinctions. Phoronida. The two genera seem to be-
As the phylum must contain at least long to a single order, for which there is
one class, there seems to be no reason for no special name (Hyman, 1959).
not using the name Echinodera at this
level. It remains effectively a synonym of Brachiopoda. There appears to be univer-
Kinorhyncha. sal agreement as to the division of this
33 Notes on Subkingdoms and Phyla
phylum into two classes. Six pairs of names disagreement on the division of this class
have been proposed for these classes, of into two orders. sometimes treated as
It is

which Inarticulata and Articulata are fa- a subclass or even as an order, of Amphi-
vored in most recent works. The Inarticu- neura or Gastropoda.
lata are usually divided into two orders, Gastropoda. Nearly all recent writers
and the Articulata were formerly divided agree on the division of this class into three
into two or three orders. There is no gen- subclasses, following Thiele (1931). The
eral agreement on the orders of Articulata, first, Prosobranchia, has been widely di-

and the opinion of Hyman (1959) and vided into three orders, but Cox (1960) in
others that there is no satisfactory classifi- the Treatise (I), has combined two of these
cation is accepted here. However, one of under the new name Caenogastropoda.
the older orders is still acceptable to pale- His new arrangement is followed here, al-
and there is no satisfactory
ontologists, though in other respects Thiele's classifica-
method for combining the remaining two. tion is accepted. A
recent classification by
The older division three orders is
into Taylor & Sohl (1962) is not followed
therefore followed, until a clear alternative here because it is not accompanied by jus-
is available. tification at the ordinal level.
Bivalvia / Pelecypoda. A
considerable
MoUusca. Only one recent American variety of classifications are in use for this
work is known to list seven classes of Mol- class, •with little obvious correlation of

lusca as is done herein. There is difference groupings. At the present time it appears
of opinion among modern workers only as best to present the more common arrange-
to the status of the groups sometimes re- ment of neontologists and also the custom-
ferred to as Isopleura. Many texts have ary paleontological one (in the footnotes),
omitted the fossil Monoplacophora and until a single scheme has been accepted by

treated the Solenogastres as an incertae both groups of workers. The first of these is
sedis group. This leaves the Amphineura the scheme of Lankester (1906) and most
or Placophora as a fifth class. It also leaves later textbooks. The second is the scheme

the classification of the phylum incomplete of Cotton & Godfrey (1938) with the or-
and therefore unsatisfactory. ders raised to subclasses as by Cox (1960).
volume I of the Treatise of Inverte-
In (Cox's more numerous orders may repre-
brate Paleontology, Yonge (1960) dis- sent a more natural arrangement, but it is
cusses the features of these groups and con- not yet known whether they will be accept-
cludes that there are seven classes. This able to other workers.)
arrangement is followed also in the Traite Scaphopoda. Apparently no names are
de Zoologie and is accepted herein. available for the single class and the single
Monoplacophora. Until 1957 this order. Therefore, Scaphopoda is here used
class was known only as fossils. Its divi- for all three levels, as they are coextensive.
sion into three orders is taken from the Cephalopoda. There is little agree-
Treatise (I, 1960). ment among recent workers as to the sub-
Amphineura / Polyplacophora. There division of this class, although many of the
are three substantially different classifica- subgroups are found in all schemes. The
tions of this group among recent works. arrangement followed here is that of Cot-
These are: orders Lepidopleurida and
1]
ton & Godfrey (1940) and many of the
Chitonida, by Thiele (1935), etc.; 2] or- older textbooks. The numerous nautiloid
ders Eoplacophora, Mesoplacophora, Iso- "orders" of Flower & Kiimmel (1950) are
placophora, and Teleoplacophora, by Cot- based largely on relative characters and
ton & Godfrey (1940); and 3] orders
have not been clearly established as enti-
Paleoloricata and Neoloricata, by Smith tled to ordinal rank.

(1960). There seems to be little direct cor-


relation between these systems. Sipunculoidea. The ten genera of this phy-
Smith's system is ostensibly based on lum are not separated into classes or or-
Pilsbry's early classification, modified by ders. Separate names are not available for

paleontological data. It is unfamiliar to the resulting single class and order.


neontologists, but it may be readily under-
stood when it is seen that all living chitons Echiuroidea. This phylum, which has of-
are placed in the order Neoloricata (to ten been placed in or appended to the An-
which many fossil forms belong as well). nelida, is divided into two classes as in
It is adopted here as the best available Boettger (1952). The class Echiurida is di-
classification of a neglected group. vided into three orders as in Pearse
Aplacophora. There appears to be no (1949).
CLASSIFICATION OF THE ANIMAL KINGDOM 34
Myzostomida. The two orders are ac Onychophora. Division of the phylum
cepted from Prenant in the Traite de Zo- into two orders is accepted from Boettger
ologie (1959). (1952) and the Treatise (O, 1959), fol-
lowing earlier writers.
Annelida. Many recent books
the list

Polychaeta and Oligochaeta as separate Arthropoda. Some recent classifications


classes. The differences between the two of this largest of all phyla have been com-
are largely relative, and the two together plicated by attempts to include the Penla-
can be described in detail with few con- stomida, Onychophora, and even the Tar-
flicts. For this reason, the two groups are digrada. When this is done, the phylum
here combined in the class Chaetopoda, can no longer be defined, because it would
as in many of the older works. Their dif- include a variety of body cavities, nervous
ferences may then be brought out at the systems, respiratory systems, excretory
subclass level. systems, integuments, etc. With these
No really satisfactory classification of groups removed, the arthropods can read-
this phylum or its classes has been found. ily be classified into eleven classes, and
Polychaeta. The usual division into these can be grouped into three subphyla.
two orders has been followed, with the ad- Much of the classification of the
dition of one extinct order. Arthropoda above the ordinal level is in an
Oligochaeta. There being no generally unsettled state. There seems to be no sin-
accepted subdivision of this group, the one gle complete classification that is adequate
given without Latin names by Avel (1959) for both Recent and fossil groups. The one
in the Traite (attributed by him to Mi- presented here is therefore made up from
chaelson, 1930) and without acceptable many most of which are at least
sources,
single names by Pearse (1949) is followed in part drawn from earlier sources. Listed
here. The names are emended to conform below are the recent works that are ac-
to custom nomenclature.
in Latin cepted at each level, but they usually are
Hirudinea. Division of this class into not the original proposal of the arrange-
four orders was proposed by Harant & ment.
Grasse (1959) using three orders pro- Subphyla are accepted from Pearse
posed by Caballero (1952) and adding the (1949), classes in the Trilobitomorpha
"Acanthobdelliformes nom.nov." These from the Treatise (O, 1959), in the Che-
names and groupings were not new there, licerata from Pearse (1949) and Moore
except in ending. Rhynchobdellida dates (1952), and in the Mandibulata from
from Blanchard (1887), Gnathobdellida Pearse (1949) with exclusion of super-
from Vaillant (1890), and Pharyngobdel- classes.
lida from Johannson (1913). These same Orders in the Trilobitomorpha are
four orders were recognized by Lowenstein from the Treatise (O, 1959), in the Mero-
(1954), using the name Acanthobdellida stomata from the Treatise (P, 1955), in
and using the name Herpobdellida instead the Pycnogonida from the Treatise (P,
of Pharyngobdelliformes. 1955), in the Arachnida from the Treatise
It is not clear whether these names (P, 1955) which is the system of Petrunke-
were independently proposed, but there vitch, in the Crustacea from Waterman &
seems to be no need for the -iformes end- Chace (1960), in the Pauropoda and Sym-
ings here. The four orders are accepted phyla from Brues & Melander (1954), in
here, the spellings of Caballero and of the Diplopoda from Boettger (1952), in
Harant & Grasse are rejected, and Pharyn- the Chilopoda from Pearse (1949), and in
gobdellida is accepted because of apparent the Insecta from Brues & Melander (1954).
priority. The Eurypterida consists of a
class
Archiannelida. There is apparently single order, for which the synonym Gi-
general agreement that there is only one gantostraca can reasonably be used.
order in this class. No ordinal name is
available.
Clicu'ioi^natlia. This
another of the one-
is

class one-order phyla. Boettger(1952) has


Tardigrada. The division into three orders used the name Sagittoidea for both class
in two classes is taken from Pearse (1949), and order, but it seems to be more appro-
after Richters (1926) and Marcus (1927). priate to restrict this name to the ordinal
level and to use the phylum name also at
Penlastomida. The orders are from Boett- the class level. The other two synonyms
ger(1952). seem to be inappropriate as they were
.

35
Notes on Suhkingdorns and Phyla
originally employed in somewhat different et al. (1952); and in the Holothurioidea,
context. from Shrock & Twenhofel ( 1 953 )

Pogonophora. This is probably the newest Pterobrancliia. Many writers agree on di-
phylum in point of knowledge of the ani- viding the pterobranchs into two groups.
mals themselves, as the first species was The difficulties of describing Rhabdopleura
described in 1914, and most of the present and Cephalodisciis together leads to skep-
day knowledge is less than a score of years ticism that they should be considered or-
old. Only one classification has been un- ders rather than classes. In deference to
dertaken, by Ivanov (1955), and his divi- workers in this field, the usual division is
sion into two orders is accepted here. adopted, with some misgivings (as in Hy-
Inasmuch as the synonym Brachiata is man, 1959).
not well known for these animals, it has
Enteropneusta. Only one class and one
not been used for the one class.
order is known in this group. It is some-
Echinodermaia. No single satisfactory clas- times left without a name, but Balanoglos-
sification has been found for this varied sida is available. It is here used for the or-
phylum. The subphyla here accepted (see der, as the phylum name is also familiar
footnotes) are those of Shrock & Twen- at the class level.
hofel (1953). The classes in the Pelmato-
Planctosphaeroidea. This phylum consists
zoa, Homalozoa, and Haplozoa are those
of a single species of what appear to be
of Shrock & Twenhofel (1953), being ex-
larvae.
tinct except for Crinoidea; in the
the
Eleutherozoa the of Shrock &
scheme
Tiinicata. There appears to be general
Twenhofel is modified according to the agreement on the assignment of the tuni-
views of Hyman (1955) to omit the Stel-
cates to three major classes. The subclasses
leroidea and treat Asteroidea and Ophiu-
and orders accepted here are those of
roidea as classes. This makes it necessary
Pearse (1949) and others, with addition
also to treat as classes the extinct groups
to the Ascidiacea of an order Octacnemida
Auluroidea and Somasteroidea. This ar-
after Harant (1948).
rangement is admittedly a compromise
with more familiar classifications. Cephalochordata. This group, which is
This phylum contains a large number usually included in the Chordata, has at
of extinct subgroups, and it is difficult to least four available names. Those adopted
give full attention to the extinct sub- are the ones most often used at the respec-
groups without confusing the classification tive levels.
of Recent ones. The orders accepted here
are as follows: In Pelmatozoa, Homalozoa, Vertebrata. The restriction of this phy-
and Haplozoa, from Shrock & Twenhofel lum to the craniates explained in a pre-
is

(1953); in Asteroidea, from Hyman vious section. The classes of Vertebrata are
(1955); in Ophiuroidea, from Boettger taken from Romer (1945) and Colbert
(1952) except for removal of Auluroidea; (1955). The orders of these classes are
in Echinoidea, from Shrock & Twenhofel taken from these same works, except that
(1953) for the Regularia and from Hy- Romer is generally followed where he
man (1955) for the Irregularia (except for treats as orders the groups which Colbert
removal of Bothriocidaroida after Moore lists as subclasses or superorders.
Complete List

Subk Series Phylum

AN MALIA
I

(Zoa)

EOZOA (Plastidozoa)
Protozoa cam-rec
PARAZOA
Porifera (Spongiario, Spongeae) cam-rec
Cyathospongia (Pleospongia, Archaeocyatha) cam
AGNOTOZOA
Mesozoa (Plamdoidea, Mionelminthes , Moruloidea rec
HISTOZOA (Metazoa, Eumetazoa)
Enterocoela (Enterozoa, Protaxonia)
Monoblastozoa rec
Graptozoa cam-mis
Conularida cam-tri
Coelenterata (Cnidaria, Nematozoa, Nematophora) cam-rec
Ctenophora {Acnidaria, Collaria, Ctenarea, Ctenoph- rec
oraria)
Acoelomata
Platyhelminthes (Platodes) rec
Rhynchocoela (Nemertinea, Aplocoela, Miocoela) rec
Pseudocoelomata {Pseudocoelia, Autoscolecida, Protoneph-
ridozoa)
Acanthocephala rec
Rotifera (Rotatoria) rec
Gastrotricha rec
Kinorhyncha (Echinodera) rec
Priapuloidea rec
Nematoda (Nemata) rec
Gordiacea (Gordioidea, N ematomorpha) car-rec
Calyssozoa (Entoprocta, Endoprocta, Kamptozoa) rec
of Phyla'

Subk Series Phylum

Coelomata (Eucoelomata)
Bryozoa (Polyzoa, Ectoprocta) ord-rec
Phoronida REC
Brachiopoda (Spirobranchia, Palliobranchiopoda) CAM-REC
Mollusca (Palliata, Malacozoa, Saccata, Heterogan- cam-rec
gliata, Otocardia)
Sipunculoidea REC
Echiuroidea REC
Myzostomida jur-rec
Annelida (Annulata, Coelhelminthes) cam-rec
Tardigrada Rec
Pentastomida (Linguatulida) REG
Onychophora (Polypoda, Protracheata, Ceratophora, cam-rec
Malacopoda)
Arthropoda (Euarthropoda) cam-rec
Chaetognatha (Homalopterygia, Oesthelminthes) cam-rec
Pogonophora (Brachiata) REC
Echinodermata cam-rec
Pterobranchia ord-rec
Enteropneusta (Helminthomorpha) REG
Planctosphaeroidea REG
Tunicata (Urochordata) REG
Cephalochordata (Acrania, Cirrhostomi, Entomocra- REC
nia, Haplocyemata, Homomeria, Myelozoa, Pharyngo-
branchii)
Vertebrata (Euchorda, Craniata) ord-rec

^
For explanations, see preceding section.
Complete List of Classes and Orders

w\th Synonyms, Subgroups, and Geologic Range

Class Subcl Order

PROTOZOA^
FLAGELLATA ^ (Mastigophora) sil-rec
Phytomastigina ^ (Phytomastigophorea) sil-rec
Chrysomonadina {Silicoflagellata, Chrysomonada- cre-rec
ceae)
Coccolithophorida sil-rec
Cryptomonadina (Cryptomonadaceae) rec
Phytomonadina ( Volvocales, Volvocina, Volvocaceae) eoc-rec
Euglenoidina (Euglenida, Euglenaceae) eoc-rec
Chloromonadina (Chloromonadaceae) rec
Dinoflagellata * (Cilioflagellata, Catenata) jur-rec
Zoomastigina ^ (Zoomastigophorea) rec
Rhizomastigina {Pantostomatida, Rhizomastigaceae rec
Rhizofiagellata)
Protomonadina (Protomastigida, Protomastigaceae) rec
Polymastigina rec
Hypermastigina {Holomastigina) rec
SARCODINA CAM-REC
^
Rhizopoda ord-rec
Proteomyxa rec

^ Synonyms: Eozoa, Plastidozoa. Subphyla: Plasmodroma (Homokaryota,


Gymnomyxa, Cytomorpha) = flagellata + sarcodina + sporozoa. Ciliophora
(Infusoria, Heterokaryota, Cytoidea) = ciliata + suctoria. Includes: Corticata
(flagellata + SPOROZOA + CILIOPHORA).
^ Includes: Lissoflagellata, Monadidea, Heteromastigina, Chromomonadina,
Choanoflagellata, Craspedomonadina, Phalansteriina, Phytoflagellata, Chlamydo-
monadina, Euflagellata.
^ Includes: Heterochlorida.
^ Includes: Prorocentraceae, Adinida, Peridiniaceae, Gymnodiniaceae, Di-
niferidea, Cystoflagellata, Rhynchoflagellata.
^ Includes: Trichomonadida.
^ Includes: Labyrinthulidea, Lobosa, Filosa.
"

COMPLETE LIST OF CLASSES AND ORDERS 40


Class Subcl Order

Mycetozoa ''
{Myxomycetes, Myxogasteres) REC
Amoebozoa (Amoebina, Amoebaea, Nuda, Gymna- rec
moebaea)
Testacea (Thecamoebae) eoc-rec
Foraminifera ® {Reticularia, Polythalamia, Thalamo- ord-rec
phora)
Actinopoda cam-rec
®
Heliozoa ple-rec
Radiolaria " cam-rec
(Po/jc}75fma, Cytophora, Echinocystida)
SPOROZOA "' (Gregarina) REC
Telosporidia ^^ (Cytosporidia, Ectospora, Amoebosporidea) rec
Gregarinida rec
Coccidia ^* (Coccidiomorpha) REC
Haemosporidia ^^ (Haemocytozoa, Acystosporidia, rec
Gymnosporidia )
Cnidosporidia REC
Myxosporidia (Amoebogeniae) REC
Actinomyxidia REC
Microsporidia REC
Helicosporidia REC
Sarcosporidia (Sarcocysddea) REC
Sarcosporidia REC
Globidia REC
Haplosporidia (Aplosporidia) REC
Haplosporidia REC

"^Includes: Euplasmodida, Eumycetozoa, Sorophora, Acrasiae, Amauro-


chaetaceae, Amaurochaetineae, Amaurosporales, Anemineae, Arcyriaceae, Cal-
carineae, Calonemineae, Ceratiomyxaceae, Dlctyosteliaceae, Didymiaceae, Endo-
sporeae, Exosporae, Guttulinaceae, Heterodermaceae, Lamprosporales, Liceaceae,
Lygogalaceae, Margaritaceae, Physaraceae, Reticulariaceae, Stemonitaceae, Tri-
chiaceae, Tubulinaceae.
^ Includes: Allogromidiaceae, Astrorhizidea, Cheilostomellaceae, Chilosto-
mellidea, Globigerinidea, Gromiidea, Lagenidea, Lituolidea, Miliolidia, Num-
mulitidea, Rotalidea, Textularidea, Xenophyophoridea.
^ Includes: Actinophrydea, Aphrothoraca, Centrohelidia, Chalarothoraca,
Chlamydophora, Desmothoraca, Helioflagellida.
'^°
Includes: Acantharia, Acanthometrida, Acanthophractida, Actipylaea,
Cyrtellaria, Botryoidea, Cyrtoidea, Spyroidea, Monopylaria, Nasselaria, Osculo-
sida, Merotrypasta, Peripylaria, Spumellaria, Collodaria, Sphaerellaria, Sphaero-
zoa, Plectellaria,Plectoidea, Stephoidea, Porulosida, Holotrypasta, Tripylaria,
Phaeodaria, Phaeoconchia, Phaeocystina, Tripylaea, Phaeogromia, Phaeo-
sphaeria).
^^ Neosporidea (Histosporidia, Endospora) = cnidosporidia + sarcospori-
dia + haplosporidia. Acnidosporidia = sarcosporidia + haplosporidia.
41 Protozoa — Porifera
Class Subcl Order

CILIATA JUR-REC
Protociliata (Ciliatoidea) rec
Opalinida rec
Euciliata jur-rec
Holotricha ^® (Aspirigera, Aspirotricha) rec
^^
Spirotricha (Spirigera) jur-rec
Chonotricha rec
Peritricha rec
SUCTORIA (Acineta, Acinetaria, Tentaculifera) rec
Suctoria rec
PORIFERA ^«

CALCAREA ^^
(Calcispongea) cam-rec
Solenida (Asconosa) cam-rec
Lebetida (Syconosa) jur-rec
Pharetronida per-rec
Thalamida (Sphinctozoa) pen-cre
HYALOSPONGEA ^°
(Hexactinellida, Triaxonidd) cam-rec
Lyssakina (Lyssacina) cam-rec
Dictyonina (Dictyida, Inermia) ord-rec
Lychniskophora (Lychniskida) jur-rec
Heteractinida cam-car
DEMOSPONGEA ^^
(Desmospongea) cam-rec
Myxospongida rec
^^
Keratosida {Ceratospongida, Ceratosa, Ceratosida, car-rec
Euceratosa)
^^ Includes: Rhabdogeniae, Serosporidia, Exosporidia, Piroplasmidea.
^^ Includes: Eugregarinida, Schizogregarinida.
^^ Includes: Adeleida, Eimeriida.
^^ Includes: Babesiida, Plasmodiida.
'^^
Includes: Apostomina, Aspirotrichaceae, Astomata, Astomina, Gymnosto-
mata, Hymenostomata, Thigmotrichina, Trichostomata.
" Includes: Ctenostomina, Entodiniomorphina, Heterotricha, Hypotricha, 0!i-
gotrichina, Polytricha, Tintinnina.
^^ Synonyms: Spongiaria, Spongeae. Includes: Octactinellida, Heteractinel-
lida. Silicospongiae (Non-calcarea) = HEXACTiNELLroA + demospongea. Niduli-
tida,sometimes included here, is probably a calcareous alga.
Asconosa, Syconosa, Leuconosa are structural types, not groups. Includes:
^^

Sycones, Dialytina, Lithonina, Homocoelida, Heterocoelida.


