Gonad : ovarium : menghasilkan sel telur testes : menghasilkan sperma Proses pematangan telur berupa perubahan struktur, kedudukan,

n, sitoplasma, dan mencakup kematangan fungsi dan fisiologis. Sitoplasma akan menjadi tempat cadangan makanan : kuning telur (egg yolk) dan butiran minyak (oil globule) Chorion : lapisan pembungkus telur paling luar Pada chorion terdapat lubang sempit sbg pintu masuk spermatozoa ke dalam telur, disebut micropyle Spermatozoa masuk hanya kepalanya (inti) saja Setelah itu micropyle menutup untuk menghalangi masuknya sperma lain. Setelah telur dilepas ke dalam air dan dibuahi, chorion akan mengeras untuk melindungi embrio yang masih sangat sensitif

Fig.4.General pattern of development of sexual products in fishes

Perkembangan telur melalui beberapa tahap, a.l: (contoh pada ikan Mas) Stage I: The primitive egg cells (ovogonium or archovogonium) are very small, their size being hardly bigger than that of other cells (812 microns). They multiply by normal mitosis. Stage II: The egg cells grow to a size of 1220 microns, and a follicle begins forming around each egg cell. The follicle, whose function is to nurture and protect the developing egg, eventually becomes a double layer of cells. Stage III: During this stage, the egg cell grows significantly larger to attain a size of 40200 microns and becomes enclosed by the follicle. These first three stages mark the period prior to the accumulation of nutrients in the developing eggs.

Stage IV: During this stage the production and accumulation of the yolk begins; this is a process known as vitellogenesis. The egg continues to grow to a size of 200350 microns with the accumulation of drops of lipoid materials in its cytoplasm. Stage V: This marks the second phase of vitellogenesis. The cytoplasm is now full of lipoid drops and yolk production begins. The egg size reaches 350500 microns. Stage VI: This is the third phase of vitellogenesis, during which the yolk plates push the lipoid drops toward the edge of the cell where two rings begin forming. The nucleoli, which take part in protein synthesis and the accumulation of nutrients are seen adhering to the membrane of the nucleus. The size of the egg is now 600900 microns. Stage VII: The process of vitellogenesis is completed during this stage and the egg attains a size of 9001 000 microns. When the yolk accumulation ends, the nucleoli withdraw into the centre of the nucleus. The micropyle (a small opening on the egg shell) develops during this stage.

Stages IV, V, VI, and VII are the stages of vitellogenesis, when yolk is synthesized and accumulated in the egg cell. The egg is now materially ready. To reach this stage of development, the female fish needs a lot of protein in its food and a favourable temperature range. On the completion of stage VII, the egg may remain as such for several months without any change, and this forms the dormant or resting phase. This resting or dormant phase will either end in ovulation if favourable conditions occur, or in follicular putrefaction and resorption in the absence of such conditions (Figure 6).

Figure 6.The fate of developing eggs

Figure 7 The course of natural spawning Figure 7 The course of natural spawning Figure 7 The course of natural spawning

Figure 7 The course of natural spawning

Perkembangan Embrio Ikan Cleavage : pembelahan zigot sec cepat menjadi unit sel yg lebih kecil Blastulasi : proses yg menghasilkan blastula, yaitu campuran sel-sel blastoderm yang membentuk rongga penuh cairan sebagai blastocoel. Gastrulasi : proses pembelahan bakal organ yang sudah terbentuk pd saat blastulasi. Organogenesis : proses pembentukan berbagai organ tubuh. Berbagai macam organ terbentuk dari beberapa bakal organ yang terbentuk saat gastrulasi. -Usus, rongga kupffer, subnotochord rod : dari endoderm -Notochord,jantung,ginjal,gonad,sirip dada : dari mesoderm -Insang, linea literalis serta lipatan-lipatan sirip : dari ektoderm

Name of fish

Optimal temperature of incubation, C

Number of days or hours

Day-grade

Common carp (Cyprinus carpio) Pike-perch (Stizostedion lucioperca) Pike (Esox lucius) European catfish (Silurus glanis) Tench (Tinca tinca) Grass carp (Ctenopharyngodon idella) Silver carp (Hypophthalmichthys molitrix) Bighead carp (Aristichthys nobilis) Rohu (Labeo rohita) Catla (Catla catla) Asian catfish (Pangasius sutchi)

2022 1015 815 2225 2225 2225 2225 2326 2430 2430 2829

3.54 days 711 days 812 days 2.53 days 3 days 11.5 days 11.5 days 11.5 days 1420 hours 1420 hours 2325 hours

6070 100110 120 5060 6070 2430 2430 2630 2022 2022 ?

Clarias macrocephalus
Giant gourami (Osphronemus goramy) Channel catfish (Ictalurus punctatus) Grey mullet (Mugil cephalus) Coporo (Prochilodus mariae)

2630
28 2430 2022 2526

1820 hours
4448 hours 1420 hours 5060 hours 1718 hours

?
? 2022 ? 1820

Sapoara falsa (Curimata sp.)

Cachama (Colossoma oculus)

2526
2526

1516 hours
1819 hours

1618
?