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A PROPOSED SOLUTION
SHORT-CUT METHODS
After consideration of a number of methods for measuring various
factors which might be associated with forage weight, we finally
selected two short-cut methods as being the most likely to fit our
needs. One, the line-transect, has been used in various fields and was
adapted to forage measurements by Canfield (2, 3). The other, which
simply involves the estimation of forage weight on sample plots (5),
has been tried by a number of investigators.
If used without quantitative control, neither of these methodspro-
vides data on forage weight directly, and they are not capable of
rigorous test to determine the accuracy of information on forage ob-
tained by their use. By the clipping of forage on someof the estimated
plots and in a belt around some of the line transects, however, the
data obtained by these short-cut methods could be converted to
quantitative weight estimates. Both of the methods seemed to prom-
ise considerable reductions in field time requirements, and both ap-
peared simple and easy to apply. And’finally, quantitative estimates
of the error of forage weights calculated from the field data could be
obtained by double-sampling analysis.
For this approximate calculation the travel cost per plot may be
taken as essentially constant with varying numbers of sample observa-
tions, within the limits imposed by a fixed total cost of sampling.
WILM, ET AL. : ESTIMATING FORAGEYIELD I97
DESCRIPTION OF EXPERIMENTS
In order to fit the studies of sampling methods into active range experiments
with maximumefficiency, we tried out the two methods at two different locations.
The line-transect trial was made in experimental pastures at the Manitou Experi-
mental Forest, in the headwaters of the South Platte River, while the efficiency
of weight estimates was tested in a study of the influence of gopher removal on
forage production, located on the Grand Mesa National Forest in western Colo-
rado. Thus, line transects and weight estimates could not be compared directly
with each other, but the efficiency of each method could be compared with that
obtained by clipping quadrats.
LINE-TRANSECT STUDY
Experimental area.--At the Manitou Experimental Forest six pastures, each
25o to 3oo acres in size, have been established to study the influence of three in-
tensities of grazing by cattle on forage production, beef yields, erosion, and in-
filtration.
The forage on these pastures is primarily a bunchgrass type existing as an
understory in an open ponderosa pine stand. Park-like areas from a few square
rods to several acres in extent are dominated by bunchgrasses. In areas occupied
by open stands of ponderosa pine the same species persist but in lighter density
than in the park-like areas. In spots where the canopy is dense, and directly be-
neath the pines, a few sedge plants are the only species present.
Field procedure.--In ascertaining forage production, a total of 36 plots was
sampled in each pasture. At each plot were measured the ground-level diameter
(to the nearest o.o~ foot) and the average height (to the nearest o.Io foot) of
portion of every plant which touched a 3o-foot cable stretched between two iron
stakes. These plants were classified as to whether grazed or ungrazed, and segre-
gated into four classes, v/z., tall bunchgrasses, short bunchgrasses, single-stemmed
and sod-forming grasses and grasslike plants, and weeds (forbs). Browse species
were ignored, as they form a minute part of the palatable forage in this cover type.
Six of thh 36 transects were selected at random for clipping. After the line-
transect data were obtained, the vegetation was clipped from a 6-inch by 3o-foot
belt transect surrounding the 3o-foot line-transect. The forage was sorted by
classes, air-dried, and then weighed to the nearest gram. In addition to these data,
each crew kept accurate notes on the amount of time consumed in clipping, tally-
ing, travel between plots, and handling the clipped forage.
WEIGHT-ESTIMATE STUDY
Experimental area.--The Grand Mesa experiment is factorial in design. Its
objective is to discover .the influence of grazing by cattle and gophers, separately
and together, on forage production and related factors. Sixteen I-acre areas have
been established, four in each of four locations or blocks. The four treatments,
cattle and gophers, cattle alone, gophers alone, and no grazing, have been assigned
at random to the 4 acres in each block.
The study area is on comparatively level to rolling land at an elevation of ap-
proximately lO,5OOfeet, where the vegetation consists of open grassland and sage-
brush interspersed with stands of Engelmann spruce. All observations were con-
fined to the park-like areas of sagebrush and grass-weed communities, since the
ground beneath the spruce trees is practically devoid of forage. Grasses constitute
approximately 2o% of the vegetal cover, varying from 8 to 3o%, depending
1.98 JOURNAL OF THE AMERICAN SOCIETY OF AGRONOMY
largely on the intensity of use in the past by grazing animals. Weedsare abundant,
constituting from 55 to more than 90% of the cover. On approximately one half
of the study area sagebrush is the dominant plant, the understory consisting of
grasses and weeds.
Field procedure.--For sampling the forage, each I-acre area.was subdivided into
nine strata and two sample plots were assigned at random to each stratum. These
plots, each 12. 5 square feet in drea, provided the large sample. For the small
sample, five plots were drawn from the 18 plots in each acre.
Estimates of green forage weight were obtained by one man, who had previously
trained himself by estimating the forage 6n a number of plots and checking his
estimates by clipping and weighing the forage. In sampling the experimental plots,
each species was estimated separately; then the species estimates were combined
into three classes, grasses, weeds, and shrubs.