^° Includes: Hexasterophora, Amphidiscophora,
Uncinataria.
21
Includes: Tetractinellida, Tetraxonida, Desmophora, Monaxonida, Sigmato-
monaxonellida, Monactinellida.
22
Considered by some to belong to Hadromerida. Sometimes placed with
some others of Demospongea as Cornacuspongia (with orders Protorhabdina,
Poikilorhabdina, Phthinorhabdina, Aporhabdina). Includes: Dictyoceratina, Mo-
noceratina, Dendroceratina, Hexaceratina.
COMPLETE LIST OF CLASSES AND ORDERS
43 Cyathospongia — Coelenterata
Class Subcl Order

GRAPTOZOA
GRAPTOLITHIDA CAM-MIS
Dendroidea CAM-MIS
Graptoloidea ORD-SIL
Tuboidea ORD-SIL
Camaroidea ORD
Stolonoidea ORD

CON U LARI DA

CONULATA CAM-TRI
Conulariida CAM-TRI

COELENTERATA
PROTOMEDUSAE
COMPLETE LIST OF CLASSES AND ORDERS 44
Class Subcl Order

SCYPHOZOA ^^
(Scyphomedusae, Acalephae, Neoscyphozoa, jur-rec
Acraspeda)
Stauromedusae (Lucernaria, Lucernariidea, Cyclico- rec
zoa, Calycozoa)
Cubomedusae (Carybdeida, Charybdeida, Marsupi- jur-rec
alia)

Coronatae (Peromedusae, Corona, Coronatida) jur-rec


Semaeostomeae (Semostomeae, Semaeostomatida) jur-rec
Lithorhizostomeae (Lithorhizostomatida) JUR
Rhizostomeae (Rhizostomatida) jur-rec
ANTHOZOA (Endoaria, Actinozoa, Actinoidea, Oecioa, Polycy- cam-rec
clia, Corallaria, Monocyclica, Coralligena, Corolla,
Scyphopolypi, Anthozoariae)
Alcyonaria ^^ {Octocorallia, Octactinia, Zoophytaria) per-rec
Stolonifera cre-rec
Telestacea REC
Alcyonacea jur-rec
Trachypsammiacea per
Coenothecalia cre-rec
Gorgonacea cre-rec
Pennatulacea cre-rec
Zoantharia ^* (HexacoralUa, Helianthoida, Zoanthactiniaria, cam-rec
Actinanthida, Dodecacorallia, Zoantha)
Zoanthiniaria (Zoanthidea) rec
Corallimorpharia (Stichodactylina, Asclerocorallia) rec
Actiniaria (Actiniidea, Malacactiniae Edwardsiidea) cam-rec
,

Rugosa (Stauracea, Pterocorallia, Tetraseptata, Tetra- ord-per


coelia, Tetracorallia)

Heterocorallia (Dicoelia) car


Scleractinia (Madreporaria, Poly actinia, Cyclocoral- tri-rec
lid)

Tabulata (Aseptata, Trichocorallia, Tubulosa, Tetra- ord-per


dida, Schizocoralla, Heliolitida, Multisolenida, Thal-
locoralla, Chaetetida)

Discomedusae (Discophora)
^2 = coronatae + semaeostomeae + rhizo-
stomeae.
^^ Includes: Pseudaxonia, Axifera, Stelechotokea, Protalcycnacea, Synal-
cyonacea.
^* Includes: Paramera, Cryptoparamera, Ptychodactiaria, Proactiniae.
Synonyms: Acnidaria, Collaria, Ctenophoraria, Ctenarea.
^^

Synonyms: Platodes, Platyelmia, Plathelminthes, Platyhelmia. Cestoidea


^® =
CESTODA + CESTODARIA.
45 Coelenterata — Platyhelminthes
Class Subcl Order

Ceriantipatharia mio-rec
Antipatharia (Hexactinia, Anticorallia, Antipathidea) mio-rec
Ceriantharia (Paranemata, Tetractiniae, Heterocoral- rec
la, Cerianthiae, Cerianthidea)

CTENOPHORA^'
TENTACULATA (Micropharyngea) REC
Cydippida (Saccata) REC
Lobata (Bolinopsidea) rec
Cestida (Cestoidea) REC
Platyctenea (Tjalfiellidea, Ctenoplanidea) REC
NUDA {Macropharyngea, Atentaculata) REC
Beroida REC

PLATYHELMINTHES'"'
TURBELLARIA (Planaria) REC
Acoela rec
^^
Rhabdocoela rec
Alloeocoela (Alloiocoela) rec
Tricladida REC
Polycladida (Cryptocoela) rec
TREMATODA rec
Monogenea ^^
(Monogenetica, Hetewcotylea, Etero- rec
cotylea, Herocotylida, Polystomea, Pectobothrii, Cryp-
tocoela, Ectoparasitica)
Aspidogastrea (Aspidobothria, Aspidocotylea) rec
Digenea ^^ (Digenetica, Malacocotylea, Distomea, Ma- rec
lacobothrii, Strigeata)

CESTODA *° (Eucestoda, Cestoidea, Merozoa, Polyzoa, Tomioso- rec


ma)
Proteocephala (Proteocephaloidea) rec
Tetraphyllidea (Phyllobothrioidea) rec
Disculicepitidea rec
Lecanicephala {Lecanicephaloidea, Diphyllidea) rec
"^
Includes: Temnocephaloidea, Dactylifera, Catenulida, Macrostomida.
^* Includes: Monopisthocotylea, Polyopisthocotylea.
^^ Includes: Gasterostoma, Prosostomata.
^° Includes: Aphylles, Monophyllidea, Heterophyllidea, Cystica, Caryophyl-
lacea.
COMPLETE LIST OF CLASSES AND ORDERS 46
Class Subcl Order

Trypanorhyncha {Tetrarhyncha, Tetrarhynchoidea, rec


Phyllorhyncha)
Cyclophyllidea (Taenioidea, Tetracotylea, Tetrabothri- rec
diata)
Aporidea REC
Nippotaeniidea REC
Caryophyllidea REC
Spathebothridea REC
Pseudophyllidea (Bothriocephaloidea, Dicestoda, Di- rec
bothridiata)
CESTODARIA (Monozoa, Atomiosoma) REC
Amphilinidea REC
Gyrocotylidea REC
Biporophyllidea REC

RHYNCHOCOELA"
NEMERTINEA *==
(Nemertea) REC
Palaeonemertea (Paleonemertea, Palaeonemertini) rec
Heteronemertea {Schizonemertini, Trimyaria, Eupoli- rec
id a)

Hoplonemertea (Metanemertini) rec


Bdellonemertea (Bdellomorpha) rec

ACANTHOCEPHALA
ACANTHOCEPHALA ''
REC
Archiacanthocephala REC
Palaeacanthocephala rec
Eoacanthocephala ^ rec

ROTIFERA*'
SEISONIDEA REC
Seisonacea rec
BDELLOIDEA REC
Bdellacea (Bdelloidaceae) rec
" Synonyms: Aplocoela, Miocoela.
*^ Includes: Dimyaria, Protonemertini, Mesonemertini, Anopla, Enopla.
^^ Metacanthocephala = archiacanthocephala + palaeacanthocephala
** Includes: Gyracanthocephala, Neoacanthocephala.
*^ Synonyms: Rotatoria. Digononta = seisonidea + bdelloidea.

^'^
Includes: Rhizota.
47
.

COMPLETE LIST OF CLASSES AND ORDERS 48


Class Subcl Order

Oxyuroidea REC
Ascaroidea REC
Strongyloidea REC
Spiniroidea (Camallanta) REC
Dracunculoidea REC
Filarioidea REC
Trichuroidea ( Trichinelloidea) REC
Dioctophymoidea REC

GOR DIACEA
NEMATOMORPHA CAR-REC
Gordioidea (Gordididea) CAR-REC
Nectonematoidea REC

CALYSSOZOA^^
ENDOPROCTA (Entoprocta) REC
Pedicellinida REC

BRYOZOA'^
PHYLACTOLAEMATA (Hippocrepia) CRE-REC
Lophopoda CRE-REC
GYMNOLAEMATA '*
(Stelmatopoda) ORD-REC
Trepostomata (Tripostomata, Monticuliporoided) ORD-PER
Cryptostomata ORD-PER
Cyclostomata (Stenostomata) ORD-REC
Ctenostomata ORD-REC
Cheilostomata (Chilostomata) CRE-REC

PHORON DA I

PHORONIDA REC
Phoronida REC
^2 Synonyms: Kamptozoa (see also class names)
^^
Synonyms: Polyzoa, Ectoprocta. Formerly included also Endoprocta; to-

gether called also Holobranchia.


^* Stenolaemata = trepostomata + cyclostomata. Cheiloctenostoma (Eu-
rystomata) = ctenostomata + cheilostomata.
^^ Synonyms: Spirobranchiopoda, Palliobranchiopoda, Branchiopoda, Bra-
chionopoda, Brachionocephala, Branchionobranchia, Spirobranchia.
^*'
Includes: Orthida, Strophomenida.
" Includes:l?entainQnda, Rhynchonellida, Spiriferida, Terebratulida, Triplesiida.
^^ Synonyms: Palliata, Malacozoa, Heterogangliata, Otocardia, Saccata. Glos-
sophora (Cephalophora) = amphineura + gastropoda + cephalopoda + sca-
"

49 Gordiacea — Mollusca
Class Subcl Order

BRACHIOPODA'^
INARTICULATA (Ecardines, Lyopomata, Pleuropygia, Gastrocau- cam-rec
lia, Tretenterata, Sarcobranchiata)
Atremata cam-rec
Neotremata cam-rec
ARTICULATA (Testicardines, Arthropomata, Apygia, Pygocaulia, cam-rec
Clistenterata)
Palaeotremata {Paleotremata) CAM

Protremata cam-rec
Telotremata cam-rec

MOLLUSCA ^«

MONOPLACOPHORA ^^
{Protogastropoda, Amphigastropoda) cam-rec
Tryblidioidea {Tryblidiacea) cam-rec
Archinacelloidea cam-sil
Cambridioidea cam
AMPHINEURA ^°
(Aculifera, Polyplacophora, Loricata, Crept- cam-rec
poda, Polyplakiphora, Polyplaxiphora, Placophora,
Polybranchiata, Lamellata, Lepidoglossa)
Paleoloricata cam-cre
Neoloricata car-rec
APLACOPHORA {Solenogastres, Telobranchia, Scolecomorpha) rec
Neomeniida (Neomeniomorpha) rec
Chaetodermatida (Chaetodermomorpha) rec
GASTROPODA*'^ {Anisopleura, Paracephalophora, Pselapho- cam-rec
cephala)
Prosobranchia (Streptoneura, Cochlides) cam-rec
^^
Archaeogastropoda cam-rec

PHOPODA. Stenolaemata = amphineura + gastropoda + bivalvia + scaphopoda.


Prorhipidoglossomorpha = gastropoda + bivalvia + scaphopoda. Isopleura =
MONOPLACOPHORA + AMPHINEURA + APLACOPHORA.
^® Includes: Cynostraca, Cochliostraca.
^^ Includes: Lepidopleurida, Chitonida, Eoplacophora, Isoplacophora, Meso-
placophora, Teleoplacophora.
'^^
Also spelled: Gasteropoda, Gastraeopoda, Gasteropodophora, Gasterozoa.
Includes: Spironotia. Euthyneura (Androgyna, Platymalakia) = opisthobranchia
+ pulmonata,
Includes: Scutibranchia, Aspidobranchiata, Diotocardia, Bellerophontacea,
'^'^

Cyclobranchia, Zygobranchia, Docoglossa, Rhipidoglossa.


COMPLETE LIST OF CLASSES AND ORDERS 50
Class Subcl Order

Caenogastropoda
^^
ORD-REC
Opisthobranchia
^* MIS-REC
Pleurocoela ^^ {Tectibranchiata, Tectobranchia, Aply- MIS-REC
siacea, Steganobranchia)

Pteropoda CRE-REC
Sacoglossa REC
Acoela ""^
EOC-REC
Pulmonata PEN-REC
Basommatophora PEN-REC
Stylommatophora PEN-REC
BIVALVIA ®^
(Pelecypoda, Lamellibranchiata, Aglossa, Lipoce- ORD-REC
phala, Conchijera, Conchophora, Acephala)
"*
Protobranchia ORD-REC
Filibranchia
'^^
ORD-REC

Eulamellibranchia SIL-REC
Septibranchia JUR-REC
SCAPHOPODA {Cirrhobranchiata, Solenoconchia, Prosopocepha- DEV-REC
la, Lateribranchiata)
Scaphopoda DEV-REC
CEPHALOPODA (Siphonopoda) CAM-REC
Tetrabranchiata (Schizosiphona, Tentaculifera) CAM-REC
^^
Nautiloidea CAM-REC
^^
Ammonitoidea ORD-CRE
Dibranchiata {Coleoidea, Acetabulifera, Holosiphona) MIS-REC
Decapoda ^^ (Decembrachiata, Decabrachia) JUR-REC
^^
Includes: Siphonobranchia, Pectinibranchia, Ctenobranchiata, Hemipoma-
tostoma, Apomatostoma, Monotocardia, Azygobranchia, Mesogastropoda, Steno-
glossa, Neogastropoda, Rachiglossa, Toxoglossa, Heteropoda, Taenioglossa, Platy-
poda, Rhachiglossa.
^Includes: Gymnosomata, Oncidiacea, Acochlidiacea, Vaginulacea, Ana-
spidea.
^^ Includes: Cephalaspidea, Thecosomata, Bullomorpha, Gyrosomata.
''^
Includes: Notaspidea, Pleurobranchomorpha, Nudibranchiata.
^''
Includes: Anisomyaria, Anatinacea, Cladocopa, Ambonodonta. Groups
sometimes recognized include: Siphonida, Asiphonida, Homomyaria, Macrocilio-
branchia, Microciliobranchia, Pteriomorpha, Colloconchida, Eutaxodonta, Isofili-
branchia, Pteroconchida, Rostroconchida, Heteroconchia, Eudesmodontida, Nai-
adida, Pachyodontida, Pantodontida, Asthenodontida, Schizodontida, Heterodon-
tida, Septibranchida. Also used as orders: Autobranchiata, Dimyaria, Eleuthero-
rhabda, Hemibranchia, Heteromya, Heteromyaria, Isedrolotila, Isomya,
Laternulacea, Macrotrachia, Monomyaria, Monomya, Neotaxodonta, Palaeo-
branchia, Palaeolamellibranchia, Parafilibranchia, Pholadacea, Pleuroconcha,
Pleurodonta, Praeheterodonta, Synaptorhabda, Veneracea.
51 Mollusca — Myzostomida
Class Subcl Order

Octopoda (Octobrachiata, Octopoida) cre-rec


Vampyromorpha rec
Belemnoida ^*
(Phragmophora) mis-eoc

SIPUNCULOIDEA
SIPUNCULOIDEA REC
Sipunculida REC

ECHIU ROI DEA


ECHIURIDA REC
Echiuroina rec
Xenopneusta rec
Heteromyota rec
SACCOSOMATIDA rec
Saccosomatida rec

MYZOSTOMIDA ^=

MYZOSTOMIDA pal-rec
Proboscidea rec
Pharyngidea rec

Arrangement preferred by paleontologists:


Subclass Prionodesmacea (orders Paleoconcha, Taxodonta, Schizodonta,
Isodonta, Dysodonta).
Subclass Anomalodesmacea (orders Septibranchia, Anomalobranchia).
Subclass Teleodesmacea (orders Pantodonta, Diogenodonta, Cyclodonta,
Teleodonta, Asthenodonta, Heterodonta, Pachyodonta, Desmodonta)
®* Sometimes a subclass with orders Cryptodonta, Lipodonta, Palaeotaxodonta.
^^ Includes: Pseudolamellibranchia.
^° Includes: Schizodonta, Heterodonta, Adapedonta, Anomalodesmata, Sinu-
palliata, Integripalliata.
^^ Sometimes ranked
as a separate subclass with orders Mixochoanites, Schi-
stochoanites, Orthochoanites, Cyrtochoanites; or with orders Ellesmeroceroida,
Michelinoceroida, Ascoceroida, Oncoceroida, Endoceroida, Actinoceroida, Disco-
soroida, Nautilida; or with orders Bassleroceratida, Barrandeoceratida, Centro-
ceratida, Rutoceratida, Solenochilida, Tarphyceratida.
^^ Sometimes ranked as a separate subclass with order Ammonitida; or with
orders Intrasiphonata, Extrasiphonata.
^^ Includes: Teuthoidea, Sepioidea, Loliginacea, Architeuthacea.
^* Sometimes used as subclass in place of Dibranchiata.
^° Synonyms: Myzostomaria, Myzostoma.
COMPLETE LIST OF CLASSES AND ORDERS 52
Class Subcl Order

ANNELIDA
CHAETOPODA "
53 Annelida — Arthropoda
Class Subcl Order

PENTASTOM DA I

LINGUATULIDA REC
Cephalobaenida REC
Porocephalida (Linguatulodea) REC

ONYCHOPHORA^^
PERIPATIDEA cam-rec
Protonychophora CAM
Euonychophora REC

ARTHROPODA «2

TRILOBITOIDEA ''
cam-pen
Merostomoidea (Xenopoda) cam
Limulavida {Prochelicerata, Limulava) CAM
Emeraldellida CAM
Nectaspida cam
Leanchoiliida (Pseudanostraca) CAM
Pseudonotostraca (Pseudocrustacea) cam
Burgessiida cam
Waptiida cam
Marrellomorpha ^ (Marellomorpha) CAM
Marrellida cam
Hymenocarina cam
Hymenocarina cam
Arthropleurida PEN
Arthropleurida PEN
Cheloniellida dev
Cheloniellida dev
Opabiniida CAM
Palaeanostraca cam
PHYLA + DiPLOPODA + CHiLOPODA + INSECTA. Hypaithropoda was proposed for
hypothetical ancestral forms. A separate class Arthrocephala
is listed by Boettger

in the Chelicerata, but no other reference to such an extinct group has been found.
Diantennata = trilobitomorpha + Crustacea. Branchiata = trilobitomorpha
+ CHELICERATA + MANDIBULATA. Accrata = MEROSTOMATA + ARACHNmA. Arach-
nomorpha = trilobitomorpha + Crustacea. Myriapoda = pauropoda + sym-
PHYLA + DIPLOPODA + CHILOPODA. PrOgOneata = SYMPHYLA + DIPLOPODA +
pauropoda. Antennata = pauropoda + symphyla + diplopoda + chilopoda +
INSECTA. Opisthogoneata = chilopoda + insecta. Includes: Marriocarida, of un-
known position.
^^ Homopoda = marrellina + pseudanostraca + pseudonotostraca +
hymenocarina.
^* Includes: Mimetasterida, Pygaspida, Proarthropoda.
COMPLETE LIST OF CLASSES AND ORDERS
55 Arthropoda
Class Subcl Order

Palpigradida {Microthelyphonida, Latisterna, Palpi- JUR-REC


gradi)
Thelyphonida (Uropygi, Holopeltidia) car-rec
Schizomida (Tartarides, Schizopeltidia, Colopyga, pli-rec
Schizonotida)
Kustarachnida (Kustarachne) PEN
Phrynichida (Amblypygi, Phryneides) CAR-REC
Araneida (Araneae) car-rec
Solpugida (Solifugae, Galeodea, Rostrata, Mycetopho car-rec
rae)
Ricinuleida {Rhinogastra, Meridogastra, Podogonata, car-rec
Cucullijera, Rhignogastra)
CRUSTACEA ''^
(Eucrustacea) CAM-REC
^*
Branchiopoda cam-rec
Anostraca (Euanostraca) EOC-REC
Lipostraca DEV
Notostraca PER-REC
Conchostraca DEV-REC
Cladocera REC
Cephalocarida REC
Cephalocarida REC
Ostracoda {Ostracopa, Ostrapoda) CAM-REC
^^
Archaeocopida (Archaeostraca) CAM-TRI
Leperditicopida ( Leperditiida ) ORD-DEV
Myodocopida (Myodocopa, Cladocopa) ORD-REC
Podocopida (Podocopa, Platycopa) ORD-REC
Palaeocopida (Paleocopa, Beyrichiida) ORD-PER
Mystacocarida REC
Mystacocarida REC
Copepoda (Eucopepoda) REC
Calanoida REC
Harpacticoida REC
Cyclopoida REC
^° Notostigmata, Cryptostigmata, Prostigmata,
Includes: Stomatostigmata,
Heterostigmata, Parastigmata, Mesostigmata, Metastigmata.
^^ Camarostomata = schizomida + thelyphonida + kustarachnida. Label-
lata = phrynichida + araneida.
^" Pedipalpida (scorpion-spiders) = schizomida + thelyphonida + phry-
nichida.
^^ Entomostraca = branchiopoda + ostracoda + copepoda + cirripedia.
^* Diplostraca = conchostraca + cladocera. Phyllopoda (Euphyllopoda,
Onychura) = anostraca + notostraca + conchostraca.
^^ Includes: Bradorina, Discinocarina.
COMPLETE LIST OF CLASSES AND ORDERS 56
Class Subcl Order

Notodelphyoida REC
Monstrilloida REC
Caligoida REC
Lernaeopodoida REC
Branchiura REC
Branchiura REC
Cirripedia (Thyrostraca, Eucirripedia) SIL-REC
Thoracica SIL-REC
Acrothoracica REC
Ascothoracica REC
Apoda REC
Rhizocephala REC
^^
Malacostraca PER-REC
Nebaliacea REC
Rhinocarina DEV-PEN
Ceratiocarina (Ceratocarina) CAM-PEN
Nahecarida DEV
Anaspidacea (Anomostraca, Syncarida) PEN-REC
®^
Mysidacea MIS-REC
Thermosbaenacea REC
Spelaeogriphacea REC
Lophogastridea REC
Cumacea (Sympoda) REC
Tanaidacea (Chelifera, Anisopoda) REC
Isopoda DEV-REC
Amphipoda (Laemodipoda) TER-REC
Euphausiacea REC
Pygocephalomorph a PAL
Decapoda ^^ TRI-REC
Stomatopoda (Hoplocarida) MIS-REC
PAUROPODA REC
Heterognatha REC

^^ = nebaliacea + rhinocarina + ceratioca-


Phyllocarida (Leptostraca)
rina HYMENOCARiNA. Eumalacostraca = anaspidacea + mysidacea +
+
THERMOSBAENACEA -\- CUMACEA -|- TANAIDACEA + ISOPODA + AMPHIPODA 4- EU-
PHAUSIACEA + DECAPODA -1- STOMATOPODA, Peracanda (Podophthalma, Thoraco-
straca, Schizopoda, Anaspides) = mysidacea + thermosbaenacea -|- lophogas-
tridea + cumacea + tanaidacea + isopoda + AMPHIPODA. Edriophthalma (Ar-
throstraca, Tetradecapoda) = isopoda -1- amphipoda. Eucarida = euphausia-
cea -I- decapoda + stomatopoda.
^^ Lophogastridea has sometimes been included here.
^^ Includes: Macrura, Anomura, Brachyura.
)

57 Arthropoda
Class Subcl Order

SYMPHYLA REC
Cephalostigmata REC
DIPLOPODA ^^ PEN-REC
Pselaphognatha {Penicillata) REC
Ancyrotricha REC
Lophotricha REC
^°°
Chilognatha TER-REC
Limacomorpha REC
Oniscomorpha (Armadillomorpha) TER-REC
Ascospermophora REC
Colobognatha ( Platydesmijormia TER-REC
Nematophora (Merochaeta) TER-REC
Proterospermophora REC
Opisthospermophora REC
Protosyngnatha PEN
Protosyngnatha PEN
Archipolypoda DEV-CAR
Palaeocoxopleura (Macrosterni) DEV-CAR
CHILOPODA (Syngnatha) TER-REC
Pleurostigmophora "^ (Pleurosdgma) TER-REC
Geophilomorpha (Geophylomorpha) TER-REC
Scolopendromorpha TER-REC
Lithobiomorpha TER-REC
Craterostigma REC
Notostigmophora (Notostigma) TER-REC
Scutigeromorpha TER-REC
INSECT A ^°2
(Hexapoda) DEV-REC
Apterygota (Synaptera, Apterygogenea, Aptera) DEV-REC
Protura {Myrientomata, Mirientomata, Frothy sanura, REC
Panprotura, Ellipura, Anamerentoma)
Thy sanura {Ectotrophi, Ectognatha) TRI-REC
Entotrophi (Entognatha, Diplura, Campodeoidea, Ho- mio-rec
lomerentoma, Panthy sanura, Dicellura)
^^ Includes: Eurysterna, Paleomorpha.
'^^°
Includes: Opisthandria, Proterandria, Eugnatha, Polydesmoidea, Juliformia.
Anamorpha = craterostigma
^°^ + lithobiomorpha. Epimorpha = geophil-
omorpha + scolopendromorpha.
^°^ extinct orders: Sypharopteroidea, Archodonata, Perielytrodea,
Includes
Protephemerida, Eubleptidodea, Syntonopteroidea, Permoneurodia. Pterygota
(Pterygogenea) = exopterygota + endopterygota. Palaeoptera = palaeo-
DICTYOPTERA + MEGASECOPTERA + PROTOHEMIPTERA + PROTODONATA + ODONATA
+ EPHEMERIDA. Neoptcra = EXOPTERYGOTA (eXCCpt PALAEOPTERA) + ENDOPTERY-
GOTA. Euentomata = diplura + ectotrophi + pterygota.
COMPLETE LIST OF CLASSES AND ORDERS 58
Class Subcl Order

CoUembola (Oligoentomata) dev-rec


Exopterygota ^"^
(Heterometabola) dev-rec
^°*
Palaeodictyoptera (Eopaleodictyoptera) pen-per
Megasecoptera ^°^
(Megasecopterida) pen-per
Protephemerida {Protephemeroidea) pen
"'^ per-rec
Ephemerida (Ephemeroptera, Ephemeroidea, Plec-
toptera)
Protodonata (Meganisoptera) pen-per
^°^
Odonata per-rec
Protohemiptera (Pseudohemiptera) pen-per
^°®
Protoperlaria per
Plecoptera (Perlarides, Perloidea, Perlaria, Nemurae- per-rec
des)
Protorthoptera pen-per
Caloneurodea pen-per
Diploglossata (Hemimeroidea, Dermodermaptera) pal
Grylloblattoidea (Notoptera) REC
Orthoptera (Saltatoria, Aeroplanoptera) pen-rec
Phasmidia (Phasmoidea, Ambulatoria, Gressoria, Che- tri-rec
leutoptera)
Blattaria (Blattaeformia, Oothecaria, Cursoria, Dicty- pen-rec
optera, Neoblattariae)
Mantodea {Deratoptera, Dacnostomata, Phylloptera, pal-rec
Exopterygoptera, Pandictyoptera )
Glosselytrodea per-jur
Protelytroptera (Protocoleoptera) per
Dermaptera (Labidura, Euplexoptera) jur-rec
Embioptera (Embiidina, Embioidea, Adenopoda, OH- oli-rec
goneura, Aetioptera)
Isoptera eoc-rec
^°^ Coleopteroidea =
protocoleoptera + coleoptera + strepsiptera.
= mallophaga + anoplura. Hemiptera (Rhynchota)
Phthiraptera (Ellipoptera)
= heteroptera + homoptera + protohemiptera + palaeohemiptera. Blat-
toidea = protoblattoidea + blattaria + mantodea + isoptera + zoraptera
+ psocoptera + mallophaga -1- anoplura.
^°* Includes: Hemiodonata, Anisaxia, Permodictyoptera, Breyeridea, Archae-
hymenoptera.
^^-'
Includes: Protohymenoptera, Diaphanopteroidea, Palaeohymenoptera.
^"^ Includes: Aphelophlebia, Agnatha, Odontota, Anisoptera, Archipterygota.
^°^ Includes: Paraneuroptera, Permodonata, Cryptodonata, Cryptodontia, Li-
bellulides, Libelluloidea.
^°^ Includes: Protoblattoidea, Pruvostitoptera, Mixotermitoidea, Synarmogoi-
59 Arthropoda — Chaetognatha
Class Subcl Order

Psocoptera (Corrodentia, Copeognatha) per-rec


Zoraptera (Panisoptera) rec
Mallophaga (Lipoptera) rec
Thysanoptera {Physopoda, Physapida, Thripoides, per-rec
Thripsites)
Homoptera pal-rec
Heteroptera "^ mes-rec
Anoplura ^^'^
(Siphunculata, Parasita, Pseudorhyn- ple-rec
chota)
"'^^
Endopterygota (Holometabola) per-rec
Neuroptera "^ per-rec
Mecoptera (Panorpatae, Panorpina, Mecaptera, Peta- per-rec
noptera, Paramecoptera, Protomecoptera, Proto-
diptera)
Trichoptera (Phryganoidea, Placipennes, Paratrichop- jur-rec
tera, Agnathes)
Lepidoptera (Glossata) eoc-rec
Diptera {Antliata, Halterata, Halteriptera, Haustel- jur-rec
lata)

Siphonaptera (Suctoria, Aphaniptera, Rophoteira, oli-rec


Medamoptera, Pulicina)
^^^
Coleoptera {Eleutherata, Elytroptera) per-rec
Strepsiptera ^" (Rhipiptera, Rhipidoptera, Strepsata, oli-rec
Stylopida)
Hymenoptera jur-rec

CHAETOGNATHA ^^=

CHAETOGNATHA cam-rec
Sagittoidea cam-rec

dea, Hapalopteroidea, Hadentomoidea, Reculoidea, Cnemidolestoidea, Paraple-


coptera, Miomoptera, Protocicadida, Protofulgorida.
^°^ Includes: Palaeohemiptera, Hemipsocoptera.
^^°
Includes: Pediculidea, Polyptera.
^^^ = archaeohymenoptera -f- palaeohymenoptera +
Hymenopteroidea
PROTOHYMENOPTERA HYMENOPTERA. Panorpoidea = TRICHOPTERA + LEPIDOP-
+
TERA + DIPTERA + SIPHONAPTERA + MEGASECOPTERA + PANORPATAE.
Includes: Megaloptera, Raphidioidea, Emmenognatha, Leptophya. (Plani-
'^'^'^

pennia sometimes used for all these.)


"^ Includes: Paracoleoptera.
^^* Sometimes placed in Coleoptera as family Stylopidae.
^^^
Synonyms: Homalopterygia, Oesthelminthes.
^^® Synonyms: Brachiata, Pogonofora.
COMPLETE LIST OF CLASSES AND ORDERS 60
Class Subcl Order

POGONOPHORA^^«
POGONOPHORA REC
Thecanephria REC
Athecanephria REC

ECHINODERMATA
CYSTOIDEA ORD-PER
Hydrophoridea "^ (Cystidea) ord-dev
Rhombifera ord-dev
Diploporita ord-dev
^"
Blastoidea ord-per
Eublastoidea sil-per
Coronata ord-sil
Parablastoidea ord
CRINOIDEA ^^° {Brachiata, Actinoidea, Eucrinoidea) ord-rec
^^^
Inadunata ord-tri
Disparida (Disparata) ord-per
Hybocrinida ord-sil
Cladida (Cladoidea) ord-tri
Flexibilia (Ichthyocrinacea) ord-per
Taxocrinida (Taxocrinoidea) ord-per
Sagenocrinida (Sagenocrinoidea) sil-per
Camerata (Sphaeroidocrinacea, Adunata) ord-per
Diplobathrida (Diplobathra) ord-mis
Monobathrida (Monobathra) ord-per
^^^
Articulata tri-rec
Isocrinida tri-rec
Millericrinida tri-eoc
Cyrtocrinida jur-rec
Uintacrinida cre
Roveacrinida tri-cre
^^^ = cystoidea + blastoidea + crinoidea + eocri-
Subphyla: Pelmatozoa
NOIDEA PARACRINOIDEA + EDRIOASTEROIDEA. HomaloZOa = CARPOIDEA + MA-
+
chaeridia. Haplozoa = single class haplozoa. Eleutherozoa = somasteroidea +
ASTEROIDEA + OPHIUROIDEA + ECHINOIDEA + BOTHROCIDAROroEA + OPHIOCYSTO-
IDEA -|- HOLOTHURioiDEA. Includes: Heterostelea (carpoidea + Amphoridea), As-
terozoa, Stelleroidea, Stelliformia, Hypostoma, Echinozoa, Cystocidaroidea, Apo-
rita.
^^^Includes: Eucystoidea, Cystechinoidea, Dichoporita, Cystocrlnoidea.
"^ Sometimes separated into orders Regulares and Irregulares, or orders Eubla-
stoidea and Protoblastoidea. Sometimes as a separate class.
^^° Includes: Palaeocrinoidea, Inarticulata, Coadunata, Costata, Testacea, Mo-
6i Pogonophora — Echinodermata
Class Subcl Order

Comatulida jur-rec
'^
EOCRINOIDEA cam-ord
Eocrinoidea cam-ord
PARACRINOIDEA ord
Paracrinoidea ord
EDRIOASTEROIDEA (Thecoidea, Cystasteroidea, Thyroidea, cam-pen
A gelacrinoidea )
Edrioasteroidea cam-pen
CARPOIDEA CAM-DEV
Carpoidea cam-dev
MACHAERIDIA ^'^ ord-dev
Machaeridia ord-dev
^^
HAPLOZOA CAM
Cyamoidea cam
Cycloidea cam
SOMASTEROIDEA ord
Goniactinida ord
ASTEROIDEA '"' (Cirrigrada) ord-rec
Hemizonida ord-car
Platyasterida ord-dev
Phanerozonea ord-rec
^^®
Spinulosa rec
^^°
Forcipulata rec
AULUROIDEA ord-mis
Lysophiuroida (Lysophiurae, Aegophiurida) ord-mis
Streptophiuroida (Streptophiurae) ord-mis
OPHIUROIDEA ^''
(Spinigrada) ord-rec
Myophiurida ord-dev
Ophiocystiida ord-dev
Aganasterida ?-rec
Phrynophiurida ?-rec
Laemophiurida ?-rec
Gnathophiurida ?-rec
nocyclica, Dicyclica, Irregularia, Regularia, Larvata, Tesselata, Canaliculata,
Semi-articulata.
^^^
Sometimes divided into Larviformia (Haplocrinacea) and Fistulata (Cyatho-
crinacea).
^^^ Includes: Pentacrinoidea, Pentacrinacea, Stomatocrinoidea, Neocrinoidea.
123
Perhaps not a natural group.
^2*

12°
Groups of doubtful relationships, — possibly not even Echinodermata.
Includes: Encrinasteriae, Euasteriae, Cryptozonia.
126
Sometimes united as Cryptozonia.
1" Includes: Ophiureae, Ophiurida, Euryalae, Euryalida, Cladophiurae, Zyg-
ophiurae. Also sometimes divided into orders Stenurida and Ophiurida.
COMPLETE LIST OF CLASSES AND
63 Pterobranchia — Cephalochordata
Class Subcl Order

^^^
E N T E R P N E U STA

ENTEROPNEUSTA rec
Balanoglossida rec

PLANCTOSPHAEROIDEA
PLANCTOSPHAEROIDEA rec
Planctosphaeroidea rec

^^*
T U N I CATA

LARVACEA (Copelata, Appendicularia, Atremata, Perennichor- rec


data)
Larvacea rec
ASCmiACEK^^^ {Tethyodeae) rec
Stolidobranchiata (Ptychobranchia) REC
Aspiraculata REC
Phlebobranchiata (Diktyobranchia, Dictyobranchia) rec
Aplousobranchiata (Krikobranchia) rec
Octacnemida rec
THALIACEA rec
Pyrosomata Rec
Pyrosomatida (Luciae, Lucida) rec
Myosomata rec
Cyclomyaria (Doliolida) rec
Hemimyaria (Salpida) rec
Desmomyaria rec

CEPHALOCHORDATA ^*«

LEPTOCARDIA rec
Amphioxi (Branchiostomoidea) rec

^^^ Includes: Pelagothurida. "^ Synonyms: Helminthomorpha.


Synonyms: Urochordata. Suhphyla: Copelata
^^* = larvacea; Acopa (Caduci-
chordata) = ascidiacea + thaliacea.
^^^ Includes: Enterogona, Pleurogona.
^^^ Synonyms: Cirrhostomi, Entomocrania, Haplocyemata, Homomeria, Mye-
lozoa, Pharyngobranchii, Acrania.
COMPLETE LIST OF CLASSES AND ORDERS 64
Class Subcl Order

VERTEBRATA 137

AGNATHA "® (Monorhyncha, Monorhina) ord-rec


Cephalaspidomorpha sil-rec
^^^
Cyclostomata (Marsipobranchii, Petromyzonoidea) rec
Cephalaspida "° {Osteostraci, Aspidocephali, Osteo- sil-dev
stracoidea, Cephalaspidiformes )
Anaspida "^ (Anaspidoidea) sil-dev
^*^
Pteraspidomorpha (Pteraspides) ord-dev
Pteraspida (Heterostraci, Pteraspidijormes ord-dev
Coelolepida (Coelolepidoidea, Coelolepiformes) sil-dev
PLACODERMI ^'^
(Aphetohyoidea) sil-per
Acanthodii "* (Acanthoglossa, Acanthodioidea) sil-per
^^^ sil-dev
Arthrodira {Euarthrodira, Arthrodiroidea, Phyllo-
lepida, Ptyctodontida)

Macropetalichthyida "® (Anarthrodira) DEV


Antiarchi ^^^
(Antiarchoidea, Pterichthyes, Pterichthy- dev
omorphd)
^^^
Synonyms: Euchorda, Craniata. Superclass Gnathostomata (Amphirhyncha,
Amphirhina) = Placodermi + all "higher" vertebrates. Ichthyopterygii = placo-
DERMI + ELASMOBRANCHII + HOLOCEPHALI + OSTEICHTHYES. PisceS (Ichthya,
Eotetrapoda) = agnatha + placodermi + chondrichthyes + osteichthyes.
Haemacryma = pisces + amphibia + reptilia. Haematherma = ayes + mam-
malia. Anamnia (Ichthyopsida) = pisces + amphibia. Amniota = reptilia +
aves + MAMMALIA. Tetrapoda (Cheiropterygia) = amphibia + reptilia + ayes
+ MAMMALIA. Sauropsida = reptilia + ayes. Includes: Protichthyes, Ichthyo-
dorulites.
^^^Ostracodermi (Ostracophori) = cephalaspida + anaspida + pteraspida +
Coelolepida. Includes: Euphaneroidea.
^^^ Includes: Myxinoidea, Hyperotreti, Myxini, Myxiniformes, Petromyzontia,

Petromyzontiformes, Hyperoarti.
"" Includes: Tremataspidiformes, Lasaniiformes, Oligocnemata, Endeiolepi-
formes.
^" Includes: Birkeniiformes, Phlebolepiformes.
^*^ Includes: Astraspiformes, Psammosieiformes, Cyathaspiformes, Amphiaspi-
formes, Thelodonti.
"^ Coccostei (Coccosteomorphi) = euarthrodira + phyllolepida + mac-
ropetalichthyes.
^^^ Includes: Climatiiformes, Mesacanthiformes, Ischnacanthiformes, Gyra-
canthiformes, Cheiracanthiformes, Acanthodiformes, Acanthoessi, Acanthoessi-
formes. Conodontophorida (Conodonta) may belong here (Ord-Tri).
^^^ Includes: Acanthothoraci, Arctolepiformes, Acanthaspidomorphi, Acanth-
aspida, Coccosteiformes, Brachythoraci, Mylostomatiformes, Ptyctodontiformes,
Phyllolepiformes.
"" Includes: Gemuendiniformes, Rhenanida, Petalichthyida.
^^^ Includes: Remigolepiformes, Asterolepiformes.
6$ Vertebrata
Class Subcl Order

Stegoselachii (Stensioelliformes, Jagoriniformes, Sten- sil-mis


sioellida)

Palaeospondyloidea (Palaeospondyliformes) dev


CHONDRICHTHYES dev-rec
Elasmobranchii "^ dev-rec
Cladoselachii "^ (Pleuropterygii) dev-per
Pleuracanthodii {Ichthyotomi, Proselachii) dev-tri
Selachii ^^°
(Euselachii, Plagiostomi, Chondropterygii, dev-rec
Placoidei)
Batoidea "^ (Hypotremata, Platosomia) jur-rec
''^
Holocephali dev-rec
Bradyodonti ^^^
(Bradyodontoidea) dev-per
Chimaerae ^^*
(Chimaeroidea) jur-rec
OSTEICHTHYES (Teleostomi) dev-rec
'^^
Actinopterygii dev-rec
Chondrostei ^^"^
(Chondrosteoidea, Palaeopterygii) dev-rec
"^ PER-REC
Holostei
'^^
Teleostei jur-rec
'^'^^
Includes: Diplodonti, Diplodontiformes, Xenanthi, Xenacanthi, Xenacan-
thoidea.
'^^^
Includes: Cladodontiformes, Cladoselachoidea, Cladoselachiformes.
^^° Pleurotremata, Selachoidea, Heterodontiformes,
Includes : Pleurostomata,
Heterodontoidea, Hexanchoidea, Lamnoidea,
Hexanchiformes, Notidanoidea,
Lamniformes, Isuriformes, Galeoidei, Squaloidea, Squaliformes, Tectospondyli,
Edestidi.
^^^
Includes: Narcaciontiformes, Torpediniformes, Rajiformes.
^^^ Includes: Petalodontes, Petalodontiformes.
^^^
Includes: Eubradyodonti, Chondrenchelyes, Chondrenchelyiformes.
^^* Includes: Chimaeriformes.
^^^
Sometimes divided into superorders: chondrostei, holostei, teleostei.
Sometimes divided into infraclasses: Polypterei, chondrostei, holostei,
TELEOSTEI. Ganoidi = chondrostei + holostei. Neopterygii = holostei +
TELEOSTEI.
^^•^
Includes: Tarrasiiformes, Gymnonisciformes, Luganoidiiformes, Phanero-
rhynchiformes, Dorypteriformes, Cephaloxeniformes, Bobasatraniiformes, Red-
fieldiiformes,Calopteriformes, Perleidiformes, Platysiagiformes, Ospiiformes,
Aetheodontiformes, Pholidopleuriformes, Saurichthyiformes, Palaeoniscoidea, He-
terocerci, Palaeonisciformes, Polypterini, Brachyopterygii, Polypteriformes, Cla-
distia, Cladistioidea, Acipenseroidea, Acipenseriformes, Subholostei, Subholo-
steoidea.
^" Includes:
Semionotoidea, Lepidostei, Lepisostei, Lepidosteoidea, Lepiso-
steiformes,Ginglymodi, Rhomboganoidei, Pycnodontoidea, Pycnodontiformes,
Aspidorhynchoidea, Aspidorhynchiformes, Amioidea, Amiiformes, Pholidopho-
roidea, Pholidophoriformes, Pachycormoidea, Pachycormiformes, Protospon-
dyli.
^''s
See p. 66 for footnote 158.
COMPLETE LIST OF CLASSES AND ORDERS 66
Class Subcl Order