After the plots had been estimated on a single acre, a~lother worker clipped the
forage on the five plots forming the small sample, segregating it into the three
classes and weighing the clip in green condition. The estimator was then permitted
to compare his estimates with the actual green w.eights as a running control on
subsequent estimates. Finally, the clipped forage was air-dried and reweighed ~o
the nearest gram.
RESULTS
LINE-TRANSECT METHOD
In Table 1 are presented," together with their standard errors,
numerical estimates of the air-dry weight (in pounds per acre) of each
class of forage on the six pastures as calculated by the double-sampling
method. For simplicity and because they were similar in magnitude,
the separate pasture standard errors were pooled to provide a single
value for each forage class.
TABLE
I.--Average estimated forag~ weight in pounds per acre per pasture,
by forage classes.
Forage class
Pasture Total
No. forage
I II III IV
WRIGHT-ESTIMATE METHOD
AS shown in Table 2, the forage on these 16 l-acre areas was esti-
mated with satisfactory precision. Of the three forage classes, only
browse showed poor results; and the sampling errors for total forage
are considerably smaller (in percentages of forage weights) than those
observed in the line-transect method. By itself this observation does
not favor weight estimates, however, as the sampling was consider-
ably more concentrated than in the Manitou experiment.
TABLE
2.--Average estimated forage weight per acre, by forage classes.
Forage classes
Acre desig- Total,
nation lbs.
Grasses, lbs. Weeds, lbs. Browse, ibs,
~cime was negligible on these small areas and was therefore included
in the other two components, while the actual clipping required 68.4
man-minutes per I2.S square-foot plot as compared to 32.4 man-
minutes per ~ 5 square-foot plot in the Manitou pastures.
As observed in the line-transect study, no significant differences
existed amongthe individual acre regressions. Therefore, it was pos-
sible to use a single regression (based on the pooled "within acre"
squares and products) for estimating the mean forage production on
each of the ~6 acres. Since, without previous experience, this fact
could not have been predicted beforehand, we used a rather inefficient
proportion of the total numberof plots per acre to the numberclipped
in each acre. As calculated by the equation on page 196, this ratio
should have been about 5.7 estimated plots for each clipped plot.
Although 18 plots were estimated and 5 of these plots were clipped
on each acre, we actually used a ratio of o.22 to 1.oo (that is, 18
estimated plots to 8o clipped plots in the regression) in estimating
the mean forage production per acre. The efficiency of double-
sampling would have been materially improved if only one or two
plots had been clipped on each acre and the time thus saved had been
used in estimating a larger numberof plots per acre.
Even with this relatively inefficient arrangement, double-sampling
with weight estimates provided a substantial increase in information
as comparedto clipping all plots. Onlyabout 6.8 plots could be clipped
in the field time required for double-sampling with 5 clipped and 18
estimated plots per acre. As a result, the latter methodprovided about
37%more information on total forage than that supplied by clipped
plots. Whenthe comparison is based on total time (field and office),
the gain in information dropped to about 14%. These gains are not
greatly different from the iine-transect results. If, however, only one
plot had been clipped on each acre and the surplus time used in
estimating about 4o additional plots per acre, the variance of the
calculated mean forage weight would have been reduced by about
one third, and the relative efficiency of double-sampling ~ould have
been correspondingly increased.
DISCUSSION
As a result of these analyses, we have a definite basis for quantita-
tive comparison of each double-sampling method with the cost of
clipping sample plots in our experiments, and a qualitative basis for
.comparing the two methods with each other.
As employed in these studies, both short-cut methods provided
a substantial saving in field time and some economyon the basis of
total time expended. The achievement of similar economies in other
studies would probably depend, however, on the relative amounts of
time required for clipping, traveling to and from the survey area and
between sample plots, and applying the short-cut method. In experi-
ments requiring intensive sampling by trained men, with sample
plots located close together, either of the double-sampling methods
maybe expected to provide substantial increases in efficiency as com-
pared to clipping all sample plots. In large-scale, extensive range
202 JOURNAL OF THE AMERICAN SOCIETY OF AGRONOMY
SUMMAI~Y
Two double-sampling methods, using line-transects and forage
weightestimates,weretested to ascertaintheirrelative efficiency
in
estimatingthe amountof foragepresenton experimental areas,as
comparedto theclipping of vegetationon sampleplots.
Considering fieldworkalone,double-sampling withtheline-tran-
sectmethodprovided an increase in information of about~8~oas
compared withthe information whichcouldhavebeenobtainedby
clippingonly,duringthesameperiodof time.On thebasisof time
expendedin bothfieldandoffice,thedouble-sampling methodpro-
videdonly abouti~% moreinformation thancouldhavebeenob-
tainedby clippingalone.
The use of weightestimates in double-sampling providedabout
3 7~omoreinformation, thancouldbe obtained by straight clipping
inan equivalentamountoffieldtime.If fieldworkandoffice compila-
tionare bothconsidered the gainin information droppedto about
~4%.
Underourconditions
of intensive
samplingbothmethods provided
substantial
economies
in fieldtimeandsomesavingin totaltimeex-
pended.
In otherstudies,
however,thesesavings
wouldbe consider-
WILM, ET AL.: ESTIMATING FORAGE YIELD 203