Choanichthyes (Amphibioidea) dev-rec


^^^
Crossopterygii (Crossopterygioidea) dev-rec
^®°
Dipnoi DEV-REC
AMPHIBIA '^' (Batrachia) dev-rec
^^^
Labyrinthodontia (Stegocephalia) dev-tri
Ichthyostegalia dev-pen
Rhachitomi mis-tri
Embolomeri mis-per
Trematosauria mes
Stereospondyli tri
^^^
Salientia pen-rec
Eoanura pen
Proanura tri
Anura "^ {Euanura, Ecaudata) jur-rec
Lepospondyli (Urodeloidei, Pseudocentrophori) mis-rec
Aistopoda pen
Nectridia pen-per
Microsauria {Adelospondyli, Micramphibia) mis-per
Urodela {Caudata, Gradientia, Saurabatrachia) cre-rec
Apoda (Gymnophiona, Peromela, Caecilia) REG
^^^ Includes: Isospondyli, Isospondyloidea, Malacopterygii, Thrissomorphi,
Ostariophysi, Ostariophysoidea, Apodes, Heteromi, Heteromoidea, Mesichthyes,
Cyprinodontiformes, Acanthopterygii, Symbranchiformes, Physostomi, Physocly-
sti, Discocephalioidea, Echiniiformes, Echeniiformes, Plectognathoidea, Hyposto-

mosoidea, Cephalacanthoidea, Scleropareioidea, Percomorphoidea, Allotriogna-


thoidea, Anacanthoidea, Apodoidea, Batrachoidea, Batrachoidiformes, Haplodoci,
Gobiesociformes, Xenopterygii, Lampridiformes, Lophiiformes, Lyomeroidea,
Pediculati, Pediculatiformes, Pegasiformes, Salmopercoidea, Solenichthyoidea,
Synentognathoidea, Tetradontiformes, Tetrodontiformes, Thoracostoidea, Thun-
niformes, Icosteioidea, Icosteiformes, Malacichthyes, Chaudhurioidea, Chaudhuri-
iformes, Mastacembelioidea, Mastacembeliformes, Opisthomi, Synbranchioidea,
Perciformes, Dactylopteriformes, Pleuronectoidea, Pleuronectiformes, Heteroso-
mata, Beryciformes, Berycomorphoidea, Zeiformes, Zeomorphoidea, Mugilifor-
mes, Polynemiformes, Ophiocephaliformes, Clupeiformes, Bathyclupeoidea, Ba-
thyclupeiformes, Galaxiiformes, Cypriniformes, Anguilliformes, Halosauriformes,
Notacanthiformes, Beloniformes, Myctophiformes, Iniomi, Scopeliformes, Ateleo-
piformes, Ateleopoidea, Giganturoidea, Giganturiformes, Saccopharyngiformes,
Mormyroidea, Mormyriformes, Gadiformes, Macruriformes, Gasterosteiformes,
Syngnathiformes, Cyprinodontoidea, Microcyprini, Phallostethiformes, Percopsei-
formes, Stephanoberyciformes.
^^^ Includes: Porolepiformes, Osteolepiformes, Osteolepides, Osteolepidoti, Ho-
loptychiformes, Megalichthyiformes, Rhipidistia, Actinistla, Coelacanthini, Coela-
canthiformes.
^^° Includes: Dipteri, Dipteriformes, Dipneumona, Dipneusti, Phaneropleuri-
67
COMPLETE LIST OF CLASSES AND ORDERS 68
Class Subcl Order
^^*
Saurischia (Theropoda, Sauropoda, Compsognatha) tri-cre
Omithischia (Orthopoda, Ornithopoda, Predentata tri-cre
Stegosauria)
^^^
Synapsida pen-jur
Pelycosauria pen-tri
^^^
Therapsida per-tri
Ictidosauria tri-jur
Mesosauria (Proganosauria) per
AVES jur-rec
Archaeornithes (Saururae) jur
Archaeopteryges (Archaeopterygiformes) jur
^^^
Neoraithes (Ornithae, Ornithurae) cre-rec
Hesperomithes (Odontolcae, Hesperornithijormes) cre-eoc
Ichthyornithes (Odontormae, Ichthyornes, Ichthyor- cre
nithijormes, Ichthyornijormes )
Sphenisci (Sphenisciformes, Impennes) oli-rec
Caenagnathae (Coenagnathiformes) cre
Struthiones {Struthioniformes) pli-rec
Rheae (Rheiformes) pli-rec
Casuarii (Casuariiformes) ple-rec
Dinornithes (Dinornithiformes) ple
Aepyornithes (Aepyornithiformes) eoc-ple
Apteryges (Apterygiformes) ple-rec
Crypturi (Tinami, Tinamijormes, Crypturiformes) pli-rec
Gaviae (Gaviiformes) eoc-rec
Podicipedes (Podicipediformes, Podicipiformes, Colym- oli-rec
bae, Colymbiformes, Pygopodes)
Procellariae {Procellariijormes, Cecomorphae, Turbi- eoc-rec
nares, Tubinares)

^^* Includes: Hallopoda.


'^''^
Promammalia (Protodonta),
Includes: at one time placed in subclass Proto-
theria, of Mammalia.
^''^
Includes: Dromosauria, Deinocephalia, Dicynodontia, Theriodontia, Ano-
modontia, Theromorpha.
^" Includes: Dromeognathae, Megistanes, Gastornithes, Stereornithes, Cari-
natae, Colymbomorphae, Pelargomorphae, Alectoromorphae, Coraciomorphae,
Limicolae, Pterocletes, Picariae, Musophagi. Neognathae (Euornithes) = all ex-
cept HESPERORNiTHEs \- ICHTHYORNITHES + IMPENNES. Superorder Odonto-

gnathae =
hesperornithes (+ ichthyornithes). Superorder Palaeognathae =
caenagnathae + struthiones 4- rheae -1- casuarii + dinornithes 4- aepyor-
nithes -I- apteryges. Ratitae included struthiones, rheae, casuarii, dinor-
69 Vertebrata
Class Subcl Order

Steganopodes (Pelecani, Pelecaniformes, Pelicanifor- cre-rec


mes)
^^^
Ciconiae (Ciconiiformes, Gressores, Herodiones cre-rec
Herodii)
Anseres {Anseriformes, Chenomorphae) cre-rec
Falcones "® (Falconiformes, Accipitres, Accipitrijor- eoc-rec
mes)
^^°
Galli (Galliformes, Gallinae) eoc-rec
^^^
Grues (Gruiformes) eoc-rec
Diatrymae (Diatrymiformes) Eoc
^*"
Charadriae (Charadriiformes, Laro-limicolae) eoc-rec
Columbae (Columbiformes) mio-rec
Psittaci (Psittaciformes) mio-rec
Cuculi (Coccyges, Cuculiformes) oli-rec
Striges (Strigiformes) eoc-rec
Caprimulgi (Caprimulgiformes) pli-rec
Macrochires (Macrochiriformes, Apoda, Apodijormes, oli-rec
Micropodi, Micropodiformes)
Colli (Colliformes) rec
Trogones (Trogoniformes) oli-rec
^^^
Coraciae (Coraciiformes) eoc-rec
Pici (Piciformes) eoc-rec
^^*
Passeres (Passeriformes) eoc-rec
MAMMALIA ^^^
(Mammifera) jur-rec
Prototheria ple-rec
Monotremata (Omithodelphia) ple-rec
^^^
Allotheria jur-eoc
Multituberculata jur-eoc
Triconodonta jur

NITHES, AEPYORNITHES, APTERYGES, CURSORES, PROCERES, BREVIPENNES, CRYPTURI.


^^®
Includes: Phoenicopteri, Phoenicopteriformes, Amphimorphae, Pelargi-
formes.
^^^ Includes: Cathartidiformes.
^^^ Includes: Opisthocomi, Opisthocomiformes, Heteromorphae, Grallae.
^^^ Includes: Ralliformes, Heliornithiformes, Hemipodii, Telmatomorphormes.
^^^ Includes: Alciformes, Lariformes.
^^^ Includes: Alcedines, Bucerotes.
^^* Includes: Eurylaemi, Menurae. Scansores = pici + psittaci in part.
^^^
Epiacentalia =
prototheria + allotheria + pantotheria. Theria = pan-
TOTHERiA + metatheria + eutheria. Promammalia transferred to Reptilia (Sy-
naptida).
^^^ Includes: Plagiaulacoidea, Tritylodontoidea.
COMPLETE LIST OF CLASSES AND ORDERS 70
Class Subcl Order

Pantotheria jur
Trituberculata jur
Symmetrodonta jur
^^'^
Metatheria cre-rec
Marsupialia (Didelphia) cre-rec
^^^
Eutheria (Monodelphia, Placentalia, Placentaria) cre-rec
Insectivora cre-rec
Dermoptera eoc-rec
Chiroptera eoc-rec
^^^
Primates eoc-rec
Tillodontia eoc
Taeniodonta (Ganodonta, Stylinodontia) eoc
Edentata {Xenarthra, Paratheria, Bruta) eoc-rec
Pholidota (Squamata, Nomarthrd) oli-rec
Lagomorpha (Duplicidentata) eoc-rec
Rodentia^®*' eoc-rec
^^^
Cetacea (Mutica, Cete) eoc-rec
^^^
Carnivora eoc-rec
Condylarthra eoc
Litoptema eoc-ple

^^'' Includes: Polyprotodontia, Caenolestoidea, Diprotodontia, Paucitubercu-


lata.
^^^ Includes: Ancylopoda, Chalicotheria, Taxeopoda, Quadrumana, Daubento-
mioidea, Subungulata, Pithecoidea, Platyrhina, Simiae, Toxodontia, Barytheria,
Carnivores, Edentates, Ungulates, Bunotheria. Unguiculata = insectivora +
DERMOPTERA CHIROPTERA + PRIMATES + TILLODONTIA + TAENIODONTA + EDEN-
4-

TATA -|- PHOLIDOTA. Ungulata = Protungulata (condylarthra -\- litopterna +


NOTOUNGULATA + ASTRAPOTHERIA + TUBULIDENTATA) + Paenungulata (PANTO-
DONTA + DINOCERATA -\- xenungulata + pyrotheria + proboscidea + embri-
THOPODA + HYRACOIDEA + DESMOSTYLIFORMES + SIRENIA -1- PERISSODACTYLA -f-
artiodactyla). Glires = lagomorpha + rodentia. Mutilata = cetacea + sire-
NiA. Protungulata = condylarthra + litoptera + notoungulata + astrapo-
THERIA + TUBULIDENTATA. Therictoidea = insectivora + Ferae. Archonta =
Menotyphla + dermoptera -\- chiroptera + primates. Paenungulata = panto-
DONTA + DINOCERATA -1- PYROTHERIA + PROBOSCIDEA + EMBRITHOPODA + HYRA-
COIDEA 4- SIRENIA. Diplarthra = perissodactyla + artiodactyla.
^^^ Includes: Anthropoidea, Lemuroidea, Prosimii.

^^° Includes: Simplicidentata.


^^^ Includes: Zeuglodontia, Archaeoceti, Odontoceti, Mystacoceti.
^^2 Includes: Fissipedia, Creodonta, Pinnipedia, Ferungulata, Ferae.
^^^ Includes: Taxodontia.
^®* Includes: Amblydactyla, Taligrada, Coryphodontia.
^^^ Includes: Myohyracoidea.
yi Vertebrata
^^^
Notoungulata eoc-ple
Astrapotheria eoc-mio
Tubulidentata eoc-rec
^^*
Pantodonta (Amblypoda) eoc-oli
Dinocerata (Uintatheria) eoc
Pyrotheria eoc-oli
Proboscidea eoc-rec
Embrithopoda (Barypoda) oli
^^^
Hyracoidea (Hyraces) oli-rec
Sirenia eoc-rec
Perissodactyla {Mesaxonia) eoc-rec
Artiodactyla (Paraxonia) eoc-rec
BIBLIOGRAPHY

MANY BOOKS, pamphlets, and articles have been consulted in the preparation of
this classification. A listing of all of these appears to serve no essential purpose
here. The Ust given below therefore contains principally recent works in which a
formal classification is presented for the entire animal kingdom or a major part of

it. The list is intended merely as background for the present classification. It does
not contain references to all authors and works cited in the text.

Boettger, C. R.
1952. Die Stamme des Tierreichs in ihrer systematischen Gliederung. Abhandl.
Braunschweigischen Wiss. Gesell., Band iv, pp. 238-300.
Borradaile, L. A., Potts, F. A., Eastham, L. E. S., and Saunders, J. T.
1951. The Invertebrata. A manual for the use of students, 2nd edition, 725 pp.
Cambridge Univ. Press, Cambridge.
Brues, C. T., Melander, A. L., and Carpenter, F. M.
1954. Classification of insects. Museum of Comparative Zoology, Bull. 108,

pp. 1-917.
Colbert, E. H.
1955. Evolution of the vertebrates. A history of the backboned animals
through time, 479 pp. John Wiley & Sons, New York.
Dawes, B.
1956. The Trematoda, with special reference to British and other European
forms, 644 pp. Cambridge Univ. Press, Cambridge.
Forster-Cooper, C.
1951. A text -book of zoology, by T. J. Parker and W. A. Haswell, 6th edi-
tion, vol. 2, 758 pp. Macmillan & Co., London.
Hadzi, J.

1953. An attempt to reconstruct animal classification. Systematic Zoology, vol.


2, pp. 145-154.
Hall, R. P.
1953. Protozoology, 682 pp. Prentice-Hall, New York.
Hyman, L. H.
1940. The invertebrates: Protozoa through Ctenophora, 726 pp. McGraw-Hill
Book Co., New York.
1951. The invertebrates: Platyhelminthes and Rhynchocoela. The acoelomate
Bilateria, vol. n, 550 pp. McGraw-Hill Book Co., New York.

1951. The invertebrates: Acanthocephala, Aschelminthes, and Entoprocta. The


Bibliography
y3
pseudocoelomate Bilateria, vol. in, 572 pp. McGraw-Hill Book Co.,
New York.
1955. The invertebrates: Echinodermata. The Coelomate Bilateria, vol. iv,

763 pp. McGraw-Hill Book Co., New York.


1959. The invertebrates: Smaller coelomate groups. Chaetognatha, Hemichor-
data, Pogonophora, Phoronida, Ectoprocta, Brachiopoda, Sipunculida.
The coelomate Bilateria, vol. v, 783 pp. McGraw-Hill Book Co., New
York.
Kerkut, G. A.
1958. The Invertebrata, by Borradaile, Potts, et al., 3rd edition, 795 pp. Cam-
bridge Univ. Press, Cambridge.
Kudo, R. R.
1954. Protozoology, 4th edition, 966 pp. Charles C. Thomas, Springfield,
Illinois.

Lowenstein, O.
1954. A text-book of zoology, by T. J. Parker and W. A. Haswell, 6th edi-
tion, vol. 1, 770 pp. Macmillan & Co., London.
Mayr, E. and Amadon, D.
1951. A classification of recent birds. American Museum Novitates, No. 1496,
pp. 1^2.
Mayr, E., Linsley, E. G., and Usinger, R. L.
1953. Methods and principles of systematic zoology, 328 pp. McGraw-Hill
Book Co., New York.
Moore, R. C. (editor)
1953-1961. Treatise of invertebrate paleontology.
(Issued in parts with numerous contributors)
D. Protista 3. Protozoa (chiefly Radiolaria and Tintinnina), 195 pp. 1954. (By
A. S. Campbell and R. C. Moore)
E. Archaeocyatha and Porifera, 122 pp. 1955. (By V. J. Okulitch and M. W.
de Laubenfels)
P. Coelenterata, 498 pp. 1956. (By F. M. Bayer, H. Boschma, H. J. Harring-
ton, D. Hill, L.H. Hyman, M. Lecompte, E. Montanaro-Gallitelli, R. C.
Moore, E. C. Stumm, J. W. Wells)
G. Bryozoa, 253 pp. 1953. (By R. S. Bassler)
I. Mollusca 1. Mollusca —
General features. Scaphopoda. Amphineura. Mo-
noplacophora. Gastropoda —General features. Archaeogastropoda and
some (mainly Paleozoic) Caenogastropoda and Opisthobranchia, 351 pp.
1960. (By J. B. Knight, L. R. Cox, A. M. Keen, A. G. Smith, R. L.
Batten, E. L. Yochelson, N. H. Ludbrook, R. Robertson, C. M. Yonge,
R. C. Moore)
L. Mollusca IV. Cephalopoda, Ammonoidea, 490 pp. 1957. (By W. J. Arkell,
W. M. Furnish, B. Kummel, A. K. Miller, R. C. Moore, O. H. Schinde-
wolf, P. C. Sylvester-Bradley, C. W. Wright)
O. Arthropoda 1. Arthropoda —
General features. Protarthropoda. Euarthrop-
oda —
General features. Trilobitomorpha, 560 pp. 1959. (By H. J. Har-
rington, G. Henningsmoen, B. F. Howell, V. Jaanusson, C. Lochman-
BIBLIOGRAPHY 74
Balk, R. C. Moore, C. Paulsen, F. Rasetti, E. Richter, R. Richter,
H. Schmidt, K. Sdzuy, W. Struve, L. St0rmer, C. J. Stubblefield,
R. Tripp, J. M. Weller, H. B. Whittington)
P. Arthropoda 2. Chelicerata with sections on Pycnogonida and Palaeoisopus,
181 pp. 1955. (By L. St0rmer, A. Petrunkevitch, J. W. Hedgpeth)
Q. Arthropoda 3. Crustacea, Ostracoda, 442 pp. 1961. (By R. H. Benson, J. M.

Berdan, W. A. van den Bold, T. Hanai, I. Hessland, H. V. Howe, R. V.


Kesling, S. A. Levinson, R. A. Reyment, R. C. Moore, H. W. Scott,

R. H. Shaver, I. G. Sohn, L. E. Stover, F. M. Swain, P. C. Sylvester-


Bradley, J. Wainwright)
V. Graptolithina with sections on Enteropneusta and Pterobranchia, 101 pp.
1955. (By O. M. B. Bulman)
Moore, R. C, Lalicker, C. G., and Fischer, A. G.
1952. Invertebrate fossils, 766 pp. McGraw-Hill Book Co., New York.
Noble, G. K.
1931. The biology of the Amphibia, 577 pp. McGraw-Hill Book Co., New
York. (Reprinted 1954, Dover Publications, New York)
Parker, T. J. and Haswell, W. A.
1897. A text-book of zoology, vol. i, 779 pp. Macmillan & Co., London.
Pearse, A. S.
1949. Zoological names. A list of phyla, classes, and orders, 24 pp. Zoology
Department, Duke Univ., Durham, N. C.
Piveteau, J.

1953. Traite de Paleontologie, tome iii, 1063 pp. Masson et Cie., Paris.

(Onychophores, Arthropodes, fichinodermes, Stomocordes)


Romer, A. S.

1945. Vertebrate paleontology, 2nd edition, 687 pp. Univ. of Chicago Press,
Chicago.
Rothschild, (Lord).
1961. A classification of living animals, 106 pp. Longmans, Green & Co., Ltd.
London.
Shrock, R. R. and Twenhofel, W. H.
1953. Principles of invertebrate paleontology, 816 pp. McGraw-Hill Book
Co., New York.
Simpson, G. G.
1945. The
principles of classification and a classification of mammals. Bull.
American Museum Natural History, vol. 85, 350 pp.
Wardle, R. A. and McLeod, J. A.
1952. The zoology of tapeworms, 780 pp. Univ. of Minnesota Press, Min-
neapolis.
Waterman, T. H. and Chace, F. A., Jr.
1960. General crustacean biology, pp. 1-33, in The physiology of Crustacea,
edited by T. H. Waterman, vol. 1 , 670 pp. Academic Press, New York.
Wetmore, A.
1960. A classification for the birds of the World. Smithsonian Miscellaneous
Collections, vol. 139, No. 1 1, pp. 1-37,
8 1

INDEX TO COMMON NAMES

aardvarks, 25 beavers, 25 bugs (Continued)


acorn-worms, 22 bed-bugs, 20 sowbugs, 19
albatrosses, 24 bee-eaters, 24 spittle-bugs, 20
alligators, 23 bees, 21 bulbuls, 25
alpacas, 25 beetles, 20 buntings, 25
amphibians, 23 bird-lice, 20 bush-babies, 25
angel-fishes, 23 birds, 23 bustard-quails, 24
angle-worms, 17 antbirds, 25 bustards, 24
animalcules blackbirds, 25 butcher-birds, 25
bear-animalcules, 7, 17 bowerbirds, 25 butterflies, 20
sun-animalcules, 9 butcher-birds, 25 butterfly-shells, 16
wheel-animalcules, 6, frigate-birds, 24
13 humming-birds, 24
animals mockingbirds, 25 Caddis-flies,20
one-celled animals, 6 mouse-birds, 24 caecilians,23
anis, 24 oil-birds, 24 cake-urchins, 22
annelids, 16 ovenbirds, 25 calcareous sponges, 10
antbirds, 25 puflFbirds, 24 camels, 25
anteaters, 25 secretary-birds, 24 campodeids, 20
antelopes, 25 snakebirds, 24 capybaras, 25
ant-lions, 20 songbirds, 25 caracaras, 24
ants, 21 surf-birds, 24 cartilaginous fishes, 23
velvet-ants, 21 tropicbirds, 24 case-flies, 20
white-ants, 20 wattlebirds, 25 cassowaries, 24
apes, 25 weaverbirds, 25 caterpillars, 20
aphids, 20 birds-of-paradise, 25 cats, 25
arachnids, 7, 18 biting-lice, 20 cattle, 25
argonauts, 16 bitterns, 24 caymans, 23
armadillos, 25 bivalves, 16 centipedes, 19
arrow-worms, 7, 21 blackbirds, 25 cephalopods, 16
arthropods, 17 black-corals, 12 chalky sponges, 10
ascidians, 22 black-widows, 18 chiggers, 18
avocets, 24 bloodsuckers, 17 chigoes, 20
awks, 24 blue-corals, 11 chimaeras, 23
aye-aye, 25 blues, 20 chimpanzees, 25
boat-shells, 16 chinchillas, 25
bony fishes, 23 chipmunks, 25
boobies, 24 chitons, 15
babblers, 25 book-lice, 20 cicadas, 20
baboons, 25 book-scorpions, 18 ciliates, 10
badgers, 25 bots, 20 civets, 25
bandicoots, 25 bowerbirds, 25 clams, 16
barbets, 24 brachiopods, 15 coatis, 25
bark-lice, 20 brine-shrimps, 1 coccolithophores, 9
barnacles, 18 bristle-tails, 20 coccoliths, 9
basket-stars, 21 brittle-stars, 21 cockroaches, 20
batrachians, 23 broadbills, 25 coelacanths, 23
bats, 25 bryozoans, 15 coelenterates, 1
bear-animalcules, 7, 17 bugs, 20 colies, 24
beard-worms, 7, 21 bed-bugs, 20 colugos, 25
bears, 25 croton-bugs, 20 comb-jellies, 6, 12
water-bears, 7, 17 mealy-bugs, 20 coots, 24
woolly-bears, 20 pillbugs,19 corallines, 15
11 1

INDEX TO Common names 76


corals, 6, 11 fairy -shrimps, 18 frogs, 23
black-corals, 12 falcons, 24 frog-spit, 20
blue-corals, 11 false-scorpions, 18 fulmars, 24
hexacorals, 11 feather-stars, 21
horny-corals, 11 finches, 25
soft-corals, 11 fireflies, 20 gallinules, 24
stony-corals, 11 fishes gall-wasps, 21
thorny-corals, 12 angel-fishes, 23 gannets, 24
cormorants, 24 bony fishes, 23 gars, 23
crabs, 19 cartilaginous fishes, 23 gastropods, 15
horseshoe-crabs, 17 crayfish, 19 gastrotrichs, 13
craniates, 35 cuttle-fish, 16 gavials, 23
cranes, 24 dogfishes, 23 gazelles, 25
crayfish, 19 hag-fishes, 23 geese, 24
creepers, 25 jawless fishes, 23 gibbons, 25
crickets, 20 lobe-finned fishes, 23 giraffes, 25
mole-crickets, 20 lungfishes, 23 glass-sponges, 10
crinoids, 21 rabbit-fishes, 23 gnats, 20
crocodiles, 23 ratfishes, 23 glow-worms, 20
croton-bugs, 20 ray-finned fishes, 23 goats, 25
crows, 25 silver-fish, 20 goatsuckers, 24
crustaceans, 7, 18 starfishes, 21 gordian-worms, 6, 14
cuckoos, 24 fish-flies, 20 gorgonians, 1

cuttle-fish, 16 flagellates, 9 gorillas, 25


animal-like, 9 graptolites, 27
plant-like, 9 grasshoppers, 20
flamingoes, 24 grebes, 24
daddy-long-legs, 18 flatworms, 6, 12 grosbeaks, 25
damselflies, 20 fleas, 20 grouse, 24
deer, 25 snow-fleas, 20 grylloblattids, 20
devil's-daming-needles, 20 water-fleas, 18 guanacos, 25
dinoflagellates, 9 flies, 20 guenons, 25
dippers, 25 butterflies, 20 guinea-pigs, 25
divers, 24 caddis-flies, 20 gulls, 24
dobson-flies, 20 case-flies, 20
dodos, 24 damselflies, 20
dogfishes, 23 dobson-flies, 20 hag-fishes, 23
dogs, 25 dragonflies, 20 hammerheads, 24
dolphins, 25 fireflies, 20 hares, 25
dormice, 25 fish-flies, 20 sea-hares, 15
doves, 24 ichneumon-flies, 21 harriers, 24
dragonflies, 20 lantern-flies, 20 harvest-men, 18
drongos, 25 mayflies, 20 hawks, 24
ducks, 24 orl-flies, 20 heart-urchins, 21,22
dust-lice, 20 salmon-flies, 20 hedgehogs, 25
sawflies, 20 herons, 24
scorpion-flies, 20 hexacorals, 1

serpent-flies, 20 hippopotamuses, 25
eagles, 24 snake-flies, 20 hoatzins, 24
ear-shells, 15 stone-flies, 20 honey-creepers, 25
earthworms, 17 trout-flies, 20 honey-guides, 24
earwigs, 20 white-flies, 20 hoopoes, 24
echidna, 25 flukes, 12 hornbills, 24
echinoderms, 21 flycatchers, 25 hornets, 21
eels flying-lemurs, 25 horntails, 21
slime-eels, 23 foraminiferans, 9 horny-corals, 1

elaters, 20 forams, 9 horny-sponges, 10


elephants, 25 fowls, 24 horsehair-worms, 6, 14
embiids, 20 foxes, 25 horses, 25
emus, 24 24
frigate-birds, horseshoe-crabs, 17
endoprocts, 14 frogmouths, 24 humming-birds, 24
INDEX TO Common names
11
hydroids, 6 lions nematodes, 14
hydrozoans, 31 ant-lions, 20 newts, 23
hyenas, 25 sea-lions, 25 night-crawlers,17
hyraxes, 25 lizards,23 nudibranchs, 16
llamas, 25 nuthatches, 25
lobe-finned fishes, 23
lobsters, 19
Ibises, 24 locusts, 20 Octopuses, 16
ichneumon-flies, 21 24 oil-birds, 24
loons,
insectivores, 25 oHgochaetes, 17
lories, 24
insects, 7, 20 lorises, 25 opalinids, 10
leaf-insects, 20 lungfishes, 23 opossums, 25
scale-insects, 20 opossum-shrimps, 19
lyrebirds, 25
stick-insects, 20 orangutans, 25
orioles, 25
macaques, 25 orl-flies, 20
macaws, 24 ospreys, 24
Jacamars, 24
maggots, 20
jacanas, 24 ostriches, 23
magpies, 25 25
japygids, 20 otters,
mammals, 25 ovenbirds, 25
jawless fishes, 23
mammoths, 25 owls, 24
jays, 25
mantids, 20
jellyfishes, 6, 11 oyster-catchers, 24
mantis-shrimps, 19
oysters, 16
marmosets, 25
marmots, 25
kagus, 24 marsupials, 25
pacas, 25
kangaroo-rats, 25 mastodons, 25 pandas, 25
kangaroos, 25 mayflies, 20 pangolins, 25
katydids, 20 meal-worms, 20 parakeets, 24
kingfishers, 24 mealy-bugs, 20 parrots, 24
kinglets, 25 medusae, 6, 11 peacocks, 24
kinkajous, 25 megapodes, 24 pearly-nautilus, 16
kiwis, 24 men, 25 peccaries, 25
koalas, 25 mice, 25 pelecypods, 16
krill, 19 pocket-mice, 25
pelicans, 24
microscorpions, 18 penguins, 23
microsporidians, 10 petrels, 24
midges, 20 phalangers, 25
lace-wings, 20
millepores, 11
lampreys, 23 phalaropes, 24
millers, 20 pheasants, 24
lamp-shells, 7, 15
millipedes, 19
lancelets, 7. 23 phoronids, 15
minks, 25 phyllopoda, 18
land-snails, 16
mites, 18
langurs, 25 piculets, 24
mockingbirds, 25 pigeons, 24
lantern-flies, 20
mole-crickets,20 pigs, 25
larks, 25
moUusks, 7, 15 pikas, 25
lars, 25
monkeys, 25 pillbugs, 19
leaf-hoppers, 20
monotremes, 25 placentals, 25
leaf-insects, 20
mosquitoes, 20
leeches, 17 plainwanderers, 24
mosquito-hawks, 20
lemmings, 25 planarians, 12
moss-animals, 7, 15
lemurs, 25 plantain-eaters, 24
moths, 20
flying-lemurs, 25 platypus, 25
motmots, 24 plovers, 24
lice, 20
mouse-birds, 24
bark-lice, 20 pocket-mice, 25
murres, 24
bird-lice, 20 polychaetes, 17
musk-oxen, 25
biting-lice, 20 polyclads, 12
muskrats, 25
book-lice, 20 porcupines, 25
mussels, 16
dust-lice, 20 porpoises, 25
sucking lice, 20 potoos, 24
limpets, 15 nautilus, 16 praying-mantis, 20
limpkins, 24 nemas, 14 prawns, 19
1 1 8 8

INDEX TO Common names 78


proboscis-worms, 6, 13 sea-cucumbers, 22 songbirds, 25
pronghoras, 25 sea-fans, 11 soothsayers, 20
proturans, 20 sea-feathers, 1 sowbugs, 19
pseudoscorpions, 18 sea-hares, 15 spiders, 18
psocids, 20 sea-lilies, 21 black-widows, 18
psyllids, 20 sea-lions, 25 scorpion-spiders, 55
pteropods, 16 sea-urchins, 21 sea-spiders, 17
24
puflFbirds, seals, 25 sun-spiders, 18
punkies, 20 sea-mats, 15 spiny-headed-worms, 6, 13
sea-mosses, 15 spittle-bugs, 20
sea-pansies, 11 sponges, 6, 10
quails, 24 sea-pens, 11 calcareous sponges, 10
sea-spiders, 17 chalky sponges, 10
sea-squirts, 7, 22 glass-sponges, 10
rabbit-fishes, 23 sea-stars, 21 horny-sponges, 10
rabbits, 25 sea-walnuts, 6, 12 stone-sponges, 10
raccoons, 25 secretary-birds, 24 spoonbills, 24
radiolarians, 9 segmented-worms, 7, 16 springtails, 20
rails, 24 serpent-flies, 20 squids, 16
ramshoms, 16 serpent-stars, 21 squirrels, 25
ratfishes, 23 sharks, 23 starfishes, 21
rats, 25 shearwaters, 24 starlings, 25
kangaroo-rats, 25 sheep, 25 stick-insects, 20
ray-finned fishes, 23 shells stilts, 24
rays, 23 boat-shells, 16 stony-corals, 11
reptiles,23 butterfly-shells, 16 stone-flies, 20
rheas, 24 ear-shells,15 stone-sponges, 10
rhinoceroses, 25 lamp-shells, 7, 15 storks, 24
rhizopods, 9 tooth-shells, 16 stromatoporoids, 31
ribbon- worms, 6, 13 tusk-sheUs, 16 stylopids, 21
roaches, 20 shrews, 25 sucking lice, 20
roadrunners, 24 tree-shrews, 25 sun-animalcules, 9
rock-jumpers, 20 shrikes, 25 sunbitterns, 24
rollers, 24 shrimps, 19 sungrebes, 24
rotifers, 6, 13 brine-shrimps, 18 sun-spiders, 18
round- worms, 6, 14 fairy-shrimps, 1 surf-birds, 24
mantis-shrimps, 19 swallows, 25
opossum-shrimps, 19 swans, 24
Salamanders, 23 silicoflagellates, 9 swifts, 24
salmon-flies, 20 silver-fish, 20
sand-dollars, 21, 22 siphonophores, 11
sandgrouse, 24 sipunculid worms, 16 tailless-whip-scorpions, 1

sandpipers, 24 skates, 23 tanagers, 25


sand-stars, 21 skimmers, 24 tapeworms, 12
sandworms, 17 skippers, 20 tapirs,25
sawflies, 21 skunks, 25 tarantula-hawks, 21
scale-insects, 20 slime-eels, 23 tarantulas, 18
scuds, 19 slime-molds, 9 tarsiers, 25
scorpion-flies, 20 sloths, 25 Tasmanian-wolf, 25
scorpions, 18 slugs, 15, 16 tenrecs, 25
book-scorpions, 18 snails, 15 termites, 20
false-scorpions, 18 land-snails, 16 terns, 24
microscorpions, 18 snakebirds, 24 thrashers, 25
pseudoscorpions, 18 snake-doctors, 20 thread-worms, 6, 14
tailless-whip-scorpions, snake-flies, 20 thrips,20
18 snakes, 23 thorny-corals, 12
whip-scorpions, 18 snipe, 24 thrushes, 25
scorpion-spiders, 55 snow-fleas, 20 ticks, 18
screamers, 24 soft-corals, 1 tinamous, 24
sea-anemones, 6, 11 solitaires, 24 tintinnids, 10
sea-cows, 25 solpugids, 18 titmice, 25
79 INDEX TO Common names
toads, 23 voles, 25 wood-wasps, 21
todies,24 vultures, 24 woolly-bears, 20
tongue-worms, 22 worms
tooth-shells, 16 acorn-worms, 22
tortoises, 23 Wagtails, 25 angle-worms, 17
toucans, 24 walking-sticks, 20 arrow-worms, 7,21
touracos, 24 walruses, 25 beard-worms, 7, 21
tree-hoppers, 20 warblers, 25 earthworms, 17
tree-shrews, 25 wasps, 21 flatworms, 6, 12
triclads, 12 gall-wasps, 21 glow-worms, 20
trogans, 24 tarantula-hawks, 21 gordian- worms, 7, 14
tropicbirds, 24 wood-wasps, 20 horsehair-worms, 7, 14
trout-flies,20 water-bears, 7, 17 meal-worms, 20
trumpeters, 24 water-fleas, 18 proboscis-worms, 6, 13
tubeworms, 17 water-pennies, 20 ribbon-worms, 6, 13
tunicates, 22 wattlebirds, 25 round-worms, 6, 14
turbans, 15 waxwings, 25 segmented-worms, 7,
turkeys, 24 webspinners, 20 16
turnstones, 24 weasels, 25 spiny-headed-worms,
turtles, 23 weaverbirds, 25 6, 13
tusk-shells, 16 weevils, 20 sandworms, 17
whales, 25 sipunculid worms, 16
wheel-animalcules, 6, 13 tapeworms, 12
Urchins whip-scorpions, 18 thread-worms, 6, 14
cake-urchins, 22 white-ants, 20 tongue-worms, 22
heart-urchins, 21,22 white-flies, 20 tubeworms, 17
sea-urchins, 21 white-grubs, 20 wire-worms, 20
wire-worms, 20 wrens, 25
wolverines, 25 wrentits, 25
Vampires, 25 wolves, 25
velvet-ants, 21 woodcock, 24
vertebrates, 7, 23 woodhewers, 25 Zebras, 25
vireos, 25 woodpeckers, 24 zorapterans, 20
INDEX TO LATIN NAMES

Acalephae, 44 Actinoceroida, 51 Amblydactyla, 70


Acantharia, 40 Actinoidea Amblypoda, 71
Acanthaspida Coelenterata, 44 Amblypygi, 55
Graptozoa, 42 Echinodermata, 60 Ambonodonta, 50
Vertebrata, 64 Actinomyxidia, 9, 40 Ambulatoria, 58
Acanthaspidomorpha, 64 Actinophrydea, 40 Amiiformes, 65
Acanthinocyathida, 42 Actinopoda Amioidea, 65
Acanthistida, 42 Echinodermata, 62 Ammonitida, 51
Acanthobdellida, 17, 52 Protozoa, 9, 40 Ammonitoidea, 50
Acanthobdelliformes, 52 Actinopterygii, 23, 65 Amniota, 64
Acanthocephala, 6, 13, 36, Actinozoa, 44 Amoebaea, 40
46 Actipylaea, 40 Amoebina, 40
Acanthocyatha, 42 Aculifera, 49 Amoebogeniae, 40
Acanthodiformes, 64 Acystosporidia, 40 Amoebosporidia, 40
Acanthodii, 64 Adapedonta, 51 Amoebozoa, 9, 40
Acanthodioidea, 64 Adeleida, 41 Amphiaspiformes, 64
Acanthoessi, 64 Adelochorda, 29 Amphibia, 23, 66
Acanthoessiformes, 64 Adelospondyli, 66 Amphibioidea, 66
Acanthoglossa, 64 Adenopoda, 58 Amphicoela, 67
Acanthometrida, 40 Adinida, 39 Amphidiscophora, 41
Acanthophractida, 40 Adunata, 60 Amphigastropoda, 49
Acanthopterygii, 66 Aegophiurida, 61 Amphilinidea, 12, 46
Acanthothoraci, 64 Aepyornithes, 68 Amphimorphae, 69
Acari, 54 Aepyornithiformes, 68 Amphineiira, 15,49
Acarida, 18, 54 Aeroplanoptera. 58 Amphioxi, 23, 63
Acarina, 54 Aeropneusta, 54 Amphipoda, 19. 56
Accipitres, 69 Aetheodontiformes, 65 Amphirhina, 64
Accipitriformes, 69 Aetioptera, 58 Amphirhyncha. 64
Acephala, 50 Aganasterida. 21.61 Amphoridea, 60
Acerata, 53 Agelacrinoidea, 61 Anacanthoidea, 66
Acetabulifera, 50 Aglaspida, 54 Anamerentoma, 57
Acineta, 41 Aglossa, 50 Anamnia. 64
Acinetaria, 41 Agnatha Anamorpha, 57
Acipenseriformes, 65 Arthropoda, 58 Anapsida, 23, 67
Acipenseroidea, 65 Vertebrata, 23, 64 Anarthrodira, 64
Acnidaria, 36, 44 Agnathes, 59 Anaspida. 64
Acnidosporidia, 40 Agnostida, 54 Anaspidacea, 19, 56
Acochlidiacea, 50 Agnotozoa, 6, 36 Anaspidea, 50
Acoela Aistopoda, 66 Anaspides, 56
Mollusca, 16,50 Ajacicyathida, 42 Anaspidoidea, 64
Platyhelminthes, 12, Alcedines, 69 Anatinacea, 50
45 Alciformes, 69 Ancylopoda, 70
Acoelomata, 6, 36 Alcyonacea, 11,44 Ancyrotricha. 19, 57
Acopa, 63 Alcyonaria, 11,44 Androgyna. 49
Acrania, 37, 63 Alectoromorphae, 68 Anemineae, 40
Acrasiae, 40 Alloeocoela, 12,45 Anguilliformes, 66
Acraspeda, 44 AUogromidiaceae, 40 Anguilluloidea, 47
Acrosauria, 67 Alloiocoela, 45 Angusticoela, 67
Acrothoracica. 18,56 Allotheria, 69 Animalia, 6, 36
Actinanthida, 44 Allotriognathoidea, 66 Anisaxia, 58
Actiniaria, 1 1, 44 Amaurochaetaceae, 40 Anisomyaria, 50
Actinildea, 44 Amaurochaetineae, 40 Anisopleura, 49
Actinistia, 66 Amaurosporales, 40 Anisopoda, 56
8i INDEX TO Latin names
Anisoptera, 58 Apostomina, 41 Ascoceroida, 51
Annelida, 7, 16, 37, 52 Appendicularia, 63 Asconosa, 41
Annulata, 37, 52 Apsidospondyli, 67 Ascorhynchomorpha, 54
Anoecia, 43 Aptera, 57 Ascospermophora, 19, 57
Anomalobranchia, 51 Apterygea, 24, 68 Ascothoracica, 19, 56
Anomalodesmacea, 51 Apterygiformes, 24, 68 Aseptata, 44
Anomalodesmata, 51 Apterygogenea, 57 Asiphonida, 50
Anomocoela, 67 Apterygota, 20, 57 Aspidobothria, 45
Anomodontia, 68 Apygia, 49 Aspidobranchiata, 49
Anomomeristica, 52 Arachnida, 18, 54 Aspidocephah', 64
Anomostraca, 56 Arachnoidea, 54 Aspidochirota, 22, 62
Anomura, 56 Arachnomorpha, 53 Aspidochirotida, 62
Anopla, 46 Araeolaimoidea, 14, 47 Aspidocotylea, 45
Anoplura, 20, 59 Araeoscelida, 67 Aspidogastrea, 12,45
Anostraca, 18, 55 Araneae, 18,55 Aspidorhynchiformes, 65
Anseres, 24, 69 Araneida, 8, 55 Aspidorhynchoidea, 65
Anseriformes, 24, 69 Archaehymenoptera, 58 Aspidospondyli, 67
Antennata, 53 Archaeoceti, 70 Aspiraculata, 22, 63
Anthocyaiha, 42 Archaeocopida, 55 Aspirigera, 41
Anthomedusae, 43 Archaeocyatha, 36, 42 Aspirotricha, 41
Anthomorphida, 42 Archaeogastropoda, 15,49 Aspirotrichaceae, 41
Anthozoa, 11,44 Archaeophyllida, 42 Asplanchnaceae, 47
Anthozoariae, 44 Archaeopteryges, 68 Asteroidea, 21, 61
Anthracomarti, 54 Archaeopterygiformes, 68 Asterolepiformes, 64
Anthracomartida, 54 Archaeornithes, 68 Asterozoa, 60
Anthracomorphoidea. 67 Archaeostraca, 55 Asthenodonta, 51
Anthracosauria, 67 Archiacanthocephala, 13, Asthenodontida, 50
Anthracosauroidea, 67 46 Astomata, 41
Anthropoidea, 70 Archiannelida, 17, 52 Astomina, 41
Antiarchi, 64 Archichaetopoda, 52 Astrapotheria, 71
Antiarchoidea, 64 Archinacelloidea, 49 Astraspiformes, 64
Anticorallia, 44 Archipolypoda, 57 Astromonaxonellida, 42
Antipatharia, 12,44 Archipterygota, 58 Astrophora, 42
Antipathidea, 44 Architarbi, 54 Astrorhizidea, 40
Antliata, 59 Architarbida, 54 Ateleopiformes, 66
Anura, 23, 66 Architeuthacea, 51 Ateleopoidea, 66
Aphacellae, 43 Archodonata, 57 Atelostomata, 62
Aphaneura, 52 Archonta, 70 Atentaculata, 45
Aphaniptera, 59 Archosauria, 67 Athecanephria, 21, 60
Aphasmidia, 47 Archosauromorpha, 67 Athecata, 43
Aphelophlebia, 58 Arctolepiformes, 64 Atomiosoma, 46
Aphetohyoidea, 64 Arcyriaceae, 40 Atremata
Aphrothoraca, 40 Armadillomorpha, 57 Brachiopoda, 15,49
Aphylles, 45 Arthrocephala, 53 Tunicata, 63
Aplacophora, 15,49 Arthrodira, 64 Aulodonta, 62
Aplocoela, 36, 46 Arthrodiroidea, 64 Auluroidea, 61
Aplosporidla, 40 Arthropleurida, 53 Autobranchiata, 50
Aplousobranchiata, 22, 63 Arthropoda, 7, 17,37,53 Autoscolecida, 36
Aplysiacea, 50 Arthropomata, 49 Aves, 23,68
Apoda Arthrostraca, 56 Axifera, 44
Amphibia, 23, 66 Arthrotardigrada, 17, 52 Azygobranchia, 50
Arthropoda, 19, 56 Articulata
Aves, 24,69 Brachiopoda, 15, 49
Echinodermata, 22, 62 Coelomata, 27 Babesiida, 41
Apodes, 66 Echinodermata, 21,60 Balanoglossida, 22, 63
Apodiformes, 69 Artiodactyla, 25,71 Barrandeoceratida, 51
Apodoidea, 66 Ascaroidea. 14, 48 Barypoda, 71
Apomatostoma, 50 Aschelminthes, 28 Barytheria, 70
Aporhabdina, 41 Ascidiacea, 22. 63 Basommatophora, 16, 50
Aporidea, 12, 46 Ascidiastella, 62 Bassleroceratida, 51
Aporita, 60 Asclerocorallia, 44 Bathyclupeiformes, 66
INDEX TO Latin names 82
Bathyclupeoidea, 66 Bunotheria, 70 Cephalocarida, 18, 55
Bathyuriscidea, 54 Burgessiida, 53 Cephalochordata, 7, 23, 37,
Batoidea, 23, 65 63
Batrachia, 23, 66 Cephalodiscidea, 22, 62
Batrachoidea, 66 Caducichordata, 63 Cephalophora, 48
Batrachoidiformes, 66 Caecilia, 66 Cephalopoda, 16, 50
Batrachosauria, 67 Caenagnathae, 68 Cephalostigmata, 19, 57
Bdellacea, 13,46 Caenagnathiformes, 68 Cephaloxeniformes, 65
Bdelloidaceae, 46 Caenogastropoda, 15, 50 Ceratiocarina, 56
Bdelloidea, 13,46 Caenolestoidea, 70 Ceratiomyxaceae, 40
Bdellomorpha, 46 Calanoida, 18, 55 Ceratocarina, 56
Bdellonemertea, 13,46 Calcarea, 10,41 Ceratodonti, 66
Belemnoida, 51 Calcarineae, 40 Ceratodontiformes, 67
Bellerophontacea, 49 Calcispongea, 10, 41 Ceratophora, 37, 52
Beloniformes, 66 Caligoida, 18,56 Ceratosa, 41
Beroida, 12,45 Calonemineae, 40 Ceratosida, 41
Beryciformes, 66 Caloneurodea, 58 Ceratospongida, 41
Berycomorphoidea, 66 Calopteriformes, 65 Ceriantharia, 12,45
Beyrichiida, 55 Calycophora, 43 Cerianthiae, 45
Bilateria, 27 Calycozoa, 44 Cerianthidea, 45
Biporophyllidea, 12,46 Calyptoblastea, 43 Ceriantipatharia, 12, 45
Birkeniiformes, 64 Calyssozoa, 6, 14, 36, 48 Cestida, 12,45
Bivalvia, 16, 50 Camallanta, 48 Cestoda, 12,45
Blastoidea, 60 Camarodonta, 62 Cestodaria, 12, 46
Blattaeformia, 58 Camaroidea, 43 Cestoidea
Blattaria, 20, 58 Camarostomata, 55 Ctenophora, 45
Blattoidea, 58 Cambridioidea, 49 Platyhelminthes, 45
Bobasatraniiformes, 65 Camerata, 60 Cetacea, 25,70
Bolinopsidea, 45 Campanulariae, 43 Cete, 70
Bothriocephaloidea, 46 Campodeoidea, 57 Chaetetida, 44
Bothriocidaroida, 62 Canaliculata, 61 Chaetodermatida, 15,49
Bothriocidaroidea, 62 Caprimulgi, 24, 69 Chaetodermomorpha, 49
Botryoidea, 40 Caprimulgiformes, 24, 69 Chaetognatha, 7,21,37,59
Brachiata Carinatae, 68 Chaetonotidea, 13, 47
Echinodermata, 60 Carnivora, 25, 70 Chaetonotoidea, 13,47
Pogonophora, 37, 59 Carnivores, 70 Chaetopoda, 17, 52
Brachionocephala, 48 Carnosida, 10, 42 Chaetosomatida, 47
Brachionoidea, 47 Carpoidea, 61 Chalarothoraca, 40
Brachionopoda, 48 Carybdeida, 44 Chalicotheria, 70
Brachiopoda, 7, 15, 37,49 Caryophyllacea, 45 Charadriae, 24, 69
Brachyopterygii, 65 Caryophyllidea, 12, 46 Charadriiformes, 24, 69
Brachythoraci, 64 Cassiduloida, 22, 62 Charybdeida, 44
Brachyura, 56 Casuarii, 24, 68 Chaudhuriiformes, 66
Bradorina, 55 Casuariiformes, 24, 68 Chaudhurioidea, 66
Bradyodonti, 65 Capitelliformia, 52 Cheiloctenostoma, 48
Bradyodontoidea, 65 Catenata, 39,42 Cheilostomata, 15,48
Branchiata, 53 Catenulida, 45 Cheilostomellaceae, 40
Branchionobranchia, 48 Cathartidiformes, 69 Cheiracanthiformes, 64
Branchiopoda Caudata, 66 Cheiropterygia, 64
Brachiopoda, 48 Caulogastra, 18, 54 Cheleutoptera, 58
Crustacea, 18, 55 Cecomorphae, 68 Chelicerata, 52
Branchiostomoidea, 63 Centrechinoida, 21,62 Chelifera, 56
Branchiotremata, 29 Centroceratida, 51 Chelonethida, 54
Branchiura, 18,56 Centrohelidia, 40 Chelonia. 23, 67
Breyeridea, 58 Cephalacanthoidea, 66 Cheloniellida, 53
Brooksellida, 43 Cephalaspida, 64 Chenomorphae, 69
Bruta, 70 Cephalaspidea, 50 Chernetes, 54
Bryozoa, 7, 15,37,48 Cephalaspidiformes, 64 Chernetidea, 54
Bucerotes, 69 Cephalaspidomorpha, 23, Chilognatha, 19, 57
BuUomorpha, 50 64 Chilophiurida, 21,62
Bunodomorpha, 54 Cephalobaenida, 17, 53 Chilopoda, 19, 57
83 INDEX TO Latin names
Chilostomata, 48 Coccidiomorpha, 40 Coronata, 60
Chilostomellidea, 40 Coccolithophorida, 9, 39 Coronatae, 11,44
Chimaerae, 23, 65 Coccostei, 64 Coronatida, 44
Chimaeriformes, 65 Coccosteiformes, 64 Corrodentia, 20, 59
Chimaeroidea, 65 Coccosteomorphi, 64 Corticata, 39
Chiroptera, 25,70 Coccyges, 69 Corynexochida, 54
Chitonida, 49 Cochlides, 49 Coryphodontia, 70
Chlamydomonadina. 39 Cochliostraca, 49 Costata, 60
Chlamydophora, 40 Coelacanthiformes, 66 Cotylosauria, 67
Chloromonadaceae, 39 Coelacanthini, 66 Craniata, 37, 64
Chloromonadina, 9, 39 Coelenterata, 6, 11, 36, 43 Craspedomonadina, 39
Choanichthyes, 23, 66 Coelhelminthes, 37, 52 Craterostigma, 19, 57
Choanoflagellata, 39 Coelolepida, 64 Creodonta, 70
Chondrenchelyes, 65 Coelolepidoidea, 64 Crepipoda, 49
Chondrenchelyiformes, 65 Coelolepiformes, 64 Crinoidea, 21, 60
Chondrichthyes, 23, 65 Coelomata, 7, 37 Crocodilia, 23, 67
Chondrophora, 43 Coenothecalia, 11,44 Crossopterygii, 23, 66
Chondropterygii, 65 Coleoidea, 50 Crossopterygioidea, 66
Chondrostei, 23, 65 Coleoptera, 20, 59 Crustacea, 18, 55
Chondrosteoidea, 65 Coleopteroidea, 58 Cryptocephala, 52
Chonotricha, 10, 41 Colii, 24,69 Cryptocoela
Chordata, 30 Coliiformes, 24, 69 Trematoda, 45
Choristida, 10,42 Collaria, 36,44 Turbellaria, 45
Chromadoroidea, 14, 47 Collembola, 20, 58 Cryptodonata, 58
Chromomonadina, 39 Colloconchida, 50 Cryptodonta, 51
Chrysomonadaceae, 39 Collodaria, 40 Cryptodontia, 58
Chrysomonadina, 9, 39 Collothecacea, 13,47 Cryptomonadaceae, 39
Ciconiae, 24, 69 Colobognatha, 19, 57 Cryptomonadina, 9, 39
Ciconiiformes, 24, 69 Colopyga, 55 Cryptoparamera, 44
Cidaroida, 21, 62 Colossendeomorpha, 54 Cryptostigmata, 55
Ciliata, 10,41 Columbae, 24, 69 Cryptostomata, 48
Ciliatoidea, 41 Columbiformes, 24, 69 Cryptozonia, 61
Cilioflagellata, 39 Colymbae, 24,68 Crypturi, 24, 68
Ciliophora, 39 Colymbiformes, 24, 68 Crypturiformes, 24, 68
Cirrhobranchiata, 50 Colymbomorphae, 68 Ctenarea, 36, 44
Cirrhostomi, 37, 63 Comatulida, 21, 61 Ctenobranchiata, 50
Cirrigrada, 61 Compsognatha, 68 Ctenodipterini, 66
Cirripedia, 18, 56 Conchifera, 50 Ctenodontiformes, 67
Cladida, 60 Conchophora, 50 Ctenophora, 6, 12, 36,45
Cladistia, 65 Conchorhaga, 47 Ctenophoraria, 36,44
Cladistioidea, 65 Conchostraca, 18, 55 Ctenoplanidea, 45
Cladocera, 18,55 Condylarthra, 70 Ctenostomata, 15,48
Cladocopa Conoclypina, 62 Ctenostomina, 41
Arthropoda, 55 Conocoryphida, 54 Cubomedusae, 11,44
Mollusca, 50 Conodonta, 64 Cuculi, 24, 69
Cladodontiformes, 65 Conodontophorida, 64 Cuculiformes, 24, 69
Cladoidea, 60 Conularida, 36, 43 Cucullifera, 55
Cladophiurae, 61 Conulariida, 43 Cucumariida, 62
Cladoselachiformes, 65 Conulata, 43 Cumacea, 19,56
Cladoselachii, 65 Copelata, 63 Cursoria, 58
Cladoselachoidea, 65 Copeognatha, 59 Cyamoidea, 61
Climatiiformes, 64 Copepoda, 18, 55 Cyathaspiformes, 64
Clistenterata, 49 Coraciae, 24, 69 Cyathocrinacea, 61
Clitellata, 52 Coraciiformes, 24, 69 Cyathospongia, 36, 42
Clupeiformes, 66 Coraciomorphae, 68 Cyclicozoa, 44
Clypeastroida, 22, 62 Coralla, 44 Cyclobranchia, 49
Cnemidolestoidea, 59 Corallaria, 44 Cyclocorallia, 44
Cnidaria, 6, 11,36,43 Coralligena, 44 Cyclodonta, 51
Cnidosporidia, 9, 40 Corallimorpharia, 11,44 Cycloidea, 61
Coadunata, 60 Cornacuspongia, 41 Cyclomyaria, 22, 63
Coccidia, 9,40 Corona, 44 Cyclophyllidea, 12, 46
INDEX TO Latin names 84
Cyclopoida, 18, 55 Diapsida, 23, 67
Cyclorhaga, 47
Cyclostomata
Bryozoa, 15,48
Vertebrata, 23, 64
Cydippida, 12,45
Cynostraca, 49
Cypriniformes, 66
Cyprinodontiformes, 66
Cyprinodontoidea, 66
Cyrtellaria, 40
Cyrtochoanites. 51
Cyrtocrinida, 21,60
Cyrtoidea, 40
Cystasteroidea, 61
Cystechinoidea, 60
Cystica, 45
Cystidea, 60
Cystocidaroidea, 60
Cystocrinoidea, 60
Cystoflagellata, 39
Cystoidea, 60
Cytoidea, 39
Cytomorpha, 39
C)^ophora, 40
Cytosporidia, 40

Dacnostomata, 58
Dactylifera, 45
Dactylopteriformes, 66
Daubentomioidea, 70
Decabrachia, 50
Decapoda
Arthropoda, 19, 56
Mollusca, 16,50
Decembrachiata, 50
Deinocephalia, 68
Delobranchia, 54
Demospongea, 10,41
Dendroceratina, 41
Dendrochirota, 22, 62
Dendrochirotida, 62
Dendroidea, 43
Deratoptera, 58
Dermaptera, 20, 58
Dermodermaptera, 58
Dermoptera, 25, 70
Desmodonta, 51
Desmomyaria, 22, 63
Desmophora, 41
Desmoscolecoidea, 14, 47
Desmospongea, 41
Desmothoraca, 40
Diadecta, 67
Diadectomorpha, 67
Diadematoida, 62
Diademoida, 62
Dialytina, 41
Diantennata, 53
Diaphanopteroidea, 58
85 INDEX TO Latin names
Embioidea, 58 Eucarida, 56 Ferungulata, 70
Embioptera, 20, 58 Euceratosa, 41 Filarioidea, 14, 48
Embolobranchiata, 54 Eucestoda, 45 Filibranchia, 16, 50
Embolomeri, 66 Euchorda, 37, 64 Filosa, 39
Embrithopoda, 71 Euciliata, 10,41 Fissipedia, 70
Emeraldellida, 53 Eucirripedia, 56
Fistulata, 61
Emmenognatha, 59 Eucoelomata, 37
Fistulides, 62
Emydosauria, 67 Eucopepoda, 18, 55
Flagellata, 9, 39
Encrinasteriae, 61 Eucrinoidea, 60
Endeiolepiformes, 64 Eucrustacea, 55 Flexibilia, 60
Endoaria, 44 Eucystoidea, 60 Flosculariacea, 13,47
Endobranchiata, 62 Eudesmodontida, 50 Foraminifera, 9, 40
Endoceroida, 51 Euechinoidea, 62 Forcipulata, 21,61
Endocyclica, 62 Euentomata, 57
Endoprocta, 6, 14, 36, 48 Euflagellata, 39
Endopterygota, 20, 59 Euglenaceae, 39 Gadiformes, 66
Endospora, 40 Euglenida, 39 Galaxiiformes, 66
Endosporeae, 40 Euglenoidina, 9, 39 Galeodea, 55
Enopla, 46 Eugnatha, 57 Galeoidei, 65
Enoplata, 47 Eugregarinida, 41 Galli, 24,69
Enoploidea, 14, 47 Eulamellibranchia, 16, 50 Galliformes, 24,69
Enterocoela, 6, 27, 36 Eumalacostraca, 56 Gallinae, 69
Enterogona, 63 Eumetazoa, 36 Gasterozoa, 49
Enteropneusta, 7, 22, 37, 63 Eumycetozoa, 40 Ganodonta, 70
Enterozoa, 27, 36 Eunotosauria, 67 Ganoidi, 65
Entodiniomorphina, 41 Euonychophora, 17, 53 Gasteropoda, 49
Entognatha, 57 Euornithes, 68 Gasteropodophora, 49
Entomocrania, 37, 63 Eupantopoda, 17, 54 Gasterosteiformes, 66
Entomostraca, 55 Euphaneroidea, 64 Gasterostoma, 45
Entoprocta, 14, 36, 48 Euphausiacea, 19, 56 Gastornithes, 68
Entotrophi, 20, 57 Euphyllopoda, 55 Gastraeopoda, 49
Eoacanthocephala, 13, 46 Euplasmodida, 40 Gastrocaulia, 49
Eoanura, 66 Euplexoptera, 58 Gastropoda, 15,49
Eocrinoidea, 61 Eupoliida, 46 Gastrotricha, 6, 13,36,47
Eodiscida, 54 Euryalae, 61 Gaviae, 24, 68
Eopaleodictyoptera, 58 Euryalida, 61 Gaviiformes, 24, 68
Eoplacophora, 49 Euryapsida, 67 Gemuendiniformes, 64
Eosuchia, 67 Eurylaemi, 69 Geophilomorpha, 19, 57
Eotetrapoda, 64 Eurypterida, 54 Geophylomorpha, 57
Eozoa, 6, 36, 39 Eurystema, 57 Gigantostraca, 54
Epectinata, 54 Eurystomata, 48 Giganturiformes, 66
Ephemerida, 20, 58 Euselachii, 65 Giganturoidea, 66
Ephemeroidea, 58 Eusuchia, 67 Ginglymodi, 65
Ephemeroptera, 58 Eutardigrada, 17, 52 Glires, 70
Epimorpha, 57 Eutaxodonta, 50 Globidia, 10,40
Epiparia, 54 Eutheria, 25,70 Globigerinidea, 40
Epipolasida, 10, 42 Euthyneura, 49 Glossata, 59
Eplacentalia, 69 Euxiphosura, 54 Glosselytrodea, 58
Erpobdellida, 52 Exocyatha, 42 Glossophora, 48
Errantia, 17, 52 Exocyclica, 62 Gnathobdellida, 17, 52
Eterocotylea, 45 Exocycloida, 21,62 Gnathobdelliformes, 52
Euamphibia, 67 Exopterygoptera, 58 Gnathophiurida, 21,61
Euanostraca, 55 Exopterygota, 20, 58 Gnathopoda, 54
Euanura, 66 Exosporae. 40 Gnathostomata
Eu-arachnida, 54 Exosporidia, 41 Echinodermata, 62
Euarthrodira, 64 Extrasiphonata, 51 Vertebrata, 64
Euarthropoda, 37, 52 Gobiesociformes, 66
Euasteriae, 61 Goniactinida, 61
Eublastoidea, 60 Falcones, 24, 69 Gordiacea, 6, 14, 36, 48
Eubleptidodea, 57 Falconiformes, 24, 69 Gordididea, 48
Eubradyodonti, 65 Ferae, 70 Gordioidea, 14, 36, 48
INDEX TO Latin names
86
Gorgonacea, 11,44 Heliozoa, 9,40 Hexapoda, 20,57
Gradientia, 66 Helminthomorpha, 37, 63 Hexasterophora, 41
Grallae, 69 Hemibranchia, 50 Hippocrepia, 48
Graptoblasti, 42 Hemichordata, 29 Hirudinea, 17, 52
Graptolithida, 43 Hemimeroidea, 58 Histosporidia, 40
Graptoloidea, 43 Hemimyaria, 22, 63 Histozoa, 6, 36
Graptovermida, 42 Hemiodonata, 58 Holectypoida, 22, 62
Graptozoa, 36, 43 Hemipodii, 69 Holobranchia, 48
Gregarina, 40 Hemipomatostoma, 50 Holocephali, 23, 65
Gregarinida, 9, 40 Hemipsocoptera, 59 Hologonia, 47
Gressores, 69 Hemiptera, 58 Holomastigina, 39
Gressoria, 58 Hemizonida, 61 Holomerentoma, 57
Gromiidea, 40 Herocotylida, 45 Holometabola, 20, 59
Grues, 24, 69 Herodii, 69 Holopeltidia, 55
Gruiformes, 24, 69 Herodiones, 69 Holoptychiformes, 66
Grylloblattoidea, 20, 58 Herpobdellida, 52 Holosiphona, 50
Guttulinaceae, 40 Hesperornithes, 68 Holostei, 23, 65
Gymnamoebaea, 40 Hesperornithiformes, 68 Holothurioidea, 22, 62
Gymnoblastea, 43 Hetairacyathida, 42 Holothuroidea, 62
Gymnodiniaceae, 39 Heteractinellida, 41 Holotricha, 10, 41
Gymnolaemata, 15,48 Heteractinida, 10, 41 Holotrypasta, 40
Gymnomyxa, 39 Heterocerci, 65 Homalopterygia, 37, 59
Gymnonisciformes, 65 Heterochlorida, 39 Homalorhaga, 47
Gymnophiona, 23, 66 Heterocoelida, 41 Homalozoa, 60
Gymnosomata, 50 Heteroconchia, 50 Homocoelida, 41
Gymnosporidia, 40 Heterocoralla, 45 Homokaryota, 39
Gymnostomata, 41 Heterocorallia, 44 Homomeria, 37, 63
Gyracanthiformes, 64 Heterocotylea, 45 Homomyaria, 50
Gyracanthocephala, 46 Heterocyemida, 1 1 42
, Homopoda, 53
Gyrocotylidea, 12, 46 Heterodermaceae, 40 Homoptera, 20, 59
Gyrosomata, 50 Heterodonta, 51 Homosclerophora, 42
Heterodontida, 50 Hoplocarida, 56
Heterodontiformes, 65 Hoplonemertea, 13,46
Hadentomoidea, 59 Heterodontoidea, 65 Hyalospongea, 10,41
Hadromerida, 10, 42 Heterogangliata, 37, 48 Hybocrinida, 60
Haemacryma, 64 Heterognatha, 19, 56 Hydrariae, 43
Haematherma, 64 Heterokaryota, 39 Hydrida, 43
Haemocytozoa, 40 Heteromastigina, 39 Hydrocorallinae, 11,43
Haemosporidia, 9, 40 Heterometabola, 20, 58 Hydroida, 11,43
Halichondrida, 10,42 Heteromi, 66 Hydromedusae, 43
Hallopoda, 68 Heteromoidea, 66 Hydrophoridea, 60
Halosauriformes, 66 Heteromorphae, 69 Hydropneustea, 54
Halterata, 59 Heteromya, 50 Hydrosauria, 67
Halteriptera, 59 Heteromyaria, 50 Hydrozoa, 11,43
Hapalopteroidea, 59 Heteromyota, 16,51 Hydrozoaria, 43
Haplocrinacea, 61 Heteronemertea, 13,46 Hymenocarina, 53
Haplocyemata, 37, 63 Heterophyllidea, 45 Hymenoptera, 21, 59
Haplodoci, 66 Heteropoda, 50 Hymenopteroidea, 59
Haplodrili, 52 Heteroptera, 20, 59 Hymenostomata, 41
Haplosclerida, 10, 42 Heterosomata, 66 Hyparthropoda, 53
Haplosporidia, 10,40 Heterostelea, 60 Hypermastigina, 9, 39
Haplozoa, 60, 61 Heterostigmata, 55 Hyperoarti, 64
Haptopoda, 54 Heterostraci, 64
Hyperotreti, 64
Haptopodida, 54 Heterotardigrada, 17, 52
Harpacticoida, 18,55 Hypoparia, 54
Heterotricha, 41
Haustellata, 59 Hexaceratina, 41 Hypostoma, 60
Helianthoida, 44 Hexacorallia, 44 Hypostomosoidea, 66
Helicosporidia, 10,40 Hexactinellida, 10,41 Hypotremata, 65
Helioflagellida, 40 Hexactinia, 44 Hypotricha, 41
Heliolitida, 44 Hexanchiformes, 65 Hyraces, 71
Heliornithiformes, 69 Hexanchoidea, 65 Hyracoidea, 25, 71
87 INDEX TO Latin names
Ichthya, 64 Kustarachne, 55 Limicolae
Ichthyocrinacea, 60 Kustarachnida, 55 Annelida, 52
Ichthyodorulites, 64 Vertebrata, 68
Ichthyopsida, 64 Limulava, 53
Ichthyopterygia, 67 Labechioidea, 43 Limulavida, 53
Ichthyopterygii, 64 Labellata, 55 Limulida, 54
Ichthyornes, 68 Labidura, 58 Linguatulida, 17, 37, 53
Ichthyorniformes, 68 Labyrinthodontia, 66 Linguatulodea, 53
Ichthyornithes, 68 Labyrinthulidea, 39 Lipocephala, 50
Ichthyornithiformes, 68 Lacertilia, 67 Lipodonta, 51
Ichthyosauria, 67 Laemodipoda, 56 Lipoptera, 59
Ichthyostegalia, 66 Laemophiurida, 21,61 Lipostraca, 55
Lagenidea, 40 Lissoflagellata, 39
Ichthyotomi, 65
Lagomorpha, 25, 70 10, 42
Lithistida,
Icosteiformes, 66
Icosteioidea, 66 Lamellata, 49 Lithobiomorpha, 19, 57
Ictidosauria, 68 Lamellibranchiata, 16, 50 Lithonina, 41
Impennes, 68 Lamniformes, 65 Lithorhizostomeae, 44
Inadunata, 60 Lamnoidea, 65 Lithorhizostomatida, 44
Inarticulata Lampridiformes, 66 Litopterna, 70
Brachiopoda, 15,49 Lamprosporales, 40 Lituolidea, 40
Echinodermata, 60 Lariformes, 69 Lobata, 12,45
Inermia, 41 Laro-limicolae, 69 Lobosa, 39
Infusoria, 39 Larvacea, 22, 63 Loliginacea, 51
Iniomi, 66 Larvata, 61 Lophiiformes, 66
Illoricata, 47 Larviformia, 61 Lophogastridea, 19, 56
Insecta, 20, 57 Lasaniiformes, 64 Lophopoda, 15,48
Insectivora, 25, 70 Lateribranchiata, 50 Lophotricha, 19, 57
Integricephalida, 54 Laternulacea, 50 Loricata
Integripalliata, 51 Latigastra, 18, 54 Mollusca, 49
Latisterna, 55 Rotifera, 47
Intrasiphonata, 51
Irregulares, 60 Leanchoiliida, 53 Vertebrata, 67
Irregularia Lebetida, 10,41 Lucernaria, 44
Crinoidea, 61 Lecanicephala, 12,45 Lucernariidea, 44
Echinoidea, 22, 62 Lecanicephaloidea, 45 Luciae, 63
Ischnacanthiformes, 64 Lemuroidea, 70 Lucida, 63
Isedrolotila, 50 Leperditicopida, 55 Luganoidiiformes, 65
Isocrinida, 21, 60 Leperditiida, 55 Lumbricimorpha, 52
Isodonta, 51 Lepidocentroida, 62 Lychniskida, 41
Isofilibranchia, 50 Lepidoglossa, 49 Lychniskophora, 10,41
Isomya, 50 Lepidopleurida, 49 Lygogalaceae, 40
Isoplacophora, 49 Lepidoptera, 20, 59 Lyomeroidea, 66
Isopleura. 49 Lepidosauria, 67 Lyopomata, 49
Isopoda, 19, 56 Lepidosireniformes, 67 Lysophiurae, 61
Isoptera, 20, 58 Lepidostei, 65 Lysophiuroida, 61
Isopygia, 54 Lepidosteoidea, 65 Lyssacina, 41
Isospondyli, 66 Lepisostei, 65 Lyssakina, 10,41
Isospondyloidea, 66 Lepisosteiformes, 65
Isuriformes, 65 Lepospondyli, 23, 66
Leptocardia, 23, 63 Machaeridia, 61
Leptolinae, 43 Macrochires, 24, 69
Leptomedusae, 43 Macrochiriformes, 24, 69
Jagoriniformes, 65
Leptophya, 59 Macrociliobranchia, 50
Juliformia, 57
Leptostraca, 56 Macrodasyidea, 13,47
Lernaeopodoida, 18, 56 Macrodasyoidea, 13,47
Leuconosa, 41 Macropetalichthyida, 64
Kamptozoa, 36, 48 Libellulides, 58 Macropharyngea, 45
Katapsida, 67 Libelluloiden 58 Macrosterni, 57
Kathapsida, 67 Liceaceae, 40 Macrostomida, 45
Keratosida, 10,41 Lichacea, 54 Macrotrachia, 50
Kinorhyncha, 6, 13, 36,47 Lichida, 54 Macrura, 56
Krikobranchia, 63 Limacomorpha, 19, 57 Macruriformes, 66
INDEX TO Latin names 88
Madreporaria, 11,44 Metanemertini, 46 Monopylaria, 40
Malacactiniae, 44 Metapsida, 67 Monorhina, 64
Malachichthyes, 66 Metastigmata, 55 Monorhyncha, 64
Malacobothrii, 45 Metatheria, 25, 70 Monotocardia, 50
Malacocotylea, 45 Metazoa, 6, 36 Monotremata, 25, 69
Malacopoda, 37,52 Michelinoceroida, 51 Monozoa, 46
Malacopterygii, 66 Micramphibia, 66 Monstrilloida, 18, 56
Malacostraca, 19, 56 Microciliobranchia, 50 Monticuliporoidea, 48
Malacozoa, 37, 48 Microcyprini, 66 Mormyriformes, 66
Mallophaga, 20,59 Micropharyngea, 45 Mormyroidea, 66
Mammalia, 25, 69 Micropodi, 24, 69 Moruloidea, 36, 42
Mammifera, 69 Micropodiformes, 69 Mosasauria, 67
Mandibulata, 52 Micropygia, 54 Mugiliformes, 66
Mantodea, 20, 58 Microsauria, 66 Multisolenida, 44
Marellomorpha, 53 Microsclerophora, 42 Multituberculata, 69
Margaritaceae, 40 Microsporidia, 10, 40 Musophagi, 68
Marrellida, 53 Microthelyphonida, 55 Mutica, 70
Marrellomorpha, 53 Miliolidea, 40 Mutilata, 70
Marriocarida, 53 Milleporida, 11,43 Mycetophorae, 55
Marsipobranchii, 64 Millericrinida, 60 Mycetozoa, 9, 40
Marsupialia Mimetasterida, 53 Myctophiformes, 66
Coelenterata, 44 Miocoela, 36, 46 Myelozoa, 37, 63
Vertebrata, 25,70 Miomera, 54 Mylostomatiformes, 64
Mastacembeliformes, 66 Miomoptera, 59 Myodocopa, 55
Mastacembelioidea, 66 Mionelminthes, 36, 42 Myodocopida, 18, 55
Mastigophora. 9, 39 Mirientomata, 57 Myohyracoidea, 70
Mecaptera, 59 Miskoa, 52 Myophiurida, 21,61
Mecoptera, 20, 59 Mixochoanites, 51 Myosomata, 22, 63
Medamoptera, 59 Mixotermitoidea, 58 Myriapoda, 53
Megalichthyiformes, 66 Mollusca, 7, 15,37,49 Myrientomata, 57
Megalopoda, 62 Molpadida, 62 Mysidacea, 19, 56
Megaloptera, 59 Molpadonia, 22, 62 Mystacocarida, 18, 55
Meganisoptera, 58 Monactinellida, 41 Mystacoceti, 70
Megasecoptera, 58 Monadidea, 39 Myxini, 64
Megasecopterida, 58 Monaxonida, 41 Myxiniformes, 64
Megistanes, 68 Monhysteroidea, 14, 47 Myxinoidea, 64
Melicertida, 47 Moniligastres, 52 Myxogasteres, 40
Melonechinoida, 62 Monobathra, 60 Myxomycetes, 40
Melonitoida, 62 Monobathrida, 60 Myxospongida, 10, 41
Menotyphla, 70 Monoblastidea, 11,42 Myxosporidia, 9, 40
Memirae, 69 Monoblastoidea, 11,42 Myzostoma, 51
Meridogastra, 55 Monoblastozoa, 6, 11, 36, Myzostomaria, 51
Mermithoidea, 14, 47 42 Myzostomida, 7, 16, 37, 51
Merochaeta, 57 Monoceratina, 41
Merostomata, 17, 54 Monocyatha, 42
Merostomoidea, 53 Monocyathida, 42 Nahecarida, 56
Merotrypasta, 40 Monocyclica Naiadida, 50
Merozoa, 45 Coelenterata, 44 Naidomorpha, 52
Mesaxonia, 71 Echinodermata, 60 Narcaciontiformes, 65
Mesacanthiformes, 64 Monodelphia, 70 Narcomedusae, 43
Mesichthyes, 66 Monogenea, 12,45 Nasselaria, 40
Mesogastropoda, 50 Monogenetica, 45 Nautilida, 51
Mesonacida, 54 Monogononta, 13.47 Nautiloidea, 16, 50
Mesonemertini, 46 Monomerostomata, 54 Nebaliacea. 19,56
Mesoplacophora, 49 Monomya, 50 Nectaspida, 53
Mesosauria, 68 Monomyaria, 50 Nectonematoidea, 14, 48
Mesostigmata, 55 Monophyllidea, 45 Nectridia. 66
Mesosuchia, 67 Monopisthocotylea, 45 Nemata. 36, 47
Mesozoa, 6, 11, 36, 42 Monoplacophora, 15,49 Nemathelminthes, 28
Metacanthocephala, 46 Monopneumona, 66 Nematoda, 6, 14, 36, 47
Metacyathida, 42 Monopnoa, 67 Nematoidea, 14.47
89 INDEX TO Latin names
Nematomorpha, 6, 14, 36, Odonata, 20, 58 Ostariophysoidea, 66
48 Odontoceti, 70 Osteichthyes, 23, 65
Nematophora Odontognathae, 68 Osteolepides, 66
Arthropoda, 57
19, Odontolcae, 68 Osteolepidoti, 66
Coelenterata, 36, 43 Odontopleurida, 54 Osteolepiformes, 66
Nematozoa, 36, 43 Odontormae, 68 Osteostraci, 64
Nemertea, 46 Odontota, 58 Osteostracoidea, 64
Nemertinea, 6, 13, 36,46 Oecioa, 44 Ostracoda, 18,55
Nemuraedes, 58 Oesthelminthes, 37, 59 Ostracodermi, 64
Neoacanthocephala, 46 Olenellida, 54 Ostracopa, 55
Neoblattariae, 58 Oligochaeta, 17, 52 Ostracophori, 64
Neocrinoidea, 61 Oligocnemata, 64 Ostrapoda, 55
Neogastropoda, 50 Oligoentomata, 58 Otocardia, 37, 48
Neognathae, 68 Oligomeria, 54 Oxyuroidea, 14, 48
Neoloricata, 15,49 Oligoneura, 58
Neomeniida, 15, 49 Oligotrichina, 41
Neomeniomorpha, 49 Oncidiacea, 50 Pachycormiformes, 65
Neoptera, 57 Oncoceroida, 51 Pachycormoidea, 65
Neopterygii, 65 Oncopoda, 29 Pachyodonta, 51
Neornithes, 23, 68 Oniscomorpha, 19,57 Pachyodontida, 50
Neosauromorpha, 67 Onychophora, 7, 17, 37, 53 Pachypodes, 67
Neoscyphozoa, 44 Onychura, 55 Paenungulata, 70
Neosporidea, 40 Oothecaria, 58 Palaeacanthocephala, 13,
Neotaxodonta, 50 Opabiniida, 53 46
Neotremata, 15,49 Opalinida, 10,41 Palaeanostraca, 53
Nereidiformia, 52 Ophidia, 67 Palaeobranchia, 50
Neuroptera, 20, 59 Ophiocephaliformes, 66 Palaeocopida. 55
Nidulitida, 41 Ophiocistioidea, 62 Palaeocoxopleura, 57
Nippotaeniidea, 12, 46 Ophiocystia, 62 Palaeocrinoidea, 60
Nomarthra, 70 Ophiocystiida, 21,61 Palaeodictyoptera, 58
Nomomeristica, 52 Ophiocystioidea, 62 Palaeo-echinoidea, 62
Non-calcarea, 41 Ophiureae, 61 Palaeognathae, 68
Notacanthiformes, 66 Ophiurida, 61 Palaeohemiptera, 59
Notaspidea, 50 Ophiuroidea, 21, 61 Palaeohymenoptera, 58
Nothosauria, 67 Opiliones, 18,54 Palaeolamellibranchia, 50
Notidanoidea, 65 Opilionidea, 54 Palaeonemertea, 13,46
Notodelphyoida, 18, 56 Opisthandria, 57 Palaeonemertini, 46
Notommatoidea, 47 Opisthobranchia, 15, 50 Palaeonisciformes, 65
Notoptera, 58 Opisthocoela, 67 Palaeoniscoidea, 65
Notostigma, 57 Opisthocomi, 69 Palaeopantopoda, 54
Notostigmata, 55 Opisthocomiformes, 69 Palaeoptera, 57
Notostigmophora, 19, 57 Opisthogoneata, 53 Palaeopterygii, 65
Notostraca, 18, 55 Opisthomi, 66 Palaeospondyliformes, 65
Notoungulata, 71 Opisthoparia, 54 Palaeospondyloidea, 65
Nuda Opisthopora, 17, 52 Palaeostraca. 54
Ctenophora, 12,45 Opisthospermophora, 19, 57 Palaeotaxodonta, 51
Protozoa, 40 Ornithae, 68 Palaeotremata, 49
Vertebrata, 67 Ornithischia, 68 Palechinoidea, 62
Nudibrachiata, 43 Ornithodelphia, 69 Paleoconcha, 51
Nudibranchiata, 50 Ornithopoda, 68 Paleocopa, 55
Nummulitidea, 40 Ornithosauria. 67 Paleo-echinoidea, 62
Nymphonomorpha, 54 Ornithoscelida, 67 Paleoloricata, 49
Orthida, 48 Paleomorpha, 57
Orthochoanites, 51 Paleonemertea, 46
0:tacnemida, 22, 63 Orthonectida, 11,42 Paleotremata, 49
Octactinellida, 41 Orthopoda, 68 Palliata, 37,48
Octactinia, 44 Orthoptera, 20, 58 Palliobranchiopoda, 37, 48
Octobrachiata, 51 Ornithurae, 68 Palpigradi, 55
Octocorallia,44 Osculosida, 40 Palpigradida, 18, 55
Octopoda, 16, 51 Ospiiformes, 65 Pandictyoptera, 58
Octopoida, 51 Ostariophysi, 66 Panisoptera, 59
INDEX TO Latin names 90
Panorpatae, 59 Pentamerida, 48 Pholidota
Panorpina, 59 Pentastomida, 7, 17, 37, 53 Amphibia, 67
Panorpoidea, 59 Peracarida, 56 Mammalia, 25, 70
Panprotura, 57 Perciformes, 66 Phoronida, 7, 15, 37, 48
Panthysanura, 57 Percomorphoidea, 66 Phragmophora, 51
Pantodonta Percopseiformes, 66 Phryganoidea, 59
Mollusca, 51 Perennichordata, 63 Phryneides, 55
Vertebrata, 71 Peridiniaceae, 39 Phrynichida, 18,55
Pantodontida, 50 Perielytrodea, 57 Phrynophiurida, 21, 61
Pantopoda, 54 Peripatidea, 17, 53 Phthinorhabdina, 41
Pantostomatida, 39 Peripylaria, 40 Phthiraptera, 58
Pantotheria, 70 Perischoechinoida, 62 Phylactolaemata, 15, 48
Parablastoidea, 60 Perissodactyla, 25,71 Phyllobothrioidea, 45
Paracephalophora, 49 Peritricha, 10, 41 Phyllocarida, 56
Paracoleoptera, 59 Perlaria, 58 Phyllolepida, 64
Paracrinoidea, 61 Perlarides, 58 Phyllolepiformes, 64
Paractinopoda, 62 Perleidiformes, 65 Phyllopoda, 55
Parafilibranchia, 50 Perloidea, 58 Phylloptera, 58
Paramecoptera, 59 Permodictyoptera, 58 Phyllorhyncha, 46
Paramera, 44 Permodonata, 58 Phyllospondyli, 67
Paranemata, 45 Permoneurodia, 57 Physapida, 59
Paraneuroptera, 58 Peromedusae, 44 Physaraceae, 40
Paraplecoptera, 59 Peromela, 66 Physoclysti, 66
Parapsida, 67 Petalichthyida, 64 Physophora, 43
Parareptilia, 67 Petalodontes, 65 Physopoda, 59
Pararthropoda, 29 Petalodontiformes, 65 Physostomi, 66
Parasita, 59 Petanoptera, 59 Phytoflagellata, 39
Parastigmata, 55 Petromyzonoidea, 64 Phytomastigina, 9, 39
Parasuchia, 67 Petromyzontia, 64 Phytomastigophorea, 39
Paratheria, 70 Petromyzontiformes, 64 Phytomonadina, 9, 39
Paratrichoptera, 59 Phacopida, 54 Phytosauria, 67
Paraxonia, 71 Phaeoconchia, 40 Picariae, 68
Parazoa, 6, 36 Phaeocystina, 40 Pici, 24, 69

Pareiasauria, 67 Phaeodaria, 40 Piciformes, 24, 69


Passeres, 25, 69 Phaeogromia, 40 Pinnipedia, 70
Passeriformes, 25, 69 Phaeosphaeria, 40 Piroplasmidea, 41
Paucituberculata, 70 Phalangida, 18,54 Pisces, 64
Pauropoda, 19, 56 Phalangiotarbi, 54 Pithecoidea, 70
Pectinibranchia, 50 Phalansteriina, 39 Placentalia, 25, 70
Pectobothrii, 45 Phallostethiformes, 66 Placentaria, 70
Pectinifera, 54 Phanerocephala, 52 Placipennes, 59
Pedata, 62 Phaneropleuriformes, 66 Placodermi, 64
Pedicellinida, 14, 48 Phanerorhynchiformes, 65 Placodontia, 67
Pediculati, 66 Phanerozonia, 21, 61 Placoidei, 65
Pediculatiformes, 66 Pharetronida, 10, 41 Placophora, 49
Pediculidea, 59 Pharyngidea, 16, 51 Plagiaulacoidea, 69
Pedipalpida, 55 Pharyngobdellida, 17, 52 Plagiostomi, 65
Pegasiformes, 66 Pharyngobdelliformes, 52 Plagiotremata, 67
Pelagothurida, 63 Pharyngobranchii, 37, 63 Planaria, 45
Pelargiformes, 69 Phasmidia Planctosphaeroidea, 7, 22,
Pelargomorphae, 68 Arthropoda, 20, 58 37,63
Pelecani, 24, 69 Nematoda, 47 Planipennia, 59
Pelecaniformes, 24, 69 Phasmoidea, 58 Planuloidea, 36, 42
Pelecypoda, 16, 50 Phlebobranchiata, 22, 63 Plasmodiida, 41
Pelicaniformes, 69 Phlebolepiformes, 64 Plasmodroma, 39
Pelmatozoa, 60 Phoenicopteri, 69 Plastidozoa, 36, 39
Pelycosauria, 68 Phoenicopteriformes, 69 Plathelminthes, 45
Penicillata, 57 Pholadacea, 50 Platodes, 36, 45
Pennatulacea, 11,44 Pholidophoriformes, 65 Platosomia, 65
Pentacrinacea, 61 Pholidophoroidea, 65 Platyasterida, 61
Pentacrinoidea, 61 Pholidopleuriformes, 65 Platycopa, 55
91 INDEX TO Latin names
Platyctenea, 12, 45 Polypiaria, 43 Proterandria, 57
Platydesmiformia, 57 Polyplacophora, 49 Proterospermophora, 19, 57
Platyelmia, 45 Polyplakiphora, 49 Prothysanura, 57
Platyhelmia, 45 Polyplaxiphora, 49 Protichthyes, 64
Platyhelminthes, 6, 12, 36, Polypoda, 37, 52 Protista, 7, 26
45 Polyprotodontia, 70 Protoblastoidea, 60
Platymalakia, 49 Polyptera, 59 Protoblattoidea, 58
Platypoda, 50 Polypterei, 65 Protobranchia, 16,50
Platyrhina, 70 Polypteriformes, 65 Protocicadida, 59
Platysiagiformes, 65 Polypterini, 65 Protociliata, 10,41
Plecoptera, 20, 58 Polystomea, 45 Protocoleoptera, 58
Plectellaria, 40 Polythalamia, 40 Protodiptera, 59
Plectognathoidea, 66 Polytricha, 41 Protodonata, 58
Plectoidea, 40 Polyzoa Protodonta, 68
Plectoptera, 58 Bryozoa, 37, 48 Protofulgorida, 59
Pleospongia, 36, 42 Platyhelminthes, 45 Protogastropoda, 49
Plesiocidaroida, 62 Porifera, 6, 10,36,41 Protohemiptera, 58
Plesiosauria, 67 Porocephalida, 17, 53 Protohymenoptera, 58
Plesiothecata, 5217, Porolepiformes, 66 Protomastigaceae, 39
Pleuracanthodii, 65 Porulosida, 40 Protomastigida, 39
Pleurobranchomorpha, 50 Praeheterodonta, 50 Protomecoptera, 59
Pleurocoela, 15, 50 Predentata, 68 Protomedusae, 43
Pleuroconcha, 50 Priapulida, 14, 47 Protomonadina, 9, 39
Pleurodonta, 50 Priapuloidea, 6, 14, 36, 47 Protonemertini, 46
Pleurogona, 63 Primates, 25, 70 Protonephridozoa, 36
Pleuronectiformes, 66 Prionodesmacea, 51 Protonychophora, 53
Pleuronectoidea, 66 Proactiniae, 44 Protoparia, 54
Pleuropterygii, 65 Proanura, 66 Protoperlaria, 58
Pleuropygia, 49 Proarthropoda, 53 Protorhabdina, 41
Pleurosauria, 67 Proboscidea Protorosauria. 67
Pleurostigma, 57 Mammalia, 25, 71 Protorthoptera, 58
Pleurostigmophora, 19, 57 Myzostomida, 16, 51 Protospondyli, 65
Pleurostomata, 65 Procellariae, 24, 68 Protosuchia, 67
Pleurotremata, 65 Procellariiformes, 24, 68 Protosyngnatha, 57
Pliomeria, 54 Prochelicerata, 53 Prototardigrada, 52
Ploima, 13,47 Procoela, 67 Prototheria, 25, 68, 69
Ploimoidaceae, 47 Procolophonia, 67 Protozoa, 6, 9, 36, 39
Podicipedes, 24, 68 Proganosauria, 68 Protracheata, 37, 52
Podicipediformes, 24, 68 Progoneata, 53 Protremata, 15,49
Podicipiformes, 68 Prolacertiformes, 67 Protungulata, 70
Podocopa, 55 Promammalia, 68, 69 Protura, 20, 57
Podocopida, 18, 55 Proparia, 54 Pruvostitoptera, 58
Podogonata, 18, 55 Prorhipidoglossomorpha, 49 Psammosteiformes, 64
Podophthalma, 56 Prorocentraceae, 39 Pselaphocephala, 49
Poecilopoda, 54 Proselachii, 65 Pselaphognatha, 19, 57
Poecilosclerida, 10, 42 Prosimii, 70 Pseudanostraca, 53
Poeobioidea, 52 Prosobranchia, 15, 49 Pseudaxonia, 44
Pogonofora, 59 Prosopocephala, 50 Pseudechinoidea, 62
Pogonophora, 7,21,37,60 Prosopora, 17, 52 Pseudocentrophori, 66
Poikilorhabdina, 41 Prosostomata, 45 Pseudocoelia, 36
Polyactinia, 44 Prosothecata, 17, 52 Pseudocoelomata, 6, 36
Polybranchiata, 49 Prostigmata, 55 Pseudocrustacea, 53
Polychaeta, 17,52 Protalcyonacea, 44 Pseudohemiptera, 58
Polycladida, 12,45 Protarthropoda, 29 Pseudolamellibranchia, 51
Polycyclia, 44 Protaxonia, 27, 36 Pseudonotostraca, 53
Polycystina, 40 Protelytroptera, 58 Pseudophyllidea, 12,46
Polydesmoidea, 57 Proteocephala, 12,45 Pseudorhynchota, 59
Polymastigina, 9, 39 Proteocephaloidea. 45 Pseudoscorpionida, 18, 54
Polymera, 54 Proteomyxa, 9, 39 Pseudosuchia, 67
Polynemiformes, 66 Protephemerida, 57, 58 Psittaci, 24, 69
Polyopisthocotylea, 45 Protephemeroidea, 58 Psittaciformes, 24, 69
INDEX TO Latin names 92
Psocoptera, 20, 59 Retioloidea, 42 Sagenocrinoidea, 60
Pteraspida, 64 Rhabdiasoidea, 14, 47 Sagittoidea, 21, 59
Pteraspides, 64 Rhabditoidea, 14, 47 Salientia, 23, 66
Pteraspidiformes, 64 Rhabdocoela, 12, 45 Salinella,27
Pteraspidomorpha, 64 Rhabdogeniae, 41 Salmopercoidea, 66
Pterichthyes, 64 Rhabdophora, 42 Salpida, 63
Pterichthyomorpha, 64 Rhabdopleurida, 22, 62 Sanatoria, 20, 58
Pteriomorpha, 50 Rhachiglossa, 50 Sarcobranchiata, 49
Pterobranchia, 7, 22, 37, 62 Rhachitomi, 66 Sarcocystidea, 40
Pterocletes, 68 Rheae, 24,68 Sarcodina, 9, 39
Pteroconchida, 50 Rheiformes, 24,68 Sarcosporidia, 10, 40
Pterocorallia, 44 Rhenanida, 64 Saurabatrachia, 66
Pterodactyli, 67 Rhignogastra, 55 Sauria, 67
Pteropoda, 16, 50 Rhinocarina, 56 Saurichthyiformes, 65
Pterosauria, 67 Rhinogastra, 55 Saurischia, 68
Pterygogenea, 57 Rhipidistia, 66 Sauropoda, 68
Pterygota, 57 Rhipidoglossa, 49 Sauropsida, 64
Ptychobranchia, 63 Rhipidoptera, 59 Sauropterygia, 67
Ptychodactiaria, 44 Rhipiptera, 59 Saururae, 68
Ptychopariida, 54 Rhiptoglossa, 67 Scansores, 69
Ptyctodontida, 64 Rhizocephala, 19, 56 Scaphopoda, 16, 50
Ptyctodontiformes, 64 Rhizoflagellata, 39 Schistochoanites, 51
Pulicina, 59 Rhizomastigaceae, 39 Schizocoela, 27
Pulmonata, 16, 50 Rhizomastigina, 9, 39 Schizocoralla, 44
Pycnodontiformes, 65 Rhizopoda, 9, 39 Schizodonta, 51
Pycnodontoidea, 65 Rhizostomatida, 44 Schizodontida, 50
Pycnogonida, 17, 54 Rhizostomeae, 1 1, 44 Schizogregarinida, 41
Pycnogonomorpha, 54 Rhizota, 46 Schizomida, 18, 55
Pygaspida, 53 Rhombifera, 60 Schizonemertini, 46
Pygocaulia, 49 Rhomboganoidei, 65 Schizonotida, 55
Pygocephalomorpha, 56 Rhombozoa, 42 Schizopeltidia, 55
Pygopodes, 68 Rhynchobdellida, 17,52 Schizopoda, 56
Pyrosomata, 22, 63 Rhynchobdelliformes, 52 Schizosiphona, 50
Pyrosomatida, 22, 63 Rhynchocephalia, 23, 67 Scirtopoda, 47
Pyrotheria, 71 Rhynchocoela, 6, 13, 36, 46 Scleractinia, 11,44
Pythonomorpha, 67 Rhynchodipteriformes, 66 Scleropareioidea, 66
Rhynchoflagellata, 39 Scoleciformia, 52
Rhynchonellida, 48 Scolecomorpha, 49
Quadrumana, 70 Rhynchostomi, 54 Scolopendromorpha, 19,57
Rhynchota, 58 Scopeliformes, 66
Ricinuleida, 18, 55 Scorpiones, 54
Rachiglossa, 50 Rodentia, 25,70 Scorpionida, 18, 54
Radiata, 27 Rophoteira, 59 Scutibranchia, 49
Radiolaria, 9,40 Rostrata, 55 Scutigeromorpha, 19, 57
Rajiformes, 65 Rostroconchida, 50 Scyphomedusae, 44
Ralliformes, 69 Rotalidea, 40 Scyphopolypi, 44
Raphidioidea, 59 Rotatoria, 6, 13, 36,46 Scyphozoa, 11,44
Ratitae, 68 Rotifera, 6, 13,36,46 Scytactinata, 62
Receptaculitida, 42 Roveacrinida, 60 Scytodermata, 62
Reculoidea, 59 Rugosa, 44 Sebecosuchia, 67
Redfieldiiformes, 65 Rutoceratida, 51 Sedentaria, 17, 52
Redlichiida, 54 Seisonacea, 13,46
Regulares, 60 Seisonidea, 13, 46
Regularia Saccata Selachii, 23, 65

Crinoidea, 61 Ctenophora, 45 Selachoidea, 65


Echinoidea, 21, 62 MoUusca, 37,48 Semaeostomeae, 1 1, 44
Remigolepiformes, 64 Saccocirrida, 52 Semaeostomatida, 44
Reptilia, 23, 67 Saccopharyngiformes, 66 Semi-articulata, 61
Reptiliomorphoidea, 67 Saccosomatida, 16,51 Semionotoidea, 65
Reticularia, 40 Sacoglossa, 16, 50 Semostomeae, 44
Reticulariaceae, 40 Sagenocrinida, 60 Sepioidea, 51
93 INDEX TO Latin names
Septibranchia, 16, 50, 51 Squaliformes, 65 Suctoria
Septibranchida, 50 Squaloidea, 65 Arthropoda, 59
Serosporidia, 41 Squamata Protozoa, 10,41
Serpentes, 67 Mammalia, 70 Sycones, 41
Sertularina, 43 Reptilia, 23,67 Syconosa, 41
Seymouriamorpha, 67 Stauracea, 44 Symbranchiformes, 66
Sigmatomonaxonellida, 41 Stauromedusae, 1 1 44 , Symmetrodonta, 70
Sigmatophora, 42 Steganobranchia, 50 Symphyla, 19, 57
Silicoflagellata, 39 Steganopodes, 24, 69 Sympoda, 56
Silicospongiae, 41 Stegocephalia, 66 Synalcyonacea, 44
Simiae, 70 Stegosauria, 68 Synapsida, 68
Simplicidentata, 70 Stegoselachii, 65 Synaptera, 57
Sinupalliata, 51 Stelechotokea, 44 Synaptida, 62
Siphonapiera, 20, 59 Stelleroidea, 60 Synaptorhabda, 50
Siphonida, 50 Stelliformia, 60 Synaptosauria, 67
Siphonobranchia, 50 Stelmatopoda, 48 Synarmogoidea, 58
Siphonophora, 1 1 43 , Stemonitaceae, 40 Synbranchioidea, 66
Siphonopoda, 50 Stenoglossa, 50 Syncarida, 56
Siphunculata, 59 Stenolaemata Synentognathoidea, 66
Sipunculida, 16,51 Bryozoa, 48 Syngnatha, 57
Sipunculoidea, 7, 16, 37, MoUusca, 49 Syngnathiformes, 66
51 Stenostomata, 48 Syntonopteroidea, 57
Sirenia, 25,71 Stensioellida, 65 Synxiphosurida, 54
Sirenoidei, 66 Stensioelliformes, 65 Sypharopteroidea, 57
Solenichthyoidea, 66 Stenurida, 61 Syringocnemida, 42
Solenida, 10,41 Stephanoberyciformes, 66
Solenochilida, 51 Stephoidea, 40
Solenoconchia, 50 Stereornithes, 68
Solenogastres, 15, 49 Stereospondyli, 66 Tabulata, 44
Solifugae, 18,55 Stethostomata, 54 Taeniodontia, 70
Solpugida, 18,55 Stichodactylina, 44 Taenioglossa, 50
Soluta, 54 Stirodonta, 62 Taenioidea, 46
Somasteroidea, 61 Stolidobranchiata, 22, 63 Taligrada, 70
Somphocyathida, 42 Stolonifera, 11,44 Tanaidacea, 19, 56
Sorophora, 40 Stolonoidea, 43 Tanysitrachelia, 67
Spatangoida, 22, 62 Stomatocrinoidea, 61 Tardigrada, 7, 17, 37, 52
Spathebothridea, 12, 46 Stomatopoda, 19, 56 Tarphyceratida, 51
Spelaeogriphacea, 19, 56 Stomatostigmata, 55 Tarrasiiformes, 65
Sphaeractinoidea, 43 Strepsata, 59 Tartarides, 55
Sphaerellaria, 40 Strepsiptera, 21, 59 Taxeopoda, 70
Sphaeroidocrinacea, 60 Streptoneura, 49 Taxocrinida, 60
Sphaerozoa, 40 Streptophiurae, 61 Taxocrinoidea, 60
Sphenisci, 23, 68 Streptophiuroida, 61 Taxodonta, 51
Sphenisciformes, 23, 68 Streptostylica, 67 Taxodontia, 70
Sphinctozoa, 41 Strigeata, 45 Tectibranchiata, 50
Spinigrada, 61 Striges, 24,69 Tectobranchia, 50
Spinulosa, 21, 61 Strigiformes, 24, 69 Tectospondyli, 65
Spioniformia, 52 Stromatoporidea, 43 Teleodesmacea, 51
Spiriferida, 48 Stromatoporoidea, 43 Teleodonta, 51
Spirigera, 41 Strongyloidea, 14, 48 Teleoplacophora, 49
Spirobranchia, 37, 48 Strophomenida, 48 Teleostei, 23,65
Spirobranchiopoda, 48 Struthiones, 23, 68 Teleostomi, 65
Spironotia, 49 Struthioniformes, 23, 68 Telestacea, 11,44
Spirotricha, 10,41 Stylasterina, 11,43 Telmatomorphormes, 69
Spiruroidea, 14, 48 70
Stylinodontia, Telobranchia, 49
Spongeae, 36, 41 Stylommatophora, 16, 50 Telogonia, 47
Spongiaria, 36, 41 Stylopida, 59 Telosporidia, 9, 40
Spongiomorphida, 43 Stylopidae, 59 Telotremata, 15, 49
Sporozoa, 9, 40 Subholostei, 65 Temnocephaloidea, 45
Spumellaria, 40 Subholosteoidea, 65 Temnospondyli, 67
Spyroidea, 40 Subungulata, 70 Tentaculata, 12,45
INDEX TO Latin names 94
Tentaculifera Thrissomorphi, 66 Tubulosa, 44
Mollusca, 50 Thunniformes, 66 Tunicata, 7,22,37,63
Protozoa, 41 Thyroidea, 61 Turbellaria, 12,45
Terebelliformia, 52 Thyrostraca, 56 Turbinares, 68
Terebratulida, 48 Thysanoptera, 20, 59
Terricolae, 52 Thysanura, 20, 57
Tesselata, 61 Tillodontia, 70 Uintacrinida, 60
Testacea Tinami, 24,68 Uintatheria, 71
Echinodermata, 60 Tinamiformes, 24, 68 Uncinataria, 41
Protozoa, 9, 40 Tintinnina, 41 Unguiculata, 70
Testicardines, 49 Tjalfiellidea, 45 Ungulata, 70
Testudinata, 67 Tomiosoma, 45 Ungulates, 70
Testudines, 67 Torpediniformes, 65 Uranocyatha, 42
Tethyodeae, 63 Toxodontia, 70 Urochordata, 37, 63
Tetrabothridiata, 46 Toxoglossa, 50 Urodela, 23, 66
Tetrabranchiata, 16, 50 Trachelosauria, 67 Urodeloidei, 66
Tetracoelia, 44 Trachomedusae, 43 Uronemiformes, 67
Tetracorallia, 44 Trachylinida, 11,43 Uropygi, 55
Tetracotylea, 46 Trachymedusae, 43
Tetractinellida, 41 Trachypsammiacea, 44
Tetractiniae, 45 Tremataspidiformes, 64 Vaginulacea, 50
Tetradecapoda, 56 Trematoda, 12,45 Vampyromorpha, 16,51
Tetradida, 44 Trematosauria, 66 Veneracea, 50
Tetradontiformes, 66 Trepostomata, 48 Vertebrata, 7,23,37,64
Tetraphyllidea, 12,45 Tretenterata, 49 Volvocaceae, 39
Tetrapoda, 64 Triaxonida, 41 Volvocales, 9, 39
Tetrarhyncha, 46 Trichinelloidea, 48 Volvocina, 39
Tetrarhynchoidea, 46 Trichiaceae, 40
Tetraseptata, 44 Trichocorallia, 44
Tetraxonida, 41 Trichomonadida, 39 Waptiida, 53
Tetrodontiformes, 66 Trichoptera, 20, 59 Weigeltisauria, 67
Teuthoidea, 51 Trichostomata, 41
Textularidea, 40 Trichuroidea, 14, 48
Thalamida, 10,41 Tricladida, 12, 45 Xenacanthi, 65
Thalamophora, 40 Triconodonta, 69 Xenacanthoidea, 65
Thalattosauria, 67 Trigonotarbi, 54 Xenanthi, 65
Thalattosuchia, 67 Trigonotarbida, 54 Xenarthra, 70
Thaliacea, 22,63 Trilobita, 54 Xenophyophoridea, 40
Thallocoralla, 44 Trilobitoidea, 53 Xenopoda, 53
Thecamoebae, 40 Trilobitomorpha, 52 Xenopneusta, 16, 51
Thecanephria, 21,60 Trimyaria, 46 Zeomorphoidea, 66
Thecaphora, 43 Trinucleida, 54 Xiphosura, 17, 54
Tbecata, 43 Triplesiida, 48 Xiphosurida, 17, 54
Thecodontia, 67 Tripostomata, 48
Thecoidea, 61 Tripylaea, 40
Thecosomata, 50 Tripylaria, 40 Zacanthoidacea, 54
Thelodonti, 64 Trituberculata, 70 Zeiformes, 66
Thelyphonida, 18,55 Tritylodontoidea, 69 Zeomorphoidea, 66
Therapsida, 68 Trochelminthes, 28 Zeuglodontia, 70
Theria, 69 Trochosphaerida, 47 Zoa, 36
Therictoidea, 70 Trogones, 24, 69 Zoantha, 44
Theriodontia, 68 Trogoniformes, 24, 69 Zoanthactiniaria, 44
Thermosbaenacea, 19, 56 Tryblidiacea, 49 Zoantharia, 11,44
Theromorpha, 68 Tryblidioidea, 15,49 Zoanthidea, 44
Theropoda, 68 Trypanorhyncha, 12, 46 2^anthiniaria, 11,44
Thigmotrichina, 41 Tubicola, 52 Zoomastigina, 9, 39
Thoracica, 18,56 Tubinares, 24,68 Zoomastigophorea, 39
Thoracostoidea, 66 Tuboidea, 43 Zoophytaria, 44
Thoracostraca, 56 Tubulariae, 43 Zoraptera, 20, 59
Thripoides, 59 Tubulidentata, 25,71 Zygobranchia, 49
Thripsites, 59 Tubulinaceae, 40 Zygophiurae, 61
Classification
of the
ANIMAL KINGDOM
BY Richard E. Blackwelder

The classiiTX*st!!dit df irSfflSf^r^^ a


field in which discovery and revision are continu-
ing, even after two hundred years of study. Yet,
the importance of classification in biology in-
creases every year because the experimental and
practical fields find increasing need for accurate
identification of animals and for understanding of
comparative relationships.
Classification of the Animal Kingdom is a

completely new classification system. It is de-


signed to promote interest and study, elicit dis-

cussion, and provide a usable single classification

for reference for zoology students and teachers.


This classification has three principal parts:
( 1 ) the complete classification, including lists of

the phyla and of the classes and orders of all

animals, living and fossil; (2) the justification for


unusual features in the classification system; and
(3) a simplified classification of Recent animals
for student use. A bibliography of major recent
classifications and indexes to common and Latin
names are also included.
Sponsored by the Society of Systematic
Zoology.

RICHARD E. BLACKWELDER is an Associate Pro-


fessor of Zoology at Southern Illinois University.
He formerly was Associate Curator at the Ameri-
can Museum of Natural History, New York, and
at the National Museum, Washington, D.C. In
1961 he was President of the Society of Syste-
matic Zoology. He is the author of more than
one hundred monographs and articles and com-
piler, with Ruth M. Blackwelder, of Directory of
Zoological Taxonomists of the World.

Press
Carbondale, Illinois

